Maaradactylus
Genus of anhanguerid pterosaur from the Early Cretaceous
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Maaradactylus is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period (Aptian to Albian stages) of the Romualdo Formation of northeastern Brazil.
| Maaradactylus Temporal range: | |
|---|---|
| Specimen MPSC R 2357, from the Araripe Basin, Brazil | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | †Pterosauria |
| Suborder: | †Pterodactyloidea |
| Family: | †Anhangueridae |
| Subfamily: | †Anhanguerinae |
| Genus: | †Maaradactylus Bantim et al., 2014 |
| Type species | |
| †Maaradactylus kellneri Bantim et al., 2014 | |
Discovery
Maaradactylus is based on the Museu Paleontologico de Santana do Cariri specimen MPSC R 2357, a skull, atlas, and axis discovered in 2010 in the Aptian—Albian-age Romualdo Formation of Sítio São Gonçalo, Santana do Cariri, Ceará, in the Araripe Basin of Brazil.
Maaradactylus was described by Renan Bantim and colleagues in 2014. The type species is Maaradactylus kellneri. The generic name refers to Maara, in the legends of the Cariri the daughter of a chief, by sorcery changed into a river monster with long teeth, devouring fishermen. The suffix ~dactylus is common in the names of pterosaurs and is derived from Greek δάκτυλος, daktylos, "finger", referring to the long (fourth) wing finger. The specific name honors Alexander Kellner, Brazil's foremost pterosaur expert.[1]
In 2019, the species Coloborhynchus spielbergi, also assigned as Anhanguera spielbergi, was reassigned to Maaradactylus by Megan L. Jacobs and colleagues as M. spielbergi.[2] This species was named after the filmmaker Steven Spielberg, the director of Jurassic Park by André J. Veldmeijer.[3] Later studies would retain it in the genus Anhanguera, though it has been noted the taxonomy of Araripe Basin anhanguerids requires further study.[4][5][6]
Description
Maaradactylus had one of the largest anhanguerid skulls from the Santana Group, and this indicated a wingspan estimate of about 6 meters (20 ft). This anhanguerid also has a relatively tall crest running along the midline of the premaxillary bones, which make up most of the upper beak region, and placements for 35 pairs of teeth in the upper jaws.[1]
Classification

In 2020 a study by Borja Holgado and Rubi Pêgas had recovered Maaradactylus as the sister taxon of Cearadactylus, both belonging to the subfamily Anhanguerinae. Their cladogram is shown on the right.[7]