Lipica Formation
Mesozoic geologic formation in Italy
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The Lipica Formation (Also known as Upper Aurisina Formation, "Formazione di Monrupino", Trieste Karst Limestone formation, "Tomaj Limestone", "Repen formation" and "Lipiza formation", as well includes part of the informal "Liburnia formation")[1][2][3] is a Mesozoic geologic formation in Italy and Slovenia.[3] This limestones are found in the Trieste area and are of Late Cretaceous (Santonian-Campanian) age, being a local record of Carbonate platform limestones, historically quarried by cutting large blocks using steel wire, what has allowed more access to them, as numerous quarries allow excellent exposure.[3][4] Due to being cut into large blocks and slabs, that usually end stored near the quarries, detailed, three-dimensional study of the rock's composition and fossils can be easily done.[5][6]
- Tomaj limestone
- Zolla member
| Lipica Formation | |
|---|---|
| Stratigraphic range: Santonian-Campanian ~ | |
Former Aurisina-type limestone quarry near Lipica | |
| Type | Geological formation |
| Sub-units |
|
| Overlies | Sežana Formation |
| Area | Friuli-Venezia Giulia |
| Lithology | |
| Primary | Limestone |
| Location | |
| Coordinates | 45.8°N 13.6°E |
| Approximate paleocoordinates | 23.5°N 16.2°E |
| Region | Trieste, Sežana |
| Country | Italy, Slovenia |
| Type section | |
| Named for | Lipica, Sežana |
Characteristics

The Lipica formation usually has a light grey background, sometimes appearing hazelnut in color. It is known for its high purity, compactness, and uniformity. The quarries where it is extracted consist of thick, solid layers of stone. Different varieties of Aurisina marble are distinguished by shades of grey and the distribution, size, and orientation of the organic remains within the stone. Well-known varieties include Aurisina Fiorita, Aurisina Lumachella, Aurisina Chiara, and Roman Stone, among others.[3][7][8]
Aurisina marble's chemical composition and physical properties make it ideal for use in architecture, both for structural and decorative purposes, as well as in statues and monuments. The stone is valued for its durability and aesthetic appeal in various applications.[3][7][9]
Historical Importance

The Lipica formation has been used since the Roman Republic. Evidence from ancient monuments in Aquileia, a Roman colony founded in 181 BC, suggests that quarrying began in the 1st century BC and continued until the 5th century AD. The stone was widely used in Aquileia for architectural elements, statues, and funerary monuments.[10][11] The nearby city of Tergeste (modern Trieste) also made extensive use of Aurisina Limestone for public buildings and sculptures. Its use spread across northern Italy, with artifacts found as far as Pavia.[10][9]
In the post-Roman period, Aurisina marble was notably used in the Mausoleum of Theodoric in Ravenna, built around 520 AD. However, its use declined during the medieval and Baroque periods. In Venice, it was replaced by Istrian stone, but in Trieste, it saw a revival in the 18th century as the city grew following the establishment of the free port by Emperor Charles VI. Notable examples of its use in Trieste include Palazzo Pitteri (1780), the Stock Exchange (1802), and the renovation of the Greek Orthodox Church of San Nicolò dei Greci (1819).[9][12]
Aurisina Limestone's "golden age" came under the Austro-Hungarian Empire, particularly after the completion of the Southern Railway, connecting Trieste to Vienna. The stone was used in major projects throughout the empire, including the Parliament and Hofburg in Vienna, the State Opera in Budapest, and many buildings in cities like Graz, Munich, and Ljubljana. By 1890, the industry employed over 3,000 workers.[9]
After Trieste's annexation to Italy in 1918, Aurisina Limestone was used in large-scale projects like the Military Shrine of Redipuglia and Milan's Central Station, where 38,000 tons were utilized. During this period, the marble also reached international markets, being used in Egypt and the United States.[9]
In the post-war era, Aurisina Limestone continued to be popular in foreign markets, with notable projects including the Atlanta subway in the U.S., Berlin airport, and La Défense in Paris. In 1989, it was used for Milan Metro's Line 3. More recently, it has featured in prestigious architectural projects, such as Milan's CityLife Shopping District designed by Zaha Hadid and Turin's Lavazza headquarters. Internationally, it is used in projects across Europe, Asia, Australia, and the U.S.[3][13]
Paleoenvironment


The formation is characterized by lagoonal facies with episodes of higher energy and rare bivalve patch reefs, such as Chondrodonta and Rudists, which are found throughout the entire unit. The lower section, referred to as the "Zolla member" consists of bivalve-rich limestone, including rudists and Chondrodonta, as well as foraminiferal limestone, often interbedded with dolomitized dark wackestones. In the upper part of this member, pelagic limestone with Pythonella fossils is present.[1][15] The main body of the Aurisina Limestone is characterized by peritidal carbonates, with some dark laminated facies, dominated by rudists, benthic foraminifera, cyanobacteria (Decastronema), and algae (Thaumatoporella).[16][3] The Paleoenvironment of this unit is accepted to be a succession of emerged and shallow marine carbonate platform settings, including inner lagoons, high-energy shoals, tidal channels, and rudist accumulations[1][4] The complex dolomitization and silicification in the region may be linked to a Monsoon climate with alternating wet and dry periods. Comparable dolomitization occurs across the Adriatic Platform, particularly in the Middle-Lower Cenomanian.[17]
The Villaggio del Pescatore fossil site records a progressive environmental shift from a stable shallow-marine carbonate platform to a restricted, tectonically controlled basin.[15] Initially, the area was a protected inner platform with shallow, calm seas, where early sediments accumulated in poorly oxygenated lagoons before giving way to better-ventilated marine conditions with normal salinity.[18] Rudist-bearing facies developed during this stage, though their small size suggests ecologically stressed conditions. Later, higher-energy shallow-marine settings produced bioclastic sands and rudstones under well-oxygenated conditions.[18] During the Lower Senonian, synsedimentary tectonic activity disrupted the platform, creating a small fault-bounded depression.[15] Differential block movements led to erosion of emergent areas and collapse of basin margins, generating polygenic breccias composed of reworked platform material, including clasts showing oxidation, pedogenic alteration, and evidence of subaerial exposure. As tectonic activity decreased or relative sea level rose, clastic input diminished and the basin evolved into a quiet, restricted water body.[15] The depression was progressively filled by finely laminated limestones deposited under low-energy conditions with limited circulation and oxygen-poor bottom waters. Seasonal lamination indicates cyclic sedimentation, consistent with fluctuating freshwater input and variable salinity in a brackish lagoon or shallow coastal lake. Microfossils, freshwater algae, plant traces, and slump structures reflect continental influence and ongoing instability along the basin margins.[18] These anoxic conditions enabled exceptional preservation of vertebrate remains, including hadrosaurs, crocodyliforms, fish, and crustaceans.[14] The surrounding bauxite and coal deposits indicate a humid climate and a karst landscape where surface water was scarce, making the basin a persistent water source. The depression likely functioned as a cenote-like feature that attracted animals, some of which drowned and were preserved at its stagnant base. A subsequent Upper Santonian transgression restored open-marine carbonate platform conditions, sealing the restricted basin deposits and preserving the unique fossil record.[14] More updated works in the Villaggio del Pescatore outcrops revelated beautifully preserved tidal rhythmites that pointed to steady tidal deposition in a protected paralic flat, likely semidiurnal, accumulating over roughly 40 years across 14 meters of sediment.[19] Geochemistry confirmed tidal origins, with dark packages reflecting wetter, runoff-heavy periods rich in organic matter and silt, while lighter ones indicate drier, better-oxygenated conditions with more aeolian influence. Salinity fluctuated between marine and freshwater inputs in this dynamic coastal setting.[19] The fossils were rapidly buried by tidal drapes and stabilized by microbial mats, preserving soft tissues and preventing decay or scavenging. Larger debris-flow beds with reworked clasts record high-energy flood pulses that carried in carcasses and plants from inland. Finally, palaeoclimate models support a monsoon-dominated regime at the site's subtropical latitude, with strong seasonal rainfall and tropical cyclones driving the repeated flood events.[19]
Biota
Color key
|
Notes Uncertain or tentative taxa are in small text; |
Foranimifera
Unnamed Miliolidae and Rotaliidae tests are known from Villaggio del Pescatore.[15]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Accordiella[15][20] |
|
|
Tests | Benthic zone Chrysalidinidae | |
| Calveziconus[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Dictyoconinae | |
| Cuneolina[20] |
|
Slivia quarries | Tests | Benthic zone Cuneolinidae | |
| Dicyclina[15][20] |
|
|
Tests | Benthic zone Dicyclinidae | |
| Fleuryana[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Charentiidae | |
| Goupillaudina[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Osangulariidae | |
| Keramosphaerina[15][20] |
|
|
Tests | Benthic zone Keramosphaeridae | |
| Metacuvillierinella[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Rhapydioninidae | |
| Moncharmontia[15][20] |
|
|
Tests | Benthic zone Charentiidae | |
| Murciella[15][20] |
|
|
Tests | Benthic zone Rhapydionininae | |
| Murgella[15] |
|
Villaggio del Pescatore | Tests | Benthic zone Praerhapydionininae | |
| Neorotalia[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Pararotaliinae | |
| Pilatorotalia[21] |
|
Villaggio del Pescatore | Tests | Benthic zone Calcarinidae | |
| Pseudocyclammina[20][21] |
|
|
Tests | Benthic zone Amijellinae | |
| Reticulinella[21][20] |
|
|
Tests | Benthic zone Loftusiidae | |
| Scandonea[15][20] |
|
|
Tests | Benthic zone Praerhapydionininae |
Bivalves
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Biradiolites[15][4][22] |
|
Isolated Shells | Rudist of the family Radiolitidae | ||
| Bournonia[4][22][24] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Chondrodonta[26][27] |
|
|
Isolated Shells | Scallop of the family Chondrodontidae | |
| Durania[27][28] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Eoradiolites[28] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Gorjanovicia[15][4][22][24] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Hippuritella[24] |
|
|
Isolated Shells | Rudist of the family Hippuritidae | |
| Hippurites[15][4] |
|
|
Isolated Shells | Rudist of the family Hippuritidae | |
| Katzeria[15][4][22][24] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Medeella[24][27] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Neithea[27] |
|
|
Isolated Shells | Scallop of the family Neitheoidae | |
| Paronaites[28] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Praelapeirouseia[24] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Praeradiolites[27] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Rajka[4] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Radiolites[15][24][27] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Radiolitella[22] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Requienia[25] |
|
|
Isolated Shells | Rudist of the family Requieniidae | |
| Sauvagesia[15][24][27][28] |
|
|
Isolated Shells | Rudist of the family Radiolitidae | |
| Vaccinites[22][24] |
|
|
Isolated Shells | Rudist of the family Hippuritidae |
Cephalopoda
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Placenticeratidae indet.[29][30] | Indeterminate |
|
Isolated Jaws, aptychi, Shells, associated soft parts and gut contents | Ammonite |
Arthropods
Rare to common Ostracod fragile smooth shells are seen at Villaggio del Pescatore.[15]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Caridea indet.[31][32] | Indeterminate |
|
Multiple specimens | Shrimps resembling extant Neocaridina.[33] | |
| Cirolanidae indet.[33] | Indeterminate |
|
MCSNT 57229 | Isopods similar to Cirolana.[33] | |
| Heteroptera indet.[33] | Indeterminate |
|
MCSNT 72/88 | Insects similar to modern Closterotomus.[33] | |
| Palaemon?[33] | P. sp. P. cf.antonellae |
|
Multiple specimens | Palaemonidae Shrimps similar to extant Palaemon.[33] |
Cnidaria
Stony coral polyps or fragments are seen in Villaggio del Pescatore.[15]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Jellyfish indet.[34] | Indeterminate |
|
BJ 1660, body impression | It has a diameter of 2.5 cm | |
| Hydnophora[35] | H. multilamellosa |
|
Impressions | Merulinidae stony coral |
Porifera
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Sarmentofascis[36] | S. zamparelliae |
|
Imprints | Cladocoropsidae Axinellidan |
Echinodermata
Abundant sea urchins spines/radioles and rare cases, as well Crinoid stem fragments/entrochi are known from Villaggio del Pescatore.[15]
Chondrichthyes
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Rhinobatos[34][38] | R. sp. "F" |
|
BJ 1380, single complete specimen | A Guitarfish |
Bony Fish
At least 120 small fishes (2–5 cm, apparently monospecific) and others of larger size (less than 20 cm) are known from Villaggio del Pescatore.[14]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Anguilliformes indet.[32] | Indeterminate |
|
Single complete specimen | This would be the oldest anguilliform found in Italy | |
| Cypriniformes? indet.[32] | Indeterminate |
|
Several small complete and incomplete specimens | This determination still needs to be confirmed | |
| Enchodus[32][34][39] | E. spp. |
|
Several small complete and incomplete specimens | A member of Enchodontidae | |
| Heckelichthys[37] | H. microdon |
|
BJ1521, complete specimen | A member of Ichthyodectidae, previously referred to Chirocentrites. | |
| Holocentridae indet.[32] | Indeterminate |
|
Several small complete and incomplete specimens | Represented mostly by small-sized specimens | |
| Hoplopteryx[37] | H. stachei |
|
BJ1566, subcomplete specimen | A member of Beryciformes. | |
| Ichthyodectidae indet.[37] | Indeterminate |
|
BJ2198, subcomplete specimen; Referred BJ396, BJ1659, BJ1730, BJ2198 | ||
| Palaeobalistum?[40] | P.? spp. |
|
Several small complete and incomplete specimens | A member of Pycnodontidae | |
| Parachanos[32] | P. ssp. |
|
Several isolated and articulated remains | A member of Chanidae. The attribution to this genus is dubious | |
| Polazzodus[41][42] |
|
|
Multiple complete specimens and isolated teeth | A member of Pycnodontidae | |
| Pycnodontiformes indet.[32] | Indeterminate |
|
Multiple complete specimens and isolated teeth | Some specimens have been referred to the genus Coelodus | |
| Rhynchodercetis[32][39] |
|
|
Multiple specimens, complete and incomplete | A member of Dercetidae. The most abundant genus at Polazzo | |
| Sloveniantriacanthus[43] | S. saksi |
|
PMS, VS-001, single specimen | A member of Cretatriacanthidae |
Turtles
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Chelonioidea indet.[32] | Indeterminate |
|
MPCM11398, part of a forelimb (humerus) and carapace (plate); MPCM11396, scattered remains among which a vertebra, parts of limbs and elements of the cintiscapularis can be recognized. | Overall about fifteen finds referable to sea turtles | |
| Testudinata indet.[31] | Indeterminate |
|
Incomplete specimens | Villaggio del Pescatore specimens suggest a large-sized taxon. |
Crocodiles
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Acynodon[44][45] |
|
|
MCNST 57248, A partial skeleton in anatomical connection.
Multiple referred specimens: MCNST 57032, MCSNT 57031, MCSNT 57245, MCSNT 21.S239, MCSNT 21.S239 |
A small Durophagous crocodrylomorph | |
| Crocodylia indet.[45] | Indeterminate |
|
MCSNT 57033, possibly a rib; MCSNT 57035, mandibular symphysis; MCSNT 57036, anteroposteriorly long vertebra; MCSNT 57037, fragmented ventral osteodermal surface. Referred Multiple teeth | ||
| Doratodon[39][47] |
|
|
MCSNT 57035, two partial dentaries; MPCMS97/118, isolated tooth crown; ACKK-D-8/088 & ACKK-D-8/081, teeth | Either a Notosuchian or a convergently evolved Paralligatoridae | |
| Notosuchia indet.[48] | Indeterminate |
|
MPCM 11720, shed tooth crown | The 1st record of the group on Italy, similar to Araripesuchus |
Pterosaurs
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Pterodactyloidea indet.[49] | Indeterminate |
|
MCSNT 13450, an alar metacarpal. | Shows a mix of features found in Pteranodon and some more basal pterodactyloids |
Dinosaurs
| Genus | Species | Location | Material | Description | Image |
|---|---|---|---|---|---|
| Coelurosauria? indet.[50] | Indeterminate |
|
N° BJ 1742 – feather | Referral not confirmed | |
| Hadrosauroidea indet.[46] | Indeterminate |
|
ACKK-D-8/20, 121, teeth | Dominant teeth at the locality | |
| Iguanodontidae? indet.[46] | Indeterminate |
|
ACKK-D-8/25, 8/04, teeth | Resemble Iguanodontid Teeth, but can belong to a convergently evolved Rhabdodontid | |
| Ornithopoda indet.[46] | Indeterminate |
|
ACKK-D-8/21-Z1, teeth | Different from Rhabdodon | |
| Tethyshadros[21] | T. insularis |
|
SC 57021, type specimen ("Antonio"), virtually complete specimen; SC 57022 ("Primus"), partial, articulated forelimbs. The tail and potentially other skeletal elements are still in situ; SC 57023, isolated left pubis; SC 57024, partial scapula; SC 57025, cervical vertebra; SC 57026 ("Secundus"), complete but heavily crushed skull, also a large unprepared block containing the proximal vertebral column; SC 57027, sacral vertebrae; SC 57028, a series of ten, fully prepared, articulated distal caudal vertebrae; SC 57247 ("Bruno"), articulated skeleton; SC 57256, isolated rib; SC 57257, proximal end of a femur; A series of thirteen, articulated, distal caudal vertebrae; Large articulated elements still in situ ("Tertius"); Large articulated elements ("Zdravko") | A small hadrosauromorph dinosaur | |
| Theropoda indet.[21][51] | Indeterminate |
|
SC 57030, "arctometatarsalian metatarsal"; Another specimen may represent an epiphysis of a long bone, perhaps of a femur. | Likely a small Abelisaur |
"Algae"
Freshwater Characeae gyrogonites, oogonia, and whorls/verticilli are found in Villaggio del Pescatore.[15]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Decastronema[15] | D. kotori |
|
Imprints | Mat-forming blue-green algae | |
| Thaumatoporella[20] | T. ssp. |
|
Imprints | Potentially an encrusting Green algae |
Flora
Palynology of the Villaggio del Pescatore section has been studied, dominated by a few gymnosperms and several angiospermous pollen types.[1] The distribution of plants in this outcrop includes pollen from tropical lowland plants (palms, shrubs); pollen from taller trees (conifers) from more temperate uplands; possible rhizoliths (root/stem traces of land plants along basin margins).[15]
| Genus | Species | Location | Material | Description | Images |
|---|---|---|---|---|---|
| Araucarites[34] |
|
Isolated Cones | Conifer cones related with Araucariaceae | ||
| Brachyphyllum[32] |
|
Isolated Branched Shoots | Conifer shots related with Araucariaceae or Hirmeriellaceae. | ||
| Cunninghamites[32] |
|
|
Isolated Branched Shoots | Conifer shots related with Cupressaceae | |
| Dicotyledoneae indet.[32] | Indeterminate |
|
Isolated Leaves | ||
| Equisetites[52] |
|
|
Stems | Stems of Equisetales | |
| Eucalyptus[52] |
|
|
Isolated Leaves | Angiosperms leaves of Eucalypteae | |
| Filicales indet.[52] | Indeterminate |
|
Pinnae | ||
| Frenelopsis[1][32] |
|
|
Isolated Branched Shoots | A Conifer, member of Hirmeriellaceae. The most abundant foliar remain locally | |
| Ginkgoales? indet.[32] | Indeterminate |
|
Isolated Leaves | ||
| Magnoliaephyllum[34][52] |
|
|
Isolated Leaves | A Flowering Plant, suggested to belong to Magnoliaceae | |
| Pagiophyllum[32] |
|
Isolated Branched Shoots | Conifer shots related with Araucariaceae or Hirmeriellaceae. | ||
| Pinophyta indet.[53] | Indeterminate | Isolated Branched Shoots | Includes at least a new genus and 2 species | ||
| Papillopollis[15] |
|
|
Pollen | Resembles extant members of the wind-pollinated Juglandales-Myricales | |
| Sassafras[52] |
|
|
Isolated Leaves | Angiosperms leaves of Lauraceae | |
| Sphenolepis[32] |
|
|
Isolated Cones | Conifer cones related with Cupressaceae |














