Eubulides (insect)

The representatives of Eubulides are medium-sized, very slender and only slightly or hardly spined Obriminae species. The males reach 46 to 62 centimetres (18 to 24 in), the females 52 to 92 centimetres (20 to 36 in) in length. The head is flat and, like the pronotum, hardly reinforced or only covered with small tubercles. Only on the frontal margin of the elongated mesonotum spines may be present. There may be a few tubercles on the rear of the mesonotum. The middle femura are clearly toothed, the hind legs very strongly toothed. The secondary ovipositor of the females is designed as a curved laying sting.^[2][3][4][5]

The previously known distribution area of the genus includes the Philippine islands

Luzon, Leyte, Mindanao and Polillo. On Luzon there are representatives in the provinces Ilocos Norte, Mountain Province, Kalinga, Quirino, Ifugao, Quezon, Camarines Sur and Nueva Vizcaya, on Mindanao in the provinces Bukidnon and Agusan del Sur proven.^[5][6]

In 1877, Carl Stål established the genus Eubulides in the first description of Eubulides alutaceus, which became the type species of the genus.^[1] The name is dedicated to the Greek philosopher Eubulides.^[7] William Forsell Kirby placed the genus 1904 in the subfamily Eurycanthinae, today only considered as tribe Euricanthini. He added a second species to it with the newly written Eubulides spuria, which since 2005 has been regarded as synonym of Dryococelus australis.^[8] Josef Redtenbacher continues to treat the genus 1906 as monotypical and includes it in the tribe Obrimini.^[9] Two additional species were added through descriptions by James Abram Garfield Rehn and his son John William Holman Rehn in 1939.^[2] The 2022 by Mescel S. Acola, Jeremy Carlo B. Naredo and Orlando L. Eusebio in 2022 described Eubulides manobo was transferred by Frank H. Hennemann to the genus Armadolides, which was created specifically for this species. At the same time, Hennemann described four other Eubulides species.^[4][5]

Valid species are:^[4][1]

• Eubulides alutaceus Stål, 1877
• Eubulides blaan Hennemann, 2023
• Eubulides constanti Hennemann, 2023
• Eubulides igorrote Rehn, J.A.G. & Rehn, J. W. H., 1939
• Eubulides lumawigi Hennemann, 2023
• Eubulides taylori Rehn, J.A.G. & Rehn, J. W. H., 1939
• Eubulides timog Hennemann, 2023

In 2004 Oliver Zompro raised the Obrimini to the rank of a subfamily and divided them into three tribes. One of them was that of the Eubulidini. In addition to the type genus Eubulides, he also placed in this Tisamenus, Ilocano (now synonymous with Tisamenus), Hoploclonia, Stenobrimus, Heterocopus, Pterobrimus and Theramenes.^[3] This tribe was withdrawn in 2016 by Hennemann et al and is now a synonym for the Obrimini.^[10]

In their work on the spread and relationships within the Heteropterygidae, based mainly on genetic analysis, which was published in 2021, Sarah Bank et al also examined samples from five members of the genus Eubulides. Three turned out to be conspecific and were identified as Eubulides igorrote. Two more could not be assigned to any known species and represent new species. One of them was described by Hennemann in 2023 as Eubulides timog. The second, like Eubulides blaan, also described by Hennemann,^[5] comes from Mindanao, more precisely from Mt. Kitanglad in the province of Bukidnon. Within the Obrimini the genus forms a sister group with a clade from the genera Sungaya, Trachyaretaon and Trachyaretaon negrosanon.^[6]