Exostyleae

This clade is composed of 6 genera, most of which were traditionally assigned to the tribe Swartzieae.^[4][5] However, recent molecular phylogenetic analyses circumscribed these six genera into a strongly supported monophyletic clade.^{[1][2][6][7][8][9]} Synapomorphic traits that unite the members of this clade include non-papilionate flowers, "serrate and sometimes spinescent leaflet or leaf margins, standard position variable in the floral bud, basifixed anthers, and drupaceous fruits".^[1][2][6][7] They are also united by wood anatomy, sharing an "uncommon presence of crystals in ray cells",^[1][10][11] and floral ontogeny, sharing "unidirectional initiation of five sepals, simultaneous initiation of petals, and[…]unusual antepetalous stamens initiating before the antesepalous ones."^[12]

• Exostyles Schott
• Harleyodendron R. S. Cowan
• Holocalyx Micheli
• Lecointea Ducke.
• Uribea Dugand & Romero
• Zollernia Wied-Neuw. & Nees