Agraphydrus
From Wikipedia, the free encyclopedia
Genus of beetles
| Agraphydrus | |
|---|---|
 | |
| Dorsal view of Agraphydrus sp. | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | Polyphaga |
| Infraorder: | Staphyliniformia |
| Family: | Hydrophilidae |
| Subfamily: | Acidocerinae |
| Genus: | Agraphydrus Régimbart, 1903 |
| Diversity | |
| 205 species | |
| Synonyms | |
| |
Agraphydrus is a genus of water scavenger beetle in the family Hydrophilidae represented by 205 recognized species.[1][2] It is distributed across the Afrotropical, Australasian, and Indomalayan realms.
Description
Small beetles (1.4–4.8 mm), pale/yellowish to dark brown in coloration, eyes not emarginate, labrum exposed, maxillary palps moderately long. Elytra without sutural striae, not laterally explanate; the elytral punctation ranges from very fine and subtle to coarse.
A complete diagnosis was presented by Girón and Short,[2] but also see Komarek and collaborators.[4][5][6][7][8]
Habitat
According to Girón and Short,[2] based on annotations by Komarek and collaborators:[4][5][6][7][8]
Agraphydrus can be found in an extremely broad range of habitats, from rivers, streams and forest pools, to hygropetric environments around waterfalls or seepages over rocks; a few species have been collected in terrestrial habitats by sifting moss and leaves from near water bodies, or in the gravel along the bank of a river; in many cases specimens have been found associated with floating vegetation, mosses and algae.[2]
Species
- Agraphydrus abrasus Komarek and Freitag, 2020 [7]
- Agraphydrus activus Komarek and Hebauer, 2018 [8]
- Agraphydrus acutus Komarek, 2020 [6]
- Agraphydrus aethiopicus Komarek, 2020 [6]
- Agraphydrus agilis Komarek and Hebauer, 2018 [8]
- Agraphydrus albescens (Régimbart, 1903) [9]
- Agraphydrus ampullatus Komarek and Freitag, 2020 [7]
- Agraphydrus anacaenoides Komarek, 2019 [5]
- Agraphydrus anatinus Komarek, 2018 [4]
- Agraphydrus andamanicus Komarek, 2018 [4]
- Agraphydrus andringitra Komarek, 2020 [6]
- Agraphydrus angulatus Komarek, 2019 [5]
- Agraphydrus angustatus Komarek, 2020 [6]
- Agraphydrus angustipenis Komarek, 2018 [4]
- Agraphydrus anhuianus (Hebauer, 2000) [10]
- Agraphydrus annapurnensis Komarek, 2018 [4]
- Agraphydrus arduus Komarek and Hebauer, 2018 [8]
- Agraphydrus ater Komarek, 2018 [4]
- Agraphydrus atripalpis Komarek, 2020 [6]
- Agraphydrus attenuatus (Hansen, 1999) [11]
- Agraphydrus audax Komarek and Hebauer, 2018 [8]
- Agraphydrus avita (Hansen, 1997) [12]
- Agraphydrus bacchusi Komarek, 2019 [5]
- Agraphydrus balkeorum Komarek, 2019 [5]
- Agraphydrus batak Komarek and Freitag, 2020 [7]
- Agraphydrus bhutanensis Komarek, 2018 [4]
- Agraphydrus bicoloratus Komarek, 2020 [6]
- Agraphydrus bilardoi Komarek, 2020 [6]
- Agraphydrus biltoni Komarek, 2020 [6]
- Agraphydrus biprojectus Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus borneensis Komarek, 2019 [5]
- Agraphydrus boukali Komarek, 2018 [4]
- Agraphydrus brevilobatus Komarek and Freitag, 2020 [7]
- Agraphydrus brevipenis Komarek, 2019 [5]
- Agraphydrus burmensis Komarek, 2019 [5]
- Agraphydrus calvus Komarek and Hebauer, 2018 [8]
- Agraphydrus camerunensis Komarek, 2020 [6]
- Agraphydrus cantonensis Komarek and Hebauer, 2018 [8]
- Agraphydrus carinatulus Komarek, 2019 [5]
- Agraphydrus cervus Komarek, 2019 [5]
- Agraphydrus ceylonensis Komarek, 2018 [4]
- Agraphydrus chinensis Komarek and Hebauer, 2018 [8]
- Agraphydrus cinnamum Komarek, 2018 [4]
- Agraphydrus clarus Komarek, 2019 [5]
- Agraphydrus comes Komarek and Hebauer, 2018 [8]
- Agraphydrus communis Komarek, 2018 [4]
- Agraphydrus confusus Komarek and Hebauer, 2018 [8]
- Agraphydrus congolensis Komarek, 2020 [6]
- Agraphydrus conicus Komarek and Hebauer, 2018 [8]
- Agraphydrus connexus Komarek and Hebauer, 2018 [8]
- Agraphydrus constrictus Komarek, 2018 [4]
- Agraphydrus contractus Komarek and Hebauer, 2018 [8]
- Agraphydrus coomani (d'Orchymont, 1927) [14]
- Agraphydrus coronarius Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus crassipenis Komarek, 2018 [4]
- Agraphydrus dapengensis Yang & Jia, 2021 [15]
- Agraphydrus decipiens Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus delineatus Komarek, 2019 [5]
- Agraphydrus elongatus Ribera, Hernando, and Cieslak, 2019 [16]
- Agraphydrus engkari Komarek, 2019 [5]
- Agraphydrus excisus Komarek, 2019 [5]
- Agraphydrus exedis (d'Orchymont, 1937) [17]
- Agraphydrus exiguus Komarek, 2019 [5]
- Agraphydrus falcatus Komarek, 2018 [4]
- Agraphydrus fasciatus Komarek and Hebauer, 2018 [8]
- Agraphydrus fikaceki Komarek and Hebauer, 2018 [8]
- Agraphydrus flavescens Komarek, 2020 [6]
- Agraphydrus flavipes Komarek, 2020 [6]
- Agraphydrus flavonotus Komarek, 2018 [4]
- Agraphydrus floresinus Komarek, 2019 [5]
- Agraphydrus fontis Komarek, 2020 [6]
- Agraphydrus forcipatus Komarek and Hebauer, 2018 [8]
- Agraphydrus fortis Komarek, 2018 [4]
- Agraphydrus fujianensis Komarek and Hebauer, 2018 [8]
- Agraphydrus geminus (d'Orchymont, 1932) [18]
- Agraphydrus gereckei Komarek, 2020 [6]
- Agraphydrus gilvus Komarek, 2018 [4]
- Agraphydrus glaber Komarek, 2018 [4]
- Agraphydrus globipenis Komarek and Hebauer, 2018 [8]
- Agraphydrus goldschmidti Komarek, 2020 [6]
- Agraphydrus gracilipalpis Komarek, 2018 [4]
- Agraphydrus hamatus Komarek, 2019 [5]
- Agraphydrus hanseni (Satô and Yoshitomi, 2004) [19]
- Agraphydrus heinrichi Komarek, 2018 [4]
- Agraphydrus helicopter Komarek, 2019 [5]
- Agraphydrus hendrichi Komarek, 2019 [5]
- Agraphydrus heterochromatus Komarek, 2019 [5]
- Agraphydrus hortensis Komarek, 2019 [5]
- Agraphydrus hygropetricus Komarek, 2018 [4]
- Agraphydrus igneus Komarek and Hebauer, 2018 [8]
- Agraphydrus imitans Komarek, 2019 [5]
- Agraphydrus indicus (d'Orchymont, 1932) [18]
- Agraphydrus inflatus Komarek, 2018 [4]
- Agraphydrus infuscatus Komarek, 2019 [5]
- Agraphydrus insidiator Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus ishiharai (Matsui, 1994) [20]
- Agraphydrus jaechi (Hansen, 1999) [11]
- Agraphydrus jankodadai Komarek, 2019 [5]
- Agraphydrus jilanzhui Komarek and Hebauer, 2018 [8]
- Agraphydrus kallar Komarek, 2018 [4]
- Agraphydrus kathapa Komarek, 2019 [5]
- Agraphydrus kempi (d'Orchymont, 1922)
- Agraphydrus khasiensis Komarek, 2018 [4]
- Agraphydrus kodaguensis Komarek, 2018 [4]
- Agraphydrus komareki Yang & Jia, 2021 [15]
- Agraphydrus laocaiensis Komarek, 2019 [5]
- Agraphydrus latus Komarek, 2019 [5]
- Agraphydrus longipalpus (Jia, 1998) [21]
- Agraphydrus longipenis Komarek and Hebauer, 2018 [8]
- Agraphydrus lunaris Komarek, 2019 [5]
- Agraphydrus luteilateralis (Minoshima and Fujiwara, 2009) [22]
- Agraphydrus madagascarensis Komarek, 2020 [6]
- Agraphydrus maehongsonensis Komarek, 2019 [5]
- Agraphydrus malayanus (Hebauer, 2000) [10]
- Agraphydrus malkini Komarek, 2020 [6]
- Agraphydrus manfredjaechi Komarek, 2019 [5]
- Agraphydrus masatakai Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus matoposensis Komarek, 2020 [6]
- Agraphydrus mazzoldii Komarek, 2019 [5]
- Agraphydrus meghalayanus Komarek, 2018 [4]
- Agraphydrus microphthalmus Komarek, 2019 [5]
- Agraphydrus minutissimus (Kuwert, 1890) [23]
- Agraphydrus mirabilis Komarek, 2019 [5]
- Agraphydrus montanus Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus muluensis Komarek, 2019 [5]
- Agraphydrus musculus Komarek, 2019 [5]
- Agraphydrus namthaensis Komarek, 2019 [5]
- Agraphydrus nanus Komarek, 2018 [4]
- Agraphydrus narusei (Satô, 1960) [24]
- Agraphydrus nemorosus Komarek, 2019 [5]
- Agraphydrus nepalensis Komarek, 2018 [4]
- Agraphydrus niger Komarek, 2018 [4]
- Agraphydrus nigroflavus Komarek, 2019 [5]
- Agraphydrus obesus Komarek, 2019 [5]
- Agraphydrus obscuratus Komarek, 2018 [4]
- Agraphydrus obsoletus Komarek, 2018 [4]
- Agraphydrus occultus Komarek and Freitag, 2020 [7]
- Agraphydrus ogatai Minoshima, 2016 [25]
- Agraphydrus orbicularis Komarek, 2019 [5]
- Agraphydrus orientalis (d'Orchymont, 1932) [18]
- Agraphydrus palawanensis Komarek and Freitag, 2020 [7]
- Agraphydrus pallidus Komarek, 2019 [5]
- Agraphydrus papuanus Komarek, 2019 [5]
- Agraphydrus pauculus (Knisch, 1924) [26]
- Agraphydrus pauper Komarek, 2020 [6]
- Agraphydrus pelingeni Komarek and Freitag, 2020 [7]
- Agraphydrus penangensis Komarek, 2019 [5]
- Agraphydrus piceus Komarek, 2019 [5]
- Agraphydrus politus (Hansen, 1999) [11]
- Agraphydrus praecipuus (d'Orchymont, 1937) [27]
- Agraphydrus protentus Komarek, 2018 [4]
- Agraphydrus pseudoniger Yang & Jia, 2021 [15]
- Agraphydrus pullus Komarek, 2018 [4]
- Agraphydrus punctatellus Régimbart, 1903 [3]
- Agraphydrus punctulatus Komarek, 2018 [4]
- Agraphydrus puzhelongi (Jia, 2010) [28]
- Agraphydrus pygmaeus (Knisch, 1924) [26]
- Agraphydrus raucus Komarek, 2019 [5]
- Agraphydrus reductus Komarek and Hebauer, 2018 [8]
- Agraphydrus regularis (Hansen, 1999) [11]
- Agraphydrus reticulatus Komarek, 2019 [5]
- Agraphydrus reticuliceps Komarek and Hebauer, 2018 [8]
- Agraphydrus rhodesiensis Komarek, 2020 [6]
- Agraphydrus rhomboideus Komarek, 2019 [5]
- Agraphydrus rivalis Komarek, 2020 [6]
- Agraphydrus robustus Komarek and Hebauer, 2018 [8]
- Agraphydrus rostratus Komarek, 2018 [4]
- Agraphydrus rugosus Komarek, 2018 [4]
- Agraphydrus sabulosus Yang & Jia, 2021 [15]
- Agraphydrus sarawakensis Komarek, 2019 [5]
- Agraphydrus schoedli Komarek, 2019 [5]
- Agraphydrus schoenmanni Komarek and Hebauer, 2018 [8]
- Agraphydrus scintillans Komarek, 2019 [5]
- Agraphydrus scutifer Komarek, 2020 [6]
- Agraphydrus setifer Komarek and Hebauer, 2018 [8]
- Agraphydrus shaverdoae Komarek, 2019 [5]
- Agraphydrus siamensis (Hansen, 1999) [11]
- Agraphydrus sipekorum Komarek, 2018 [4]
- Agraphydrus skalei Komarek, 2019 [5]
- Agraphydrus spadix Komarek, 2019 [5]
- Agraphydrus spinosus Komarek, 2019 [5]
- Agraphydrus splendens Komarek and Hebauer, 2018 [8]
- Agraphydrus stagnalis (d'Orchymont, 1937)
- Agraphydrus stramineus Komarek, 2019 [5]
- Agraphydrus sucineus Komarek, 2019 [5]
- Agraphydrus sundaicus Komarek, 2019 [5]
- Agraphydrus tamdao Komarek, 2019 [5]
- Agraphydrus taprobanensis Komarek, 2018 [4]
- Agraphydrus tenuipalpis Komarek and Freitag, 2020 [7]
- Agraphydrus thaiensis Minoshima, Komarek, and Ôhara, 2015 [13]
- Agraphydrus tristis Komarek, 2019 [5]
- Agraphydrus tulipa Komarek, 2019 [5]
- Agraphydrus tumidus Komarek, 2020 [6]
- Agraphydrus tumulosus Komarek, 2018 [4]
- Agraphydrus umbrosus Komarek and Hebauer, 2018 [8]
- Agraphydrus uncinatus Komarek and Hebauer, 2018 [8]
- Agraphydrus usambaraensis Komarek, 2020 [6]
- Agraphydrus uvaensis (Hebauer, 2000) [10]
- Agraphydrus vadoni Komarek, 2020 [6]
- Agraphydrus variabilis Komarek and Hebauer, 2018 [8]
- Agraphydrus vietnamensis Komarek, 2019 [5]
- Agraphydrus villiersi (Balfour-Browne, 1958) [29]
- Agraphydrus wangmiaoi Komarek and Hebauer, 2018 [8]
- Agraphydrus yunnanensis Komarek and Hebauer, 2018 [8]
- Agraphydrus zetteli Komarek and Freitag, 2020 [7]
References
- ↑ "Agraphydrus Régimbart, 1903". Catalogue of Life. Species 2000: Leiden, the Netherlands. Retrieved 7 December 2025.
- 1 2 3 4 Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species". ZooKeys (1045): 1–236. Bibcode:2021ZooK.1045....1G. doi:10.3897/zookeys.1045.63810. ISSN 1313-2970. PMC 8233300. PMID 34228772.
- 1 2 Régimbart, MA (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
- 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Komarek, A. (2018). "Taxonomic revision of Agraphydrus Régimbart, 1903 II. The Indian Subcontinent" (PDF). Koleopterologische Rundschau. 88: 103–204. Archived from the original (PDF) on 2021-06-24. Retrieved 2022-01-07.
- 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 Komarek, A. (2019). "Taxonomic revision of Agraphydrus Régimbart, 1903 III. Southeast Asia (except Philippines) and Australian Region (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 89: 151–316. Archived from the original (PDF) on 2021-06-24. Retrieved 2022-01-07.
- 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 Komarek, A. (2020). "Taxonomic revision of Agraphydrus Régimbart, IV. Africa, Western Asia, and redescription of the genus (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 90: 127–200. Archived from the original (PDF) on 2021-02-14. Retrieved 2022-01-07.
- 1 2 3 4 5 6 7 8 9 10 11 12 Komarek, A.; Freitag, H. (2020). "Taxonomic revision of Agraphydrus Régimbart, 1903 V. Philippine species and their first DNA barcodes" (PDF). Koleopterologische Rundschau. 90: 201–242. Archived from the original (PDF) on 2021-02-14. Retrieved 2022-01-07.
- 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 Komarek, A.; Hebauer, F. (2018-07-30). "Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)". Zootaxa. 4452 (1): 1–101. doi:10.11646/zootaxa.4452.1.1. ISSN 1175-5334. PMID 30314249. S2CID 52975164.
- ↑ Régimbart, M. (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophi-lidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
- 1 2 3 Hebauer, F. (2000). "The genus Megagraphydrus Hansen, 1999, with description of new species (Coleoptera: Hydrophilidae)". Acta Coleopterologica. 16: 14–22.
- 1 2 3 4 5 Hansen, Michael (1999-01-01). "Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family". Insect Systematics & Evolution. 30 (2): 121–172. doi:10.1163/187631200X00228. ISSN 1876-312X.
- ↑ Hansen, M. (1997). "A new subfamily for a remarkable new genus and species of Hydrophilidae from New Guinea (Coleoptera: Hydrophilidae)". Annales Zoologici. 47: 107–110.
- 1 2 3 4 5 6 7 Minoshima, Y. N.; Komarek, A.; Ôhara, M. (2015). "A revision of Megagraphydrus Hansen (Coleoptera, Hydrophilidae): synonymization with Agraphydrus Régimbart and description of seven new species". Zootaxa. 3930: 1–63. doi:10.11646/1-63 (inactive 9 December 2025). PMID 25781810.
{{cite journal}}: CS1 maint: DOI inactive as of December 2025 (link) - ↑ d’Orchymont, A. (1927). "Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums". Journal of the Federated Malay States Museums. 13 (4): 246–252.
- 1 2 3 4 Yang, Zhen-ming; Jia, Fenglong; Jiang, Lu; Guo, Qiang (2021-05-18). "Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae)". Deutsche Entomologische Zeitschrift. 68 (1): 189–205. doi:10.3897/dez.68.66200. ISSN 1860-1324. S2CID 236330170.
- ↑ Ribera, I.; Hernando, C.; Cieslak, A. (2019). "Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae" (PDF). Acta Entomologica Musei Nationalis Pragae. 59: 253–272. doi:10.2478/aemnp-2019-0021. hdl:10230/44839. S2CID 198381025.
- ↑ d’Orchymont, A. (1937). "Descriptions of three new Hydrophilidae from India". Records of the Indian Museum. 39: 29–31.
- 1 2 3 d’Orchymont, A. (1932). "Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali". Archiv für Hydrobiologie. Supplement Band IX: 623–714. [pl. XIV–XVII.]
- ↑ Satô, M.; Yoshitomi, H. (2004). "Discovery of a second representative of the genus Horelophopsis (Coleoptera, Hydrophilidae) from the Ryukyu archipelago, Japan". Elytra. 32: 41–49.
- ↑ Matsui, E. (1994). "Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 20: 215–220.
- ↑ Jia, F. (1998). "A new genus Pseudopelthydrus gen. n. from Hainan Island, China (Coleoptera: Hydrophilidae: Hydrophilinae)". Chinese Journal of Entomology. 18: 225–230.
- ↑ Minoshima, Y. N.; Fujiwara, J. (2009). "Discovery of water scavenger beetle genus Megagraphydrus Hansen (Coleoptera, Hydrophilidae) from Japan, with description of a new species" (PDF). Elytra. 37: 53–64.
- ↑ Kuwert, A. (1890). "Bestimmungs-Tabellen der europäischen Coleopteren. XX. Heft. Hydrophilidae. II. Abteilung: Sphaeridiini und Helophorini". Verhandlungen des naturforschenden Vereins in Brünn. 28: 159–328.
- ↑ Satô, M. (1960). "One new genus and two new species of the subtribe Helocharae from Japan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 6: 76–80.
- ↑ Minoshima, Yûsuke N. (2016). "Taxonomic review of Agraphydrus from Japan (Coleoptera: Hydrophilidae: Acidocerinae)". Entomological Science. 19 (4): 351–366. doi:10.1111/ens.12213. ISSN 1479-8298. S2CID 89394202.
- 1 2 Knisch, A. (1924). "Neue neotropische Palpicornier. (Col. Hydrophilidae. - Op. 16.)". Wiener Entomologische Zeitung. 41: 114–140.
- ↑ d’Orchymont, A. (1937). "Contribution à l'étude des Palpicornia IX". Bulletin et Annales de la Société Entomologique de Belgique. 77: 213–255.
- ↑ Jia, Fenglong (2010-06-08). "Megagraphydrus puzhelongi, sp. n., a new water scavenger beetle from China (Coleoptera: Hydrophilidae: Hydrophilinae)". Zootaxa. 2498 (1): 65. doi:10.11646/zootaxa.2498.1.6. ISSN 1175-5334.
- ↑ Balfour-Browne, J. (1958). "La Reserve naturelle integrale du Mont Nimba, Guinea. VI. Coléoptères Hydrophiloides". Mémoires de l'Institut français d'Afrique noire. 53: 169–190.
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