Paleobiota of the Koshelevka Formation

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The Koshelevka Formation is a Permian formation in Russia containing the Tshekarda lagerstätte, which preserves over 200 insect species.[1]

Holometabola

Holometabolans
Genus Species Higher taxon Notes Images
Sylvacoleus S. richteri, S. sharovi Tshekardocoleidae[2] Has a relatively long abdomen, named after the Sylva River[3]
Isolated elytron of S. richteri
Sylvacoleodes S. admirandus Tshekardocoleidae Resembles Sylvacoleus, but has a narrower head[4]
Tshekardocoleus T. magnus Tshekardocoleidae[2] Type genus of the family, wing venation quite similar to the hypothetical ancestral beetle[3]
Cavalarva C. caudata Holometabola incertae sedis Shares characteristics with larvae from numerous holometabolan orders[1]

Paraneoptera

Paraneopterans
Genus Species Higher taxon Notes Images
Graticladus G. apiatus[1] Ischnoneuridae (Hypoperlida) Placed in Hypoperlida, which was reclassified as stem-Paraneoptera in 2017[5]
Tshekardoclardus T. sparsus[1] Tococladidae Placed in Hypoperlida, which was reclassified as stem-Paraneoptera in 2017[5]

Polyneoptera (excluding Notoptera)

Non-Notopteran Polyneopterans
Genus Species Higher taxon Notes Images
Arachnophasma A. scurra Permophasmatidae Oldest stick insect known, has very long legs and a short body[1]
Uraloblatta U. insignis Archimylacridae ("Eoblattodea")[6] Has a relatively long oviscapt[7]

Notoptera

Notopterans
Genus Species Higher taxon Notes Images
Artinska A. infigurabilis[8] Lemmatophoridae
Bardapteron B. ovale Atactophlebiidae Formerly placed with paleodictyopterans, known from the less diverse Barda locality as opposed to Chekarda[8]
Chelopterum C. ultimum Chelopteridae[1]
Cucullistriga C. cucullata Idelinellidae Named after the hood-like pronotum covering its head[9]
Culiciforma C. formosa Lemmatophoridae Similar to Megorkhosa[8]
Czekardia C. blattoides Eoblattida incertae sedis Preserves stripes and spots on its tergites, only known from a nymph[10]
Depressopterum D. bardum Liomopteridae[8]
Euryptilodes E. commatulus Euryptilonidae Known from wings and exuviae of nymphs[8]
Iblatta I. attrepida Eoblattida incertae sedis Only known from a nymph, has quite short legs and a small head[10]
Jubala J. pectinata Euryptilonidae Formerly placed within Sylvardembia[1]
Khosarophlebia K. sylvaensis Liomopteridae Formerly known from a wing fragment, but had a complete specimen discovered later[8]
Kishertia K. tricubitalis Pinideliidae Distinguished from Pinidelia by wing venation and size[8]
Koshelevka K. megakhosaroides Doubraviidae Formerly included in Cerasopterum[11]
Kungurocauda K. spinosa Sylviaphlebiidae Second-smallest member of the family behind Sylvophenoptera[8]
Kungurolioma K. cancellata Liomopteridae
Lemmatonympha L. gracilis, L. gracilissima Lemmatophoridae Identifiable to family based on its tarsi and cerci[12]
Liomopterella L. kungurica Liomopteridae[8]
Micaidelia M. minutissima Ideliidae Similar to Anaidelia[8]
Neprotembia N. truncata Permulidae[13] Similar to Neraphidia[8]
Neraphidia N. mitis, N. rigida Neraphidiidae[14] Formerly only known from part of a forewing, but a complete specimen was discovered later[8]
Novokshonovus N. ignoratus Atactophlebiidae Related to Kirkorella[1]
Ornaticosta O. novokshonovi Liomopteridae[8]
Parakhosara P. coalita Megakhosaridae[8]
Parasheimia P. truncata, P. rotundata Sheimiidae Larger than Sheimia and has a wider pronotum[8]
Parastenaropodites P. stirps Mesorthopteridae[1]
Parasylvaella P. umbra, P. minor Liomopteridae[8]
Parasylviodes P. tetracladus Liomopteridae[8]
Paratillyardembia P. sepicolorata Protembiidae[15] Has an unusual pronotum[16]
Parmaptera P. permiana Parmapteridae Placed within its own monotypic family[1]
Pectinokhosara P. sylvardembiodes Megakhosaridae Has elongated wings[8]
Permedax P. effertus Tillyardembiidae Differs from all other known grylloblattid nymphs by its elongate body,[8] but this is also a character of adult tillyardembiids[12]
Permula P. tshekardensis, P. minor Permulidae Formerly placed within Sojanopermula,[8] with Permula itself being a replacement name for Allicula.[13]
Permoshurabia P. kungurica Geinitziidae One of the earliest geinitziids[1]
Permostriga P. augustalis Idelinellidae[9]
Pseudosheimia P. caudata Sheimiidae Has longer cerci than Sheimia[8]
Scutistriga S. scutata Idelinellidae Named after its shield-like pronotum[9]
Sheimia S. tshekardensis Sheimiidae Bears larger eyes than other Sheimia species[8]
Strigulla S. cuculiophoris Idelinellidae Formerly placed within Euryptilon[9]
Suksunus S. bicodex Cacurgidae Similar to Kochopteron[11]
Sylvaclinicus S. echinatus Grylloblattida incertae sedis Only known from nymphs[8]
Sylvafossor S. forcipatus Probnidae Has unusual earwig-like cerci[8]
Sylvamicropteron S. harpax Grylloblattida incertae sedis An unusual micropterous grylloblattid[8]
Sylvaphlebia S. tuberculata, S. fucata Sylvaphlebiidae Has a pigmented wing strip[8]
Sylvaprisca S. gravis, S. forta, S. focaleata[8] Lemmatophoridae Named after the lemmatophorid genus Paraprisca[17]
Sylvardembia S. matura, S. tamaena Sylvardembiidae[16]
Sylvastriga S. miranda[8] Idelinellidae
Sylvictor S. major Liomopteridae One of the largest notopterans from Koshelevka[8]
Sylviodes S. perloides Mesorthopteridae[18] Formerly placed in Ideliidae[8]
Sylvonympha S. tshekardensis Grylloblattida incertae sedis Unusually for a grylloblattid, the larva has gills[12]
Sylvophenoptera S. perlongata, S. fimbriata Sylvaphlebiidae Resembles Phenopterum but has a longer ovipositor[8]
Tillyardembia T. antennaeplana, T. ravisedorum[19] Tillyardembiidae Most common insects from Koshelevka alongside the earliest insects with preserved pollen attached to them[20]
Tshekardanympha T. lienterica, T. bardensis Grylloblattida incertae sedis Split from Tshekardites[12]
Tshekardeigma T. rasnitsyni Epideigmatidae[21]
Tshekardelia T. media Ideliidae One of the first ideliids known from a well-preserved body[22]
Tshekardites T. comitialis Grylloblattida incertae sedis[8] First established as a nymphal form taxon, before all other species were split out[12]
Tshekardomina T. maculatus, T. imbecilla, T. imbecillissima, T. subincurvata[8] Tshekardominidae (Grylloblattida) Type specimen preserves several eggs inside the abdomen, rest of specimens are all female (possibly suggesting flightless males?)[23]
Tshekardophlebia T. capitata Sylvaphlebiidae Resembles Sylvaphlebia but has a larger head[8]
Tshekardushka T. artenatis Reculida incertae sedis Only known from a nymph. Preserves an oval impression, likely representing a bolus of inorganic material[10]
Uralioma U. maxima Liomopteridae Incompletely preserved, may belong to a separate genus[8]
Uraloprisca U. uralica Lemmatophoridae Formerly included in Paraprisca[24]
Visherifera V. sylvaensis[8] Visheriferidae Originally described from a late Permian locality in the Solikamsk Formation[25]

Misc. Arthropoda

Plants

References

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