Agoudim Formation
Geological formation in Morocco
From Wikipedia, the free encyclopedia
The Agoudim Formation (also known as the "Marnes et Marno-Calcaires d'Agoudim") is a geological formation of late Toarcian to Bajocian (Lower–Middle Jurassic[1]) age in the central High Atlas and Middle Atlas (particularly around Rich and extending to areas like Midelt, Boulemane, and Beni Mellal) of Morocco.[2][3] The formation is also known as the "Formation Marneuse d’Amane-Ilila", "Marnes de Boulemane", "Marnes de Taffert" and "Calcaires à Cancellophycus".[2][4]
- R'cifa Formation
- Tagoudite Formation
- Jbel Ouchbis Formation
- Jbel Aberdouz Formation
- Jbel Choucht Formation
| Agoudim Formation | |
|---|---|
| Stratigraphic range: Late Toarcian–Bajocian | |
Agoudim village | |
| Type | Geological Formation |
| Underlies |
|
| Overlies |
|
| Thickness | 2,000 m (6,600 ft) (up to 5,000 m (16,000 ft) reported) |
| Lithology | |
| Primary | Marl, Marly Limestone, Bioclastic limestone |
| Location | |
| Location | Er-Rich |
| Coordinates | 32.25°N 4.50°W |
| Approximate paleocoordinates | 27.0°N 3.0°W |
| Region | High Atlas and Middle Atlas |
| Country | Morocco |
| Type section | |
| Named for | Agoudim |
| Named by | M. Studer |
| Year defined | 1980 |
| Region | Er-Rich, Midelt Province |
| Thickness at type section | ~4,500–5,000 m (14,800–16,400 ft) |
Description

The Agoudim Formation is a substantial geological unit primarily composed of marls and marly limestones, representing a continuation from the underlying Tagoudite or Jbel Ouchbis formations and marking a significant shift toward more clay-rich (argillaceous) sedimentation.[2][5] This shift is periodically interrupted by layers of bioclastic material (deposits rich in shell fragments and other organic debris) that increase in both frequency and thickness moving upward through the formation. It is stratigraphically divided into two main members: a lower marly member dominated by gray-blue marls with occasional nodular bioclastic limestone beds, and an upper member characterized by more calcareous and bioclastic rocks with greater lithological variability.[6] The boundary between these members is often marked by a distinctive layer of organogenic turbidites, notably the "Calcaires à Cancellophycus", which forms a prominent brown cuesta and serves as an important marker horizon for geological mapping.[5][7]
The lower member consists of monotonous, micritic marls that reflect relatively quiet hemipelagic depositional conditions, where fine clay and carbonate mud settled along with benthic material.[2] This distal sedimentation was influenced by intermittent turbiditic currents transporting detrital quartz and glauconite, indicating periodic sediment input from nearby land areas and submarine slopes. In contrast, the upper member shows increasingly energetic conditions with cycles of sedimentation starting with lumachellic limestones (shell-rich limestones with oblique lamination) and ending in thick bioclastic marls.[5][6] This member records frequent synsedimentary slumps and spectacular deformation structures like ball-and-pillow folds (phacoids), which are indicative of tectonic instability and sediment loading on unstable slopes. The evolution upward culminates with a thick limestone cap dominated by oolitic limestones and bioherms (reef-like carbonate buildups that further highlight the transition to shallower marine conditions).[6][7]
Paleoenvironment
The Agoudim Formation records a progressive transition from deep to shallow marine settings under the influence of rifting, subsidence, and environmental perturbations in a Diapiric Rift Basin.[8][9] The lower marly members were deposited in hemipelagic conditions below storm wave base, with pelagic sedimentation, fine siliciclastic influx from nearby uplifts, and reworked carbonate from Liassic platforms.[10][11] Fossils such as ammonites, radiolarians, foraminifera, small bivalves, and echinoderm debris confirm a low-energy offshore environment. Rapid subsidence and sea-level rise during the Early Bajocian led to a drowning phase of the neritic carbonate factory, favoring marl sedimentation locally enriched in organic matter.[12] Within these successions, several intervals of authigenic carbonates developed, notably yellow micritic nodules, tubular and chimney-like concretions, and conical chemoherms, interpreted as the record of hydrocarbon Cold seep. Petrographic, isotopic, and sedimentological evidence indicates that the second and third authigenic horizons formed at active seep sites, with feeder-pipes, carbonate crusts, microbialites, and bioclast-rich infills, while the yellow nodules of the lower interval likely reflect organic matter remineralization without direct seepage.[13] Strongly depleted Δ13C values (down to −19‰) point to fluids rich in dissolved inorganic carbon generated by organic matter degradation, likely through bacterial sulfate reduction and anaerobic oxidation of hydrocarbons, rather than pure methane seepage. These processes created localized hardgrounds, serpulidae and oyster colonization, and chemoherm buildups on the seafloor.[13][14] As sedimentation continued, carbonate production recovered, and three reefal complexes developed, ranging from microbial-coral patch reefs to larger coral–sponge–algal bioherms, accompanied by ooid–peloid shoals and brachiopod-rich limestones.[15][16][17] These buildups evolved from small mounds to coalescing reefs, shaped by constructive growth, storm reworking, bioerosion, and micritization.[15][16][17] The upper part of the formation is dominated by shallow platform deposits with abundant brachiopods, gastropods, corals, and reefal frameworks.[15][17][18][19] Some areas, like the Agoudim valley record excepcionally preserved platform sections, showing it´s inner ramp had high-energy ooid–peloid shoals, microbial mounds, and small patch reefs and a proximal middle ramp with mixed coral–sponge–bryozoan–brachiopod assemblages in lower energy waters.[20] The distal ramp consisted of marl–limestone alternations with storm beds and oyster–coral–sponge buildups. Local syndepositional highs and terrigenous input created semi-restricted zones with fluctuating oxygenation and nutrients.[20]
Biota
Foranimifera
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Ammobaculites[21][22] | fontinensis | Talghemt Ridge | II-IV | Tests | Lituolidae | |
| agglutinans | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II | Tests | Lituolidae | ||
| Bullopora[13] | sp. | Jebel Bou Kendill | III | Tests | Polymorphinidae | |
| Citharina[21] | serratocostata | Talghemt Ridge | IV | Tests | Vaginulinidae | |
| Citharina[21] | sp. | Talghemt Ridge | I-II | Tests | Vaginulinidae | |
| Eoguttulina[21][22] | bilocularis | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II | Tests | Polymorphinidae | |
| sp. | Talghemt Ridge | IV | Tests | Polymorphinidae | ||
| Epistommina[22] | mosquensis | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II-IV | Tests | Epistomininae | |
| Flabellammina[21][22] | althoffi | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | III-IV | Tests | Lituolidae | |
| Garantella[22] | ampasindavaensis | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II | Tests | Epistomininae | |
| stellata | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II | Tests | Epistomininae | ||
| sp. | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | II | Tests | Epistomininae | ||
| Haplophragmium[21][22] | subaequale | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | III-IV | Tests | Lituolidae | |
| aequale | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | III-IV | Tests | Lituolidae | ||
| Ichthyolaria[21] | carinata | Talghemt Ridge | I | Tests | Ichthyolariidae | |
| gr. major | Talghemt Ridge | I | Tests | Ichthyolariidae | ||
| sp. | Talghemt Ridge | I | Tests | Ichthyolariidae | ||
| Lenticulina[21][22] | argonauta | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | IV | Tests | Nodosariidae | |
| chicheryi | Talghemt Ridge | I-II | Tests | Nodosariidae | ||
| d'orbignyi | Talghemt Ridge | I-II | Tests | Nodosariidae | ||
| münsteri | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | I-II | Tests | Nodosariidae | ||
| quenstedti | Talghemt Ridge | IV | Tests | Nodosariidae | ||
| subalata | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | I-II | Tests | Nodosariidae | ||
| toarcense | Talghemt Ridge | I | Tests | Nodosariidae | ||
| polymorpha | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | III-IV | Tests | Nodosariidae | ||
| Parurgonina[22] | sp. | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | III-IV | Tests | Parurgoninidae | |
| Planularia[21] | sp. | Talghemt Ridge | IV | Tests | Vaginulinidae | |
| Proteonina[21] | gr. difflugiformis | Talghemt Ridge | IV | Tests | Hormosinidae | |
| Protoglobigerina?[21] | spp. | Talghemt Ridge | I-III | Tests | Globigerinidae | |
| Pseudonodosaria[21] | gr. pygmea | Talghemt Ridge | I | Tests | Nodosariidae | |
| multicostata | Talghemt Ridge | I | Tests | Nodosariidae | ||
| Reinholdella[22] | dreheri | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | III-IV | Tests | Ceratobuliminidae | |
| Spirillina[21] | orbicula | Talghemt Ridge | I-II | Tests | Spirillinidae | |
| Textularia[21] | jurassica | Talghemt Ridge | IV | Tests | Textulariidae | |
| Trochammina[21] | globigeriniformis | Talghemt Ridge | IV | Tests | Trochamminidae | |
| Verneuilinoides[21] | cf. subvitreus | Talghemt Ridge | I-II | Tests | Verneuilinidae |
Ichnotaxa
| Ichnogenus | Ichnospecies | Locality | Member | Made by | Images |
|---|---|---|---|---|---|
| Chondrites[23] | isp. | Assameur, Jebel Taferdout | III-IV | ||
| Planolites[23] | isp. | Assameur, Jebel Taferdout | III-IV |
|
|
| Phycodes[23] | isp. | Assameur, Jebel Taferdout | III-IV | ||
| Thalassinoides[13][24] | isp. | Jebel Bou Kendill | III | ||
| Trypanites[23] | isp. | Assameur, Jebel Taferdout | III-IV | ||
| Zoophycos[25] | (Cancellophycus) ispp. | Amellago; Talghemt Ridge; Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | I-IV | ||
Brachiopoda
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Acanthothiris[26] | oligocantha | Rich (syncline of Jebel Bou Kendill) | II | Shells | Athyrididae | |
| tenuispina | Rich (syncline of Jebel Bou Kendill) | III | Shells | Athyrididae | ||
| Cymatorhynchia[27] | quadriplicata | Jebel Assameur; Boukendil; Rich-Gourrama area | IV | Shells | Tetrarhynchiidae | |
| Kallirhynchia[28] | concinna | Amellago, Talghemt Ridge | I | Shells | Rhynchonellidae | |
| Loboidothyris[26] | sp. | Rich (syncline of Jebel Bou Kendill) | II | Shells | Loboidothyrididae | |
| Lophrothyris[26] | contracta | Rich (syncline of Jebel Bou Kendill) | II | Shells | Terebratulidae | |
| Monsardithyris[17][26] | cortonensis | Rich (syncline of Jebel Bou Kendill) | III | Shells | Terebratulidae | |
| ventricosa | Assameur n’Ait Fergane | IV | Shells | Terebratulidae | ||
| sp. cf. M. loubensis | Bou-Ouchène (northern platform); Rich | III | Shells | Terebratulidae | ||
| Quadratirhynchia[28] | vasconcellosi | Amellago | I | Shells | Tetrarhynchiidae | |
| Rhactorhynchia[26] | distracta | Rich (syncline of Jebel Bou Kendill) | III | Shells | Rhynchonellidae | |
| Rhynchonelloidea[26] | goyi | Amellago | I | Shells | Rhynchonellidae | |
| Rugitela[26] | cf. hughesi | Rich (syncline of Jebel Bou Kendill) | II–III | Shells | Zeilleriidae | |
| Zeilleria[17][26] | truncatella | Rich (syncline of Jebel Bou Kendill) | III | Shells | Zeilleriidae | |
| perlata | Assameur n’Ait Fergane, Boukendil | IV | Shells | Zeilleriidae |
Bryozoa
| Genus | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Berenicea[17] | sp. | Boukendil, Assameur | IV | Colonies | Bereniceidae | |
| Collapora[17] | cf. straminea | Boukendil | IV | Colonies | Atactotoechidae | |
| Entalophora[17] | cf. stokesi | Boukendil | IV | Colonies | Entalophoridae |
Cnidaria
| Genus | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Acropora[17] | cf.sp. | Assameur n’Ait Fergane, Boukendil | IV | Colonies | Acroporidae | |
| Aggomorphastraea[29] | "sp. 1" | Amellago | I | Colonies | Latomeandridae | |
| Allocoenia[29] | "sp. 1" | Amellago | I | Colonies | Actinastreidae | |
| "sp. 2" | Amellago | I | Colonies | Actinastreidae | ||
| Ampakabastrea[29] | ampakabensis | Amellago | I | Colonies | Astrocoeniidae | |
| Cladophyllia[9][17] | sp. | Assameur n’Ait Fergane; Imilchil | I-IV | Colonies | Caryophylliidae | |
| Comophyllia[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Comophylliidae | |
| Complexastrea[29] | "sp. 1" | Amellago | I | Colonies | Thecosmiliidae | |
| Cyathocoenia[29] | faugeresi | Amellago | I | Colonies | Actinastreidae | |
| "sp. 1" | Amellago | I | Colonies | Actinastreidae | ||
| Dendrarea[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Scleractinia (family uncertain) | |
| Dasmosmilia[29] | "sp. 1" | Amellago | I | Colonies | Dermosmiliidae | |
| Dimorpharaea[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Scleractinia (family uncertain) | |
| Edwardoseris[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Scleractinia (family uncertain) | |
| Enallocoenia[29] | "sp. 1" | Amellago | I | Colonies | Actinastreidae | |
| Isastrea[17][29] | sp. | Assameur n’Ait Fergane | IV | Colonies | Latomeandridae/Montlivaltiidae | |
| "sp. 1" | Amellago | I | Colonies | Latomeandridae | ||
| Kobymeandra[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Scleractinia (family uncertain) | |
| Latiastrea[29] | "sp. 1" | Amellago | I | Colonies | Latomeandridae | |
| Lochmaeosmilia[9] | sp. | Imilchil | I | Colonies | Calamophylliidae | |
| Microphyllia[29] | irregularis | Amellago | I | Colonies | Latomeandridae | |
| Montlivaltia[17] | sp. | Assameur n’Ait Fergane | IV | Coralla | Montlivaltiidae (solitary) | |
| Polystylidium[17] | sp. | Assameur n’Ait Fergane | IV | Colonies | Scleractinia (family uncertain) | |
| Stylosmilia[17][29] | sp. | Foum Zabel (southern margin) | II-III | Colonies | Thecosmiliidae | |
| "sp. 1" | Amellago | I | Colonies | Thecosmiliidae | ||
| Thecocyathus[29] | mactrus | Amellago | I | Solitary corallites | Caryophylliidae | |
| Vallimeandropsis[29] | lineata | Amellago | I | Colonies | Scleractinia (family uncertain) |
Porifera
| Genus | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Cheilosporites[23] | tirolensis | Assameur, Jebel Taferdout | IV | Sponge body fossils | Vaceletidan | |
| spp. | Assameur, Jebel Taferdout | IV | Sponge body fossils | |||
| Cladocoropsis[17] | spp. | Boukendil, Assameur | IV | Sponge body fossils | Axinellidan | |
| Eudea[17] | cf. globata | Boukendil, Assameur | IV | Sponge body fossils | Stellispongiida | |
| Neuropora[9] | spp. | Imilchil | I | Sponge body fossils | Neuroporidae |
Mollusks
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Actinostreon[13][16] | sp. | Assameur n’Ait Fergane, Boukendil | IV | Shells | Gryphaeidae | |
| Alocolytoceras[1][30] | ophioneum | Gheris Valley | I | Shells | Lytoceratidae | |
| dorcadis | Ait El Abbas; Azag | I | Shells | |||
| sp. | Azag | I | Shells | |||
| Ambersites[1] | sp. | Rich region | II–III | Shells | Hammatoceratidae | |
| Amphitrochus[31] | abbas | Rich region; Outarbat | III-IV | Shells | Nododelphinulidae | |
| Ancolioceras[32] | opalinoides | n’Zala (Jbel Aouja) | I–II | Shells | Graphoceratidae | |
| Asthenoceras[1] | sp. | Rich (Concavum–Discites zones) | II–III | Shells | Sonniniidae | |
| Belemnopsis[33] | baculiformis | Iwansitn | III | Rostra | Belemnopseidae | |
| pougatchevskae | Iwansitn | III | Rostra | |||
| Bositra[1] | buchi | Amellago; Rich | I–III | Shells | Posidoniidae | |
| Botula[16] | sp. | Boukendil area | IV | Shells, borings | Mytilidae | |
| Bradfordia[1] | costata | Rich region | III | Shells | Oppeliidae | |
| sp. | Rich region | III | Shells | |||
| Calabribelus[33] | combemoreli | Rich (Jbel Bou Kandil), Iwansitn (Middle Atlas) | II–III | Rostra | Holcobelidae | |
| tetramerus | Rich, Iwansitn | II–III | Rostra | |||
| Calliphylloceras[30][34] | nilssoni | Amellago | I | Shells | Phylloceratidae | |
| cf.capitanii | Azag | I-II | Shells | |||
| sp. | Azag | I-II | Shells | |||
| Catacoeloceras[34] | sp. | Ait El Abbas | I | Shells | Dactylioceratidae | |
| Catulloceras[1] | sp. | n’Zala (Jbel Aouja) | I | Shells | Graphoceratidae | |
| Chlamys[1][9] | sp. | Rich, Amellago; Imilchil | I-II–III | Shells | Pectinidae | |
| Cirrus[31] | sp. | Aberdouz | III-IV | Shells | Cirridae | |
| Conobelemnopsis[33] | sp. A | Rich, Iwansitn | III | Rostra | Belemnopseidae | |
| Collina[30] | gemma | Ait El Abbas; Azag | I | Shells | Dactylioceratidae | |
| cf.nummularia | Ait El Abbas | I | Shells | |||
| linae | Ait El Abbas | I | Shells | |||
| sp. | Azag | I | Shells | |||
| Cotteswoldia[32] | sp. | n’Zala (Jbel Aouja) | I | Shells | Graphoceratidae | |
| Crassiceras[30] | cf.latum | Ait El Abbas | I | Shells | Hildoceratidae | |
| Ctenostreon[1] | sp. | Rich, Amellago | II–III | Shells | Limidae | |
| Cypholioceras[32] | sp. | n’Zala (Jbel Aouja) | II | Shells | Graphoceratidae | |
| Dactylioceras[21][30][34] | (Eodactylites) mirabile | Amellago | I | Shells | Dactylioceratidae | |
| (Eodactylites) pseudocommune | Amellago | I | Shells | |||
| (Eodactylites) simplex | Talghemt Ridge | I | Shells | |||
| (Orthodactylites) aff. crosbeyi | Amellago | I | Shells | |||
| (Orthodactylites) cf. semiannulatum | Amellago | I | Shells | |||
| (Orthodactylites) andaluciensis | Agounou | I | Shells | |||
| sp. | Ait El Abbas | I | Shells | |||
| ?Dicoelites[33] | sp. A | Iwansitn | III | Rostra | Dicoelitidae | |
| Discohelix[31] | subaequalis | Rich region; Outarbat | III-IV | Shells | Discohelicidae | |
| pulchrior | Rich region | IV | Shells | |||
| sp. | Rich region | IV | Shells | |||
| Docidoceras[1] | wysogorskii | Rich | III | Shells | Otoitidae | |
| Eleganticeras[34] | exaratum | Amellago | I | Shells | Dactylioceratidae | |
| sp. | Amellago | I | Shells | |||
| Emileia[35] | (Emileia) polymera | Boukendil, Assameur | IV | Shells | Otoitidae | |
| Eucyclus[31] | ornatus | Rich region | IV | Shells | Eucyclidae | |
| sp. | Aberdouz | III | Shells | |||
| Eucycloidea[31] | (Pseudalaria?) subangulata | Rich region | IV | Shells | Eucyclidae | |
| cf. etheridgii | Rich region; Outarbat | III-IV | Shells | |||
| Eudmetoceras[1] | flexuosum | Rich | II–III | Shells | Hammatoceratidae | |
| masticonense | Rich | III | Shells | |||
| eudmetum | Rich | III | Shells | |||
| sp. | Rich | II–III | Shells | |||
| Euaptetoceras[1] | amaltheiforme | Rich | III | Shells | Hammatoceratidae | |
| amplectens | Rich | III | Shells | |||
| sp. | Rich | II–III | Shells | |||
| Fontannesia[1] | aurita | Rich | II | Shells | Sonniniidae | |
| montillanensis | Rich | II | Shells | |||
| sp. | Rich | II–III | Shells | |||
| Furloceras[30] | cf.chelussii | Ait El Abbas | I | Shells | Phymatoceratidae | |
| Geczyceras[30] | clausum | Ait El Abbas | I | Shells | Hammatoceratidae | |
| speciosum | Azag | I | Shells | |||
| Grammatodon[1] | sp. | Rich | II–III | Shells | Parallelodontidae | |
| Hammatoceras[34] | H. aff. insigne | n’Zala (Jbel Aouja) | I | Shells | Hammatoceratidae | |
| Harpoceras[30][34] | falciferum | Amellago | I | Shells | Hildoceratidae | |
| pseudoserpentinum | Amellago | I | Shells | |||
| serpentinum | Amellago | I | Shells | |||
| subplanatum | Amellago | I | Shells | |||
| cf.mediterraneum | Azag | I | Shells | |||
| Hildaites[30][34] | forte | Amellago | I | Shells | Hildoceratidae | |
| levisoni | Amellago | I | Shells | |||
| cf. serpentiniformis | Amellago | I | Shells | |||
| cf. subserpentinus | Amellago; Agounou | I | Shells | |||
| striatus | Amellago | I | Shells | |||
| wrighti | Amellago | I | Shells | |||
| (Martanites) cf. prorsiradiatus | Agounou | I | Shells | |||
| murleyi | Agounou | I | Shells | |||
| aff. murleyi | Ait El Abbes | I | Shells | |||
| cf. striatus | Agounou | I | Shells | |||
| undicosta | Agounou | I | Shells | |||
| Hildoceras[21][30][34] | bifrons | Amellago; Azag; Talghemt Ridge | I | Shells | Hildoceratidae | |
| lusitanicum | Amellago; Talghemt Ridge | I | Shells | |||
| semipolitum | Amellago; Talghemt Ridge | I | Shells | |||
| sublevisoni | Amellago; Azag; Talghemt Ridge | I | Shells | |||
| bonarellii | Talghemt Ridge | I–II | Shells | |||
| cf.tethysi | Azag | I | Shells | |||
| appenninicus | Agounou; Azag | I | Shells | |||
| snoussi | Ait El Abbes | I | Shells | |||
| cf.apertum | Azag | I | Shells | |||
| sp. | Amellago; Azag | I | Shells | |||
| Hyperlioceras[1] | walkeri | Rich | III | Shells | Graphoceratidae | |
| incisum | Rich | III | Shells | |||
| inclusum | Rich | III | Shells | |||
| mundum | Rich | III | Shells | |||
| sp. | Rich | III | Shells | |||
| Leioceras[32] | cf. comptum | n’Zala (Jbel Aouja) | II | Shells | Graphoceratidae | |
| costusum | n’Zala (Jbel Aouja) | II | Shells | |||
| Lithophaga[17] | sp. | Boukendil area | IV | Shells, borings | Mytilidae | |
| Ludwigia[32] | aff. haugi | n’Zala (Jbel Aouja) | II | Shells | Graphoceratidae | |
| Lytoceras[30][34][36] | siemensi | Amellago; n’Zala | I | Shells | Lytoceratidae | |
| cf.siemensi | Agounou; Ait El Abbes | I-II | Shells | |||
| subfimbriatum | n’Zala | II | Shells | |||
| francisci | Ait El Abbes | I-II | Shells | |||
| sp. | Amellago; Agounou; Ait El Abbes; n’Zala | I–II | Shells | |||
| Maconiceras[34] | soloniacense | Amellago | I | Shells | Hammatoceratidae | |
| Malladaites[1] | pertinax | Rich | III | Shells | Hammatoceratidae | |
| Merlaites[30] | alticarinatus | Ait El Abbes; Azag | I | Shells | Hildoceratidae | |
| gradatus | Ait El Abbes | I | Shells | |||
| Neotelodactylites[30] | eucosmus | Ait El Abbes | I | Shells | Dactylioceratidae | |
| Nerinea[17] | sp. | Assameur n’Ait Fergane | IV | Shells | Nerineidae | |
| Nicaniella[1] | sp. | Rich, Amellago | II–III | Shells | Astartidae | |
| Nodicoeloceras[30] | cf. merlai | Ait El Abbes | I | Shells | Dactylioceratidae | |
| Osperleioceras[30] | bicarinatum | Ait El Abbes; Azag | I | Shells | Hildoceratidae | |
| beauliziense | Azag | I | Shells | |||
| cf. authelini | Azag | I | Shells | |||
| cf. reynesi | Azag | I | Shells | |||
| O. (Pseudolioceras) sp. | Ait El Abbes | I | Shells | |||
| sp. | Azag | I | Shells | |||
| Palaeonucula[1] | hammeri | Rich, Amellago | II–III | Shells | Nuculidae | |
| Palaeoneilo[1] | sp. | Rich | III | Shells | Malletiidae | |
| Peronoceras[30] | cf. subarmatum | Ait El Abbes | I | Shells | Dactylioceratidae | |
| Phylloceras[30][34] | doderleinianum | Ait El Abbes | II | Shells | Phylloceratidae | |
| sp. | Amellago; n’Zala | I–II | Shells | |||
| Phymatoceras[30] | sp. | Azag | I | Shells | Phymatoceratidae | |
| Pholadomya[27] | (Bucardiomya) bucardium | Jebel Assameur; Boukendil; Rich-Gourrama area | IV | Shells | Pholadomyidae | |
| Planammatoceras[32] | sp. | n’Zala (Jbel Aouja) | II | Shells | Hammatoceratidae | |
| Platystrophites[30] | cf. latus | Agounou | I | Shells | Dactylioceratidae | |
| Pleurotomaria[31] | armata | Rich region | IV | Shells | Pleurotomariidae | |
| Pleydellia[32] | aalensis | n’Zala (Jbel Aouja) | II | Shells | Graphoceratidae | |
| Polyplectus[34] | pluricostatum | Amellago | I | Shells | Hammatoceratidae | |
| sp. | Amellago | I | Shells | |||
| Porpoceras[30][34] | gr. vortex-verticosum | Amellago; Talghemt Ridge | I | Shells | Dactylioceratidae | |
| cf. vortex | Ait El Abbes | I | Shells | |||
| cf. vorticellum | Ait El Abbes | I | Shells | |||
| cf. verticosum | Ait El Abbes | I | Shells | |||
| sp. | Ait El Abbes; Agounou | I | Shells | |||
| Praestrigites[1] | sp. | Rich | III | Shells | Strigoceratidae | |
| Praeoppelia[1] | sp. | Rich | III | Shells | Oppeliidae | |
| Protetragonites[36] | sp. | n’Zala | II | Shells | Lytoceratidae | |
| Protoecotraustes[1] | sp. | Rich | III | Shells | Haploceratidae | |
| Pseudocrassiceras[30] | aff.bayani | Ait El Abbes; Azag | I | Shells | Hildoceratidae | |
| bayani | Ait El Abbes | I | Shells | |||
| frantzi | Ait El Abbes | I | Shells | |||
| praegruneri | Ait El Abbes | I | Shells | |||
| cf. frantzi | Azag | I | Shells | |||
| sp. | Azag | I | Shells | |||
| Pseudogrammoceras[30] | praegruneri | Azag | I | Shells | Hildoceratidae | |
| subregale | Ait El Abbes; Azag | I | Shells | |||
| aff. fallaciosum | Azag | I | Shells | |||
| Rastellum[17] | sp. | Assameur n’Ait Fergane, Boukendil | IV | Shells | Ostreidae | |
| Rhynchocerithium[31] | sp. | Rich region | IV | Shells | Procerithiidae | |
| Riselloidea[31] | sp. | Outarbat | III-IV | Shells | Eucyclidae | |
| Sadkia[31] | richensis | Rich region | IV | Shells | Eucyclidae | |
| Saxicava[17] | sp. | Boukendil area | IV | Shells, borings | Hiatellidae | |
| Sonninia[1] | dominans | Rich | III | Shells | Sonniniidae | |
| modestum | Rich | III | Shells | |||
| nannomorpha | Rich | III | Shells | |||
| Tmetoceras[32] | T. scissum | n’Zala (Jbel Aouja) | I–II | Shells | Hammatoceratidae | |
| Toxamblyites[35] | densicostatus | Boukendil, Assameur | IV | Shells | Haploceratidae | |
| Trigonia[1] | sp. | Rich | III | Shells | Trigoniidae | |
| Trilobiticeras[1] | platygaster | Rich | III | Shells | Otoitidae | |
| trilobitoides | Rich | III | Shells | |||
| sp. | Rich | III | Shells | |||
| Tylostoma[1] | globosum | Rich | III | Shells | Tylostomatidae | |
| Vacekia[32] | sp. | n’Zala (Jbel Aouja) | I–II | Shells | Graphoceratidae | |
| Zugodactylites[30] | sp. | Ait El Abbes | I | Shells | Dactylioceratidae | |
| Zurcheria[1] | sp. | Rich | III | Shells | Hammatoceratidae |
Arthropods
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Bairdia[21] | sp. A | Talghemt Ridge | I–II | Valves | Bairdiidae | |
| aff. hilda | Talghemt Ridge | IV | Valves | Bairdiidae | ||
| Bairdiacypris[21][22] | sp. | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | I–II | Valves | Bairdiacyprididae | |
| Cytherella[21][22] | aff. concentrica | Talghemt Ridge | IV | Valves | Cytherellidae | |
| aff. fullonica | Talghemt Ridge | IV | Valves | Cytherellidae | ||
| aff. toarcensis | Talghemt Ridge | I–II | Valves | Cytherellidae | ||
| spp. | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | II | Valves | Cytherellidae | ||
| Cytherelloidea[21] | gr. catenulata | Talghemt Ridge | IV | Valves | Cytherelloididae | |
| Favreina[23] | isp. | Assameur, Jebel Taferdout | III-IV | Coprolites | Decapod Coprolites (Axiidea?) | |
| Kinkelinella[21] | K. sermoisensis | Talghemt Ridge | I–II | Valves | Kinkelinellidae | |
| gr. sermoisensis | Talghemt Ridge | I–II | Valves | Kinkelinellidae | ||
| Micropneumatocythere[22] | sp. | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | II | Valves | Podocopida | |
| Monoceratina[21] | aff. ventrale | Talghemt Ridge | IV | Valves | Monoceratidae | |
| Paracypris[21] | sp. | Talghemt Ridge | IV | Valves | Cytheridae | |
| Polycope[21] | sp. | Talghemt Ridge | IV | Valves | Polycopidae | |
| Pontocyprella[21] | sp. | Talghemt Ridge | I–II | Valves | Cyprididae | |
| Procytheridea[21][22] | sp. B | Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge | II-IV | Valves | Cytherideidae | |
| Striatojonesia[21] | S. striata | Talghemt Ridge | IV | Valves | Bythocytheridae |
Echinoderms
| Genus | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Cidaris[27] | sp. | Jebel Assameur; Boukendil; Rich-Gourrama area | IV | Spines & Specimens | Cidaridae | |
| Diplopodia[22][27] | cf. morierii | Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | IV | Spines & Specimens | Diplopodiidae | |
| Gymnocidaris[27] | pustulosa | Jebel Assameur; Boukendil; Rich-Gourrama area | IV | Spines & Specimens | Hemicidaridae | |
| Hemicidaris[22][27] | koechlini | Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | IV | Spines & Specimens | Hemicidaridae | |
| Psephechinus[22][27] | globosus | Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi | IV | Spines & Specimens | Stomechinidae |
Annelida
| Genus | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Nogrobs[35] | moroccensis | Boukendil, Assameur | IV | Quadrangular tubes | Serpulidae | |
| Serpula[9][13][17] | socialis | Assameur n’Ait Fergane, Boukendil; Imilchil; Jebel Bou Kendill (cap bed) | I-III–IV | Tubes / colonies | Serpulidae | |
| sp. | Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) | III–IV | Tubes / colonies | Serpulidae | ||
| Terebella[17] | lapilloides | Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) | III–IV | Tubes / colonies | Terebellidae |
Chondrichthyes
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Strophodus[37] | atlasensis | Talsint (eastern High Atlas) | III–IV | Articulated dentition (holotype PIMUZ A/I 5181) | Acrodontidae |
Dinoflajellates
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Kallosphaeridium[38] | sp. | Amellago | I-II | Cysts | Incertade Sedis | |
| Luehndea[38] | cirilliae | Amellago | I | Cysts | Luehndeoideae | |
| spinosa | Amellago | I | Cysts | Luehndeoideae | ||
| Mancodinium[38] | semitabulatum | Amellago | I-III | Cysts | Mancodinioideae | |
| Mendicodinium[38] | microscabratum | Amellago | I-II | Cysts | Gonyaulacales | |
| sp. 1 | Amellago | I-II | Cysts | Gonyaulacales | ||
| Moesiodinium[38] | sp. | Amellago | I-II | Cysts | Heterocapsaceae | |
| Phallocysta[38] | sp. | Amellago | I-II | Cysts | Peridiniphycideae | |
| Wallodinium[38] | cf. cylindricum | Amellago | I-II | Cysts | Incertade Sedis |
Algae
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Biscutum[10][39][40] | depravatum | Amellago; Talghemt Ridge | II–III | Coccoliths | Biscutaceae | |
| grande | Amellago; Talghemt Ridge | II–III | Coccoliths | Biscutaceae | ||
| Botryococcus[38] | sp. | Amellago | I-III | Miospore | Botryococcaceae | |
| Cayeuxia[17][18] | sp. | Assameur n’Ait Fergane; Assoul ramp | IV | Filamentous thalli | Cyanobacteria | |
| Carinolithus[10][39][40] | superbus | Amellago; Talghemt Ridge | II–III | Coccoliths | Carinolithaceae | |
| Cylindroporella[17] | arabica | Assameur n’Ait Fergane | IV | Thalli | Dasycladaceae | |
| Discorhabdus[10][39][40] | ignotus | Amellago; Talghemt Ridge | II–III | Coccoliths | Discorhabdaceae | |
| striatus | Amellago; Talghemt Ridge | II–III | Coccoliths | Discorhabdaceae | ||
| Garwoodia[20] | sp. | Amellago canyon | IV | Filamentous thalli | Cyanobacteria | |
| Girvanella[17] | sp. | Assameur n’Ait Fergane | IV | Filamentous thalli | Cyanobacteria | |
| Halimeda[17] | sp. (cf.) | Assameur n’Ait Fergane | IV | Thallus fragments | Halimedaceae | |
| Lotharingius[10][39][40] | hauffii | Amellago; Talghemt Ridge | II–III | Coccoliths | Lotharingiaceae | |
| Mitrolithus[10][39][40] | jansae | Amellago; Talghemt Ridge | II–III | Coccoliths | Mitrolithaceae | |
| Permocalculus[17] | sp. | Assameur n’Ait Fergane, Boukendil | IV | Encrusting thalli | Gymnocodiaceae | |
| Rivularia[20] | sp. | Amellago canyon | IV | Filamentous thalli | Rivulariaceae | |
| Sarfatiella[17] | dubari | Assameur n’Ait Fergane | IV | Thalli | Dasycladaceae | |
| Schizosphaerella[10][39][40] | sp. | Amellago; Talghemt Ridge | II–III | Coccoliths | Schizosphaerellaceae | |
| Sellinoporella[17] | sp. | Assameur n’Ait Fergane | IV | Thalli | Dasycladaceae | |
| Similiscutum[10][39][40] | finchii | Amellago; Talghemt Ridge | II–III | Coccoliths | Biscutaceae | |
| novum | Amellago; Talghemt Ridge | II–III | Coccoliths | Biscutaceae | ||
| Solenopora[17] | sp. (cf.) | Assameur n’Ait Fergane | IV | Encrusting thalli | Solenoporaceae | |
| Tasmanites[38] | sp. | Amellago | I-III | Miospore | Tasmanaceae | |
| Triscutum[10][39][40] | sullivanii | Amellago; Talghemt Ridge | II–III | Coccoliths | Biscutaceae | |
| Watznaueria[10][39][40] | colacicchii | Amellago; Talghemt Ridge | III | Coccoliths | Watznaueriaceae | |
| fossacincta | Amellago; Talghemt Ridge | III | Coccoliths | Watznaueriaceae |
Plants
| Taxa | Species | Locality | Member | Material | Notes | Images |
|---|---|---|---|---|---|---|
| Alisporites[38] | spp. | Amellago | I | Pollen | Corystospermaceae | |
| Callialasporites[38] | spp. | Amellago | I-II | Pollen | Araucariaceae | |
| Classopollis[38] | sp. | Amellago | I | Pollen | Hirmeriellaceae | |
| Cyathidites[38] | spp. | Amellago | I | Spores | Cyatheaceae | |
| Deltoidospora[38] | spp. | Amellago | I | Spores | Pteridophyte | |
| Kraeuselisporites[38] | reissingeri | Amellago | I | Spores | Lycopodiaceae | |
| Ischyosporites[38] | variegatus | Amellago | I | Spores | Lycopodiaceae | |
| Neoraistrickia[38] | spp. | Amellago | I-II | Spores | Selaginellaceae | |
| Quadraeculina[38] | spp. | Amellago | I | Pollen | Podocarpaceae | |
| Zonalapollenites[38] | spp. | Amellago | I-II | Pollen | Araucariaceae |
See also
- Toarcian turnover
- Toarcian formations
- Tafraout Formation, SISTER UNIT, Morocco
- Tagoudite Formation, SISTER UNIT, Morocco
- Azilal Formation, SISTER UNIT, Morocco
- Aganane Formation, Morocco
- Calcaires du Bou Dahar, Morocco
- Marne di Monte Serrone, Italy
- Podpeč Limestone, Slovenia
- El Pedregal Formation, Spain
- Mizur Formation, North Caucasus
- Sachrang Formation, Austria
- Posidonia Shale, Lagerstätte in Germany
- Irlbach Sandstone, Germany
- Ciechocinek Formation, Germany and Poland
- Krempachy Marl Formation, Poland and Slovakia
- Djupadal Formation, Central Skane
- Lava Formation, Lithuania
- Whitby Mudstone, England
- Fernie Formation, Alberta and British Columbia
- Whiteaves Formation, British Columbia
- Navajo Sandstone, Utah
- Los Molles Formation, Argentina
- Mawson Formation, Antarctica
- Kandreho Formation, Madagascar
- Kota Formation, India
- Cattamarra Coal Measures, Australia























