Heliocopris

Heliocopris (large dung beetles) is a genus of Scarabaeidae or scarab beetles in the superfamily Scarabaeoidea.^[1][2][3] Forty-seven of the fifty-two known species are found in Africa,^[2] but a few are found in southern and southeast Asia.^[4]

Phylum:Arthropoda

Class:Insecta

Order:Coleoptera

Suborder:Polyphaga

Quick facts Scientific classification, Species ...

Heliocopris
Heliocopris dominus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Scarabaeiformia
Family:	Scarabaeidae
Subfamily:	Scarabaeinae
Tribe:	Coprini
Genus:	Heliocopris Hope, 1837
Species
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Heliocopris includes some of the world's largest dung beetles (their size only rivalled by certain Catharsius and Coprophanaeus), with H. anderson, H. colossus, H. dilloni, H. dominus, H. gigas and H. tyrannus reaching up to about 7 cm (2.8 in) long, although a more typical size for the members of this genus is 3–5 cm (1.2–2.0 in).^[5] They are generally black or dark brown beetles where males tend to have horns on their head or pronotum, which females lack.^[5] Some are associated with the dung of elephants, rhinos or hippos, but there are large variations among the species in the genus and dung from other animals, like domestic cattle or monkeys, is also used. The female lays her eggs in a tunnel she digs under the excrement.^[5]

Heliocopris has been placed in the tribe Dichotomiini, but Montreuil places them in Coprini.Montreuil, Olivier (1998). "Analyse phylogénétique et paraphylie des Coprini et Dichotomiini (Coleoptera: Scarabaeidae). Scénario biogéographique". Annales de la Société Entomologique de France. 34 (2): 135–148. doi:10.1080/21686351.1998.12277767. S2CID 82053231.