↑ P. Morzadec and D. Brice. 2000. The Devonian of France: a tentative tie with the GSSP of the Devonian stages. 225:115-129
1 2 3 4 5 Darras, L., Derycke, C., Blieck, A., Vachard, D. 2008. The oldest holocephalan (Chondrichthyes) from the Middle Devonian of the Boulonnais (Pas-de-Calais, France). Comptes Rendus Palevol v. 7, no. 5, pp. 297–304
