Portal:Devonian

From top to bottom and left to right, examples of sarcopterygians: Guiyu oneiros, West Indian Ocean coelacanth, Australian lungfish and the tetrapodomorph Panderichthys rhombolepis.

Sarcopterygii (/ˌsɑːrkɒptəˈrɪdʒi.aɪ/; from Ancient Greek σάρξ (sárx) 'flesh' and πτέρυξ (ptérux) 'wing, fin')—sometimes considered synonymous with Crossopterygii (κροσσός, krossós, 'fringe')—is a clade (traditionally a class or subclass) of vertebrate animals which includes a group of bony fishes commonly referred to as lobe-finned fishes. These vertebrates are characterised by prominent muscular limb buds (lobes) within their fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fishes, the Actinopterygii, which have only skin-covered bony spines supporting the fins.

The tetrapods, a mostly terrestrial clade of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes. Their paired pectoral and pelvic fins evolved into limbs, and their foregut diverticulum eventually evolved into air-breathing lungs. Cladistically, this would make the tetrapods a subgroup within Sarcopterygii and thus sarcopterygians themselves. As a result, the phrase "lobe-finned fish" normally refers to not the entire clade but only aquatic members that are not tetrapods, i.e. a paraphyletic group.

Non-tetrapod sarcopterygians were once the dominant predators of freshwater ecosystems during the Carboniferous and Permian periods, but suffered significant decline after the Permian–Triassic extinction event. The only known extant non-tetrapod sarcopterygians are the two species of coelacanths and six species of lungfishes. (Full article...)

Archaeopteris fossil leaves

The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the spermatophytes, ancestral to both gymnosperms and angiosperms (flowering plants). They have been treated formally at the rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as Protopteridium, in which the vegetative organs consisted of relatively loose clusters of axes. Tetraxylopteris is another example of a genus lacking leaves. In more advanced aneurophytaleans such as Aneurophyton these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of progymnosperms gave rise to the first really large trees known as Archaeopteris. The latest surviving group of progymnosperms is the Noeggerathiales, which persisted until the end of the Permian.

Other characteristics:

• Vascular cambium with unlimited growth potential is present as well as xylem and phloem.
• Ancestors of the earliest seed plants as well as the first true trees.

• Strong monopodial growth is exhibited. (Full article...)

The Grey Downtonian facies occurs in the Downton Castle Sandstone Group of the British Old Red Sandstone, and more or less straddles the Devonian-Silurian boundary. The Ludlow Bone Bed and Temeside Shales are sometimes also included in the Grey Downtonian, which is also referred to as the Temeside group, part of the Downton Series. It is intermediate between the marine flagstones beneath it and the terrestrial deposits above it. The beds were deposited in a marine environment, with some material being washed in from the nearby land. (Full article...)