Alcobaça Formation

Geological formation in Portugal From Wikipedia, the free encyclopedia

The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit,[1] is a geological formation in Portugal.[2] It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals.[3][4] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.

Quick facts Type, Sub-units ...
Alcobaça Formation
Stratigraphic range: Oxfordian-Kimmeridgian
~156.9–152.5 Ma
TypeGeological formation
Sub-unitsCamadas de Guimarota
UnderliesLourinhã Formation
OverliesMontejunto Formation
Lithology
PrimaryMarl
OtherMudstone, coal, siltstone, limestone, sandstone
Location
Coordinates39.6°N 9.0°W / 39.6; -9.0
Approximate paleocoordinates33.7°N 1.6°E / 33.7; 1.6
RegionEstremadura, Leiria, Lisboa
CountryPortugal
ExtentLusitanian Basin
Type section
Named forAlcobaça, Portugal
Named byChoffat
Year defined1885
Alcobaça Formation is located in Portugal
Alcobaça Formation
Alcobaça Formation (Portugal)
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Vertebrate paleofauna

Dinosaur eggs are geographically located in Lisbon District, Portugal.[5] Dinosaur tracks are geographically located in Leiria District, Portugal.[6]

Chondrichthyes

More information Chondrichthyes of the Alcobaça Formation, Genus ...
Chondrichthyes of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
Hybodus[7] Indeterminate Guimarota Isolated Teeth
Asteracanthus[7] A. biformatus Guimarota isolated teeth, fin-spine and a head-spine
Polyacrodus[7] Indeterminate Guimarota Isolated teeth and a fin-spine
Orectolobiformes[7] Indeterminate Guimarota One isolated tooth
Batomorphi[7] Indeterminate Guimarota Isolated teeth
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Osteichthyes

More information Osteichthyes of the Alcobaça Formation, Genus ...
Osteichthyes of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
Macromesodon[7] Indeterminate Guimarota Isolated teeth
Proscinetes[7] Indeterminate Guimarota Isolated teeth and prearticulars
"Lepidotes"[7] Indeterminate Guimarota isolated lateral teeth, lower jaws, ganoid scales and isolated skull bones Two species of different size can be distinguished from the fossil material[7]
Ionoscopidae[7] Indeterminate Guimarota Isolated teeth
Macrosemiidae[7] Indeterminate Guimarota Isolated grasping tooth
Caturus?[7] Indeterminate Guimarota Isolated Teeth and a partial skull
Pachycormidae[7] Indeterminate Guimarota
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Amphibians

Albanerpetontids are one of the most numerous faunal components of the Guimarota mine with around 9000 isolated remains, including parts of the skull and limbs. Remains of indeterminate salamanders and frogs are also present but far rarer.[7]

More information Amphibians of the Alcobaça Formation, Genus ...
Amphibians of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
cf. Celtedens[7] Indeterminate Porto Pinheiro Albanerpetontid
cf. Marmorerpeton[7] Indeterminate Guimarota Indeterminate salamander
Nabia[8] N. civiscientrix Guimarota Albanerpetontid
Discoglossidae[7] Indeterminate Guimarota Indeterminate frog
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Choristoderes

More information Choristoderes of the Alcobaça Formation, Genus ...
Choristoderes of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Cteniogenys Indeterminate Guimarota
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Ornithischians

Indeterminate euornithopod remains located in Lisbon District.[5] Indeterminate stegosaurid remains present in Lisbon District.[5]

More information Ornithischians of the Alcobaça Formation, Genus ...
Ornithischians of the Alcobaça Formation
GenusSpeciesDistrictStratigraphic positionAbundanceNotesImages
Dacentrurus[5] D. armatus[5] Lisboa[5]
Stegosaurus[9] S. cf. ungulatus "...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and [calcaneus], a cervical plate, and several fragments of plates."
Trimucrodon[5] T. cuneatus[5] Lisboa[5] Later determined to be indeterminate ornithischian remains.[5]
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Saurischians

Indeterminate sauropod remains located in Leiria and Lisboa.[10]

More information Saurischians of the Alcobaça Formation, Genus ...
Saurischians of the Alcobaça Formation
GenusSpeciesDistrictStratigraphic positionAbundanceNotesImages
Allosaurus[6] Indeterminate[11] Guimarota MG 27804, an isolated maxilla belonging to a hatchling individual The ontogenetically youngest and stratigraphically oldest Portuguese Allosaurus fossil
Aviatyrannis A. jurassica Guimarota
Lusotitan[5] L. atalaiensis[5] Lisboa[5]
Ceratosaurus[5] Indeterminate[5] Lisboa[5]
Dinheirosaurus[5] D. lourinhanensis[5] Lisboa[5] "[Two] cervicals, [nine] dorsals, ribs, limb elements."[12]
Lourinhasaurus[5] L. alenquerensis[5] Lisboa[5]
Megalosaurus[5] M. insignis[5] Lisboa[5] Later determined to be indeterminate theropod remains.[5]
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Pterosaurs

More information Pterosaurs of the, Genus ...
Pterosaurs of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
aff. Rhamphorhynchus[7] Indeterminate Guimarota Isolated Fragments
Pterodactyloidea[7] Indeterminate Guimarota Teeth
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Turtles

More information Turtles of the Alcobaça Formation, Genus ...
Turtles of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Peltochelys P. duchastelli Guimarota
Platychelys Indeterminate Guimarota
Pleurosternidae Indeterminate Guimarota
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Lepidosauromorphs

More information Lepidosauromorphs of the Alcobaça Formation, Genus ...
Lepidosauromorphs of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Becklesius B. hoffstetteri Guimarota Paramacellodid lizard
Dorsetisaurus D. purbeckensis Guimarota Anguimorph lizard
Marmoretta[13] M. drescherae Guimarota Basal Lepidosauromorph
Portugalophis P. lignites Guimarota Stem-snake
Saurillodon S. proraformis Guimarota A lizard
Saurillus S. henkeli, cf. obtusus Guimarota A lizard
cf. Opisthias[14] Indeterminate Andrés, Pombal A rhynchocephalian
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Crocodyliformes

More information Crocodyliformes of the Alcobaça Formation, Genus ...
Crocodyliformes of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Goniopholis G. baryglyphaeus Guimarota A goniopholidid
Knoetschkesuchus K. guimarotae Guimarota An atoposaurid
Lisboasaurus L. mitracostatus Guimarota
Machimosaurus M. hugii Guimarota A marine machimosaurid
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Mammals

More information Mammals of the Alcobaça Formation, Genus ...
Mammals of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Bathmochoffatia B. hapax Guimarota A paulchoffatiid multituberculate
Bernardodon B. sp, B. atlanticus Porto Pinheiro A pinheirodontid multituberculate
Drescheratherium D. acutum Guimarota A paurodontid dryolestidan
Dryolestes D. leirensis Guimarota A dryolestid dryolestidan
Ecprepaulax E. anomala Porto Pinheiro A pinheirodontid multituberculate
Guimarotodon G. leiriensis Guimarota A paulchoffatiid multituberculate
Guimarotodus G. inflatus Guimarota A dryolestid dryolestidan
Haldanodon H. exspectatus Guimarota A docodont
Henkelodon H. naias, H. sp Guimarota A paulchoffatiid multituberculate
Henkelotherium H. guimarotae Guimarota A dryolestidan
Iberodon I. quadrituberculatus Porto Pinheiro A pinheirodontid multituberculate
Kielanodon K. hopsoni Guimarota A paulchoffatiid multituberculate
Krebsotherium K. lusitanicum Guimarota A dryolestid dryolestidan
Kuehneodon K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp Guimarota A paulchoffatiid multituberculate
Laolestes L. andresi Porto Pinheiro A dryolestid dryolestidan
Meketibolodon M. robustus Guimarota A paulchoffatiid multituberculate
Meketichoffatia M. krausei, M. sp Guimarota A paulchoffatiid multituberculate
Nanolestes N. krusati Porto Pinheiro A cladotherian
N. drescherae Guimarota
Paulchoffatia P. delgadoi Guimarota A paulchoffatiid multituberculate
Pinheirodon P. sp, P. vastus, P. pygmaeus Porto Pinheiro A pinheirodontid multituberculate
Plesiochoffatia P. thoas, P. peperethos Guimarota A paulchoffatiid multituberculate
Portopinheirodon P. asymmetricus Porto Pinheiro A dryolestid dryolestidan
Priacodon Indeterminate Porto Pinheiro A triconodontid
Proalbionbaatar P. plagiocyrtus Guimarota A multituberculate of uncertain placement
Pseudobolodon P. dryas, P krebsi, P. oreas Guimarota A paulchoffatiid multituberculate
Renatodon R. amalthea Guimarota A paulchoffatiid multituberculate
? Tinodon Indeterminate Porto Pinheiro A tinodontid symmetrodont
Xenachoffatia X. oinopion Guimarota A paulchoffatiid multituberculate
Paulchoffatiidae Indeterminate Guimarota 4 distinct genera with 7 species
Pinheirodontidae Indeterminate Porto Pinheiro
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Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Invertebrate Paleofauna

Ostracods

More information Ostracods of the, Genus ...
Ostracods of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
Cetacella[7] C. armata Cetacella is a typical component of Late Jurassic brackish water environments in Europe. Member of the Cypridacea[7]
C. inermis
C. striata
Theriosynoecum[7] T. wyomingense Member of the Cytheracea
Bisulcocypris[7] B. pahasapensis Member of the Cytheracea
Timiriasevia[7] T. guimarotensis Member of the Cytheracea
Poisia[7] P. bicostata Member of the Cytheracea
P. clivosa
Dicrorygima[7] D. reticulata Member of the Cytheracea
Darwinula[7] D. leguminella Member of the Darwinulacea
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Mollusca

Bivalves

More information Bivalves of the, Genus ...
Bivalves of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
Isognomon[7] I. rugosus Guimarota Dozens of well preserved shells sometimes making up entire stratigraphic layers This genus is regarded as an indicator for brachyhaline water conditions and it was likely buried in its natural habitat[7]
"Unio" "U." cf. alcobacensis Guimarota Dozens of well preserved shells with articulated valves freshwater clam
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Gastropods

More information Gastropods of the, Genus ...
Gastropods of the Alcobaça Formation
Genus Species Location Stratigraphic Position Material Notes Images
Teinostoma[7] Indeterminate Guimarota Shells Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle[7]
Cryptaulax[7] Indeterminate Guimarota Shells
Melampoides[7] M. jurassicus Guimarota Shells Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle, modern relatives like Melampus also venture onto land, which can also be expected from M. jurassicus[7]
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See also

References

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