Anapidae

• 
  Dippenaaria luxurians
• 
  D. luxurians

Spiders of this family are very small, usually less than two millimeters long, and lack a cribellum. They can have zero, six or eight eyes, the rear median eyes either reduced or missing. One species: 'Epigastrina typhlops' (Rix & Harvey, 2010) ^[4] has no eyes, an adaption to life underground.^[5] In some genera the carapace is modified so that the eyes are raised higher than usual. Color can range from reddish brown, yellowish brown to pale, pigment reduced, creams. Generally both margins of chelicerae have teeth, one exception being 'Acrobleps hygrophilus' who lacks retromarginal teeth.^[6] The legs are short and spineless, though there are some with reduced spination such as the genus: 'Teutoniella' ^[7] or with only one or two long spines such as 'Borneanapis'^[8]). The labium has a spur that extends between the chelicerae and can be seen when the chelicerae are spread.^[9]

Anapidae are found worldwide, particularly in South America, Africa, Asia, Australia and New Zealand. Few genera occur in North America or Europe. Only Comaroma simoni and the three species of Zangherella are found in Europe; Gertschanapis shantzi and Comaroma mendocino are found in the United States.^[1]

• 
  male Novanapis spinipes
• 
  male N. spinipes

The family Micropholcommatidae was synonymized with this family by Schütt in 2003^[10] and by Lopa et al. in 2011,^[11][2] a change that has been accepted by the World Spider Catalog.^[1] Similarly, the family Holarchaeidae was synonymized by Dimitrov et al. in 2017, and likewise accepted by the World Spider Catalog.^[12]

As of October 2025^[update], this family includes 59 genera and 233 species:^[1]

There are also genera only known as fossils, typically in Baltic amber:^[13]

• †Balticoroma Wunderlich, 2004
• †Balticonopsis Wunderlich, 2004
• †Cenotextricella Penney in Penney et al., 2007
• †Deanoorapsis Penney, 2020
• †Dubianapis Wunderlich, 2004
• †Flagellanapis Wunderlich, 2004
• †Fossilanapis Wunderlich, 2004
• †Palaeoanapis Wunderlich, 1988
• †Ruganapis Wunderlich, 2004
• †Saxonanapis Wunderlich, 2004
• †Tuberanapis Wunderlich, 2004