Kladno Formation

Geologic formation in the Czech Republic From Wikipedia, the free encyclopedia

The Kladno Formation is a Carboniferous formation in the Czech Republic which preserves a wide array of temnospondyls, lepospondyls and fish,[1] among other fauna.

TypeGeological formation
UnderliesTýnec Formation
RegionPilsen
CountryCzech Republic
Quick facts Type, Underlies ...
Kladno Formation
Stratigraphic range: Westphalian
TypeGeological formation
UnderliesTýnec Formation
Location
RegionPilsen
CountryCzech Republic
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Paleobiota

Invertebrates

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
 Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
More information Invertebrates, Genus ...
Invertebrates
Genus Species Higher taxon Notes Images
Necymylacris N. scudderi Blattodea Formerly included within Phylloblatta[2]
Bohemiatupus B. elegans Meganeuridae[3] One of the larger meganisopterans
Bojophlebia B. prokopi Palaeoptera Largest insect by wingspan other than palaeodictyopterans or griffinflies.[4]
Carbotriplura C. kukalovae Dicondylia Sister group to Pterygota, formerly classed as a Bojophlebia nymph.[5]
Pronaidites P. carbonarius, P. arenivorus Tubificidae "P. crenulatus" actually an ichnofossil[6]
Microconchus M. vorax, M. minimus Tentaculita Formerly classed as a spirorbid[6]
Prolimulus P. woodwardi Bellinuridae Likely clades with other genera like Alanops.[7]
Eolycosa[8] E. lorenzii Mesothelae incertae sedis[9] Spider affinities doubted in a 1996 paper[10]
Geralycosa[8] G. fricii Arthromygalidae May not be a spider[10]
Rakovnicia[8] R. antiqua Tetrapulmonata incertae sedis[9]
Pyritaranea P. tubifera Araneae incertae sedis[9] Synonymised with Eopholcus
Nyranytarbus[9][11] N. hofmanni, N. longipes Trigonotarbida Formerly placed within Hemiphrynus
Doubravatarbus D. krafti Aphantomartidae Likely arboreal, due to being found on a liana-like plant and having relatively gracile legs.[12]
Tynecotarbus T. tichaveki Trigonotarbida Collected alongside two more fragmentary indeterminate fossils[13]
Palaranea P. borassifolia Araneae incertae sedis Known from a different locality to other Kladno fauna[9]
Anthracomartus[9] A. carcinoides, A. bohemica, A. elegans Trigonotarbida
A. trilobitus fossil (from North America)
Isobuthus I. nyranensis[14] Scorpiones
Prothelyphonus P. bohemicus[15] Uropygi Synonymous with Geralinura[16]
Geralinura fossil (from North America)
Dinopilio D. gigas Arachnida incertae sedis[9] Very large fossil, hence the name
Geratarbus[8] G. bohemicus Phalangiotarbida
Orthotarbus[8] O. nyranensis Phalangiotarbida
Pleurolycosa P. prolifera Arthropoda incertae sedis[9]
Nyranius N. costulatus, N. tabulatus Xyloiuloidea Formerly within Xylobius[17]
N. costulatus fossil
Pleurojulus P. biornatus, P. levis Pleurojulida Also known from Mazon Creek[18]
P. biornatus fossil
Isojulus I. constans Pleurojulida Synonymised with two Pleurojulus species[18]
Purkynia P. lata Zosterogrammida Very poorly preserved[19]
Illustration of a P. lata fossil
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Vertebrates

More information Vertebrates, Genus ...
Vertebrates
Genus Species Higher taxon Notes Images
Archaeothyris A. sp Ophiacodontidae Only known from a jaw fragment[20]
Archaeothyris reconstruction
Brouffia B. orientalis Eureptilia Formerly placed within Gephyrostegus[21]
Brouffia skull reconstruction
Coelostegus C. prothales[21] Eureptilia Recovered as the basalmost eureptile[22]
Coelostegus skull reconstruction
Gephyrostegus G. bohemicus[23] Gephyrostegidae Formerly included another species, G. watsoni[24]
Gephyrostegus reconstruction
Solenodonsaurus S. janenschi Reptiliomorpha? Resembles diadectomorphs, yet likely not closely related to amniotes[25]
Solenodonsaurus reconstruction
Urocordylus U. angularis[26] Urocordylinae Also known from Ireland
Urocordylus reconstruction
Sauropleura S. scalaris Sauropleurinae Formerly included in Urocordylus[27]
Sauropleura life reconstruction
Scincosaurus S. crassus Scincosauridae Formerly synonymised with Keraterpeton,[26] related to diplocaulids[28]
Scincosaurus reconstruction
Keraterpeton K. galvani[26] Diplocaulidae Bore an unusually long tail
Keraterpeton reconstruction
Hyloplesion H. longicostatum "Microsauria" Synonymous with "Orthocosta" and "Seeleya"[26][29]
Hyloplesion reconstruction
Microbrachis M. pelikani Recumbirostra Synonymised with three other species in the same genus[29]
Microbrachis reconstruction
Ricnodon R. copei Recumbirostra? May be related to Saxonerpeton[29]
Crinodon C. limnophyes Tuditanidae Name derives from an anagram of Ricnodon[29]
Sparodus S. validus Gymnarthridae? Resembles Hylerpeton[29]
Diplovertebron D. punctatum Embolomeri Synonymous with "Nummulosaurus"[26]
Diplovertebron reconstruction
Capetus C. palustris[26] Temnospondyli One of the most basal temnospondyls[30]
Capetus reconstruction
Cochleosaurus C. bohemicus Cochleosauridae Adult specimens only found in 2003[31]
Cochleosaurus life restoration
Mordex M. calliprepes[26] Trematopidae Unclear position within the family[32]
Mattauschia M. laticeps Trematopidae Formerly included in Mordex,[32] includes "Potamochoston" and several "Limnerpeton" species[33]
Nyranerpeton[34] N. amilneri Micromelerpetontidae Also known from the Montceau-les-Mines lagerstätte[35]
Limnogyrinus L. elegans Micromelerpetontidae Formerly included within "Limnerpeton"[33]
Oestocephalus[36] O. granulosum, O. nanum Aistopoda Formerly included within Ophiderpeton[26]
Oestocephalus reconstruction
Phlegethontia P. longissima, "Dolichosoma scutiferum" Aistopoda Formerly included within the nomen nudum "Dolichosoma"[37][26]
Phlegethontia reconstruction
Baphetes B. orientalis Baphetidae Formerly known as "Loxomma bohemicum"[38]
Stambergichthys S. macrodens Actinopterygii Likely a high-level predator in its ecosystem[1]
Sceletophorus S. biserialis, S. verrucosus Trissolepididae[39]
Pyritocephalus P. sculptus Haplolepiformes[39] Synonymous with other Pyritocephalus species
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Plants

Non-seed plants

More information Non-seed plants, Genus ...
Non-seed plants
Genus Species Higher taxon Notes Images
Kladnostrobus K. clealii, K. psendae Lycopsida Within the monotypic family Kladnostrobaceae due to differing sporangia structure[40]
Lepidophloios L. acerosus Lepidodendrales Mistakenly spelled "Lepidofloios" in the paper reporting its presence[41]
Lepidophloios fossil (from Scotland)
Lepidodendron L. acutum, L. simile,[41] L. aculeatum, L. ophiurum,[42] L. lycopodioides[43] Lepidodendrales One of the most abundant Carboniferous plants
Lepidodendron restoration
Flemingites F. sp Lepidodendrales Cone taxon[41]
Lepidocarpon L. majus[42] Lepidodendrales Sporangium taxon[41]
Sigillaria S. rugosa, S. diploderma Lepidodendrales S. diploderma only known from a small fragment[41]
Sigillaria reconstruction
Syringodendron S. sp Lepidodendrales Stem taxon[41]
Omphalophloios O. feistmantelli[42] Isoetales Questionable specimen known from a fragment of a fertile axis[41]
Omphalophloios fossil
Stigmaria S. ficoides[41] Lepidodendrales Form taxon representing roots
Stigmaria fossil
Calamites C. carinatus, C. distachya?, C. cistii?[42] Calamitaceae Second species is closest to C. distachya, but poor preservation means its affinity is unclear[41]
Reconstruction of the entire Calamites tree
Annularia A. radiata[41] Calamitaceae Leaf taxon of Calamites stems
A. stellata fossil
Asterophylites A. longifolius, A. equisetiformis, A. grandis[43] Calamitaceae One specimen preserves the tip of a branch with dense leaves[41]
Sphenophyllum S. cuneifolium, S. pseudoaquense, S. ?majus,[43] S. priveticense[44] Sphenophyllales Very common, but often poorly preserved[41]
Sphenophyllum reconstruction
Bowmanites B. priveticensis, B. myriophyllus, B. pseudoaquensis, B. brasensis[45] Sphenophyllales Cones of Sphenophyllum[44]
Lobatopteris L. aspidioides Marattiales Only one small fragment preserved[41]
L. corsinii fossil (from Spain)
Corynepteris C. sternbergii, C. angustissima, C. essinghii[42] Zygopteridales C. sternbergii may be synonymous with C. angustissima[41]
C. angustissima fossil
Senftenbergia S. plumosa Tedeleaceae (Filicales) Common elsewhere, yet rare in Kladno[41]
S. dentata fossil (from Spain)
Zeilleria Z. delicatula Filicales incertae sedis[41] Shares its name with a Jurassic brachiopod
Renaultia R. crepinii Filicales incertae sedis Only known from a fragment[41]
R. lebachensis fossil (from Spain)
Sphenopteris S. rotundifolia, S. mixta, S. delicatula, S. acutiloba, S. ujezdensis, S. pulcherrima,[42] S. cirrhifolia,[46] S. spinosa[43] Filicales incertae sedis (rotundifolia), Lyginopteridales (others) Includes seed plants and a fern[41]
S. rotundiloba fossil
Aphlebia A. sp Indeterminate Artificial taxon; may belong to another species preserved.[41]
A. crispa fossil (accompanied by Polymorphopteris)
Noeggerathia N. foliosa, N. intermedia[47] Noeggerathiales Most common plant in the sampled fossils[42]
Drawings of N. expansa fossils
Archaeonoeggerathia A. schatzlarensis[47] Noeggerathiales
Discosoropteris D. zlatkokvacekii, D. chlupatum Filicales incertae sedis Likely a short, upright plant[48]
Spencerites S. leismanii, S. havlenae[43] Lycopsida incertae sedis A member of the "sub-arborescent" lycophytes[49][46]
Dendraena D. pinnatilobata Anachoropteridaceae (Filicales) Likely grew near river margins,[50] a very common plant[46]
Kidstonia K. heracleensis Filicales Likely a small understorey fern[46]
Desmopteris D. alethopteroides, D. longifolia[43] Filicales Likely bore procumbent (growing along the ground) rhizomes[46]
Selaginella S. gutbieri,[43] S. labutae[51] Selaginellaceae Extant genus
The extant S. selaginoides (from Estonia)
Palaeostachya P. distachya, P. gracilima Calamitaceae Cone taxon,[43] although with a complete plant known from the early Permian of China[52]
P. sp fossil (from Mazon Creek)
Pecopteris P. aspidioides, P. tuffitica[53] Psaroniaceae Likely a mid-sized tree fern[43]
P. sp fossil (from Mazon Creek)
Sonapteris S. pilsensis, S. barthelii, S. bekii[43] Botryopteridaceae (Filicales) Possibly a liana-like plant?[53]
Oligocarpia O. lindsaeoides Gleicheniales? Probably a liana-like plant[43]
O. gutbieri fossil (from Germany)
Adiantites A. sp Progymnospermopsida Likely a low-lying plant[43]
Echinosporangites E. libertite Filicopsida? Sporangium taxon, parent plant unknown[54]
Polysporia P. drabekii, P. rothwelii, P. radvanicensis, P. robusta Isoetales A sub-arborescent lycophyte[55]
Huttonia H. spicata Equisetales Cone taxon[56]
Thomasites T. serrata, T. elongatus Lycopsida T. elongatus formerly placed in Selaginellites[57]
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Seed plants

More information Seed plants, Genus ...
Seed plants
Genus Species Higher taxon Notes Images
Alethopteris A. lonchitica, A. distantinervosa, A. pseudograndinioides, A. kettneri, A. serlii Medullosales A. pseudograndinioides is a replacement name for the illegitimate "A. grandinioides"[58]
A. lonchitica fossil
Neuromariopteris N. scandens Callistophytales Likely a creeping plant growing on ridge slopes[59]
Eusphenopteris E. nummularia Lyginopteridales While a large (>1 m) frond was found, it could not be transported from the site.[41]
E. striata fossil
Lonchopteris L. rugosa Medullosales Possible record of L. chandesrisii (although based on a poorly preserved specimen) also known[58]
L. rugosa fossil
Macroneuropteris M. scheuchzeri Medullosales Formerly placed within Neuropteris[58]
M. scheuchzeri frond reconstruction
Neuropteris N. plicata Medullosales Placement of this species is unclear in relation to N. ovata.[58]
N. ovata fossil (from Kansas)
Odontopteris O. reichiana Medullosales May be synonymous with O. brardii[58]
O. genuina fossil
Callipteridium C. rubescens, C. armasii Medullosales May be synonymous with C. jongmansii[58]
C. gigas fossil
Laveineopteris L. loshii, L. tenuifolia, L. bohemica, L. lubnensis, L. hollandica Medullosales L. loshii is abundant elsewhere, while some other species are endemic to the Kladno Formation[58]
L. rarinervis fossil
Palaeoweichselia P. defrancei Medullosales Described from Kladno in a 2020 overview paper despite specimens being already known prior[58]
Linopteris L. neuropteroides, L. obliqua, L. weigelii Medullosales L. neuropteroides is split into two forms, neuropteroides and minor.[58]
L. subbrongniarti fossil
Paripteris P. linguaefolia Medullosales Included alleged records of P. gigantea[58]
Havlenaea[60] H. coriacea, H. stradonitzensis Medullosales Formerly placed within Neuropteris[58]

Mixoneura

M. muensterifolia

Medullosales

Too poorly preserved to place within a new genus, yet does not belong to Mixoneura either[58]
Rhacopteris R. elegans Pteridospermatophyta Only known from one isolated pinnule (leaf cluster)[42]
R. elegans fossil (from Italy)
Cordaites C. theodorii, C. wartmannii, C. kladnoensis,[61] C. borassifolius[43] Cordaitales C. theodorii is only known from this formation[42]
C. foliatus fossil (from France)
Palmatopteris P. furcata Lyginopteridales Probably a liana-like plant[43]
P. furcata fossil (from Germany)
Mariopteris M. muricata Lyginopteridales Likely a vining plant, as its fossils are all associated with Lepidodendron.[43]
M. muricata fossil (from Spain)
Fortopteris F. radnicensis Medullosales Not given a type specimen in its original description within Mariopteris.[62]
Rhodeites R. gutbieri Lyginopteridales Likely a canopy-dwelling climber[63]
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References

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