Pupilla
Genus of gastropods
From Wikipedia, the free encyclopedia
Pupilla is a genus of minute air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the subfamily Pupillinae of the family Pupillidae.[2]
| Pupilla Temporal range: | |
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| A right side view of a live Pupilla muscorum | |
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| Six shells of Pupilla muscorum, scale bar in mm | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| Order: | Stylommatophora |
| Superfamily: | Pupilloidea |
| Family: | Pupillidae |
| Subfamily: | Pupillinae |
| Genus: | Pupilla Fleming, 1828[1] |
| Type species | |
| Pupa marginata Draparnaud, 1801 | |
| Synonyms | |
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Shells of Pupilla species are known from terrestrial Cenozoic strata dating back to the Oligocene until the Holocene period.[3]
Distribution
This genus occurs in Eurasia, northern Africa and North America.[4]
Species
Species within the genus Pupilla include:
- † Pupilla aeoli (Hilber, 1884)
- Pupilla afghanicum Solem, 1979
- Pupilla alabiella Schileyko, 1984
- Pupilla alaskensis Nekola & Coles, 2014
- † Pupilla almuhambetovae Prysjazhnjuk, 2017
- Pupilla alpicola (Charpentier, 1837)[4]
- Pupilla annandalei Pilsbry[5]
- Pupilla anzobica Izzatullaev, 1970
- Pupilla armeniaca (Issel, 1865)
- † Pupilla bargi Prysiazhniuk, 2017
- † Pupilla belokrysi Steklov, 1966
- Pupilla bigranata (Rossmässler 1839)[4] (taxon inquirendum)
- Pupilla bipatulata Akramowski, 1943
- † Pupilla bituberculata H.-J. Wang & S.-Y. Guo, 1991
- † Pupilla blainvilleana (Dupuy, 1850)
- Pupilla blandii E. S. Morse, 1865 – Rocky Mountain column
- † Pupilla bogdanovkaense Prysiazhniuk, 2017
- Pupilla brevicostis (Benson, 1849)
- Pupilla calacharicus (Boettger, 1886) [citation needed]
- † Pupilla capitani (de Morgan, 1920)
- † Pupilla crossei (Michaud, 1862)
- † Pupilla cupella O. Boettger, 1889
- † Pupilla diezi (Flach, 1890)
- Pupilla diopsis (Benson, 1863)
- Pupilla duplicata (Preston, 1911)
- Pupilla ficulnea (Tate, 1894)[6]
- Pupilla gallae Tzvetkov, 1940
- Pupilla goniodon Pilsbry, 1927
- Pupilla guadalupensis Pilsbry, 1927
- Pupilla gutta (Benson, 1864)
- Pupilla hebes (Ancey 1881) – crestless column
- Pupilla hokkaidoensis Nekola, Coles & S. Chiba, 2014
- Pupilla hudsoniana Nekola & Coles, 2014
- Pupilla iheringi Suter, 1900
- † Pupilla impressa (F. Sandberger, 1862)
- Pupilla inequidenta Schileyko & Almuhambetova, 1979
- † Pupilla inermis Russell, 1931
- Pupilla inops (Reinhardt, 1877)
- † Pupilla iratiana (Dupuy, 1850)
- Pupilla khunjerabica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
- Pupilla kyrostriata Walther & Hausdorf, 2014
- Pupilla limata Schileyko, 1984
- Pupilla loessica Ložek, 1954[7]
- † Pupilla michaudi Wenz, 1919
- † Pupilla mlomnickii (Friedberg, 1905)
- Pupilla muscorum (Linnaeus, 1758)[4] – widespread column – type species
- † Pupilla mutabilis Steklov, 1966
- Pupilla obliquicosta Smith, 1892 – extinct[8]
- Pupilla paraturcmenica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
- † Pupilla parvula (Deshayes, 1863)
- † Pupilla perlabiata Gottschick & Wenz, 1919
- † Pupilla poltavica O. Boettger, 1889
- Pupilla profundotuberculata H.-J. Wang & S.-Y. Guo, 1991
- Pupilla pupula[9]
- Pupilla quxuensis D.-N. Chen & G.-Q. Zhang, 2001
- † Pupilla retusa (F. Sandberger, 1858)
- † Pupilla roberti Prysiazhniuk, 2017
- Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861)
- Pupilla satparanica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
- † Pupilla selecta (Thomä, 1845)
- † Pupilla selectiformis (Simionescu & Barbu, 1940)
- Pupilla seriola (Benson, 1861)
- † Pupilla shantungensis Yen, 1969
- Pupilla simplexa Y.-T. Li, 1985
- Pupilla sonorana (Sterki, 1899) – three-tooth column
- † Pupilla staszicii (Łomnicki, 1886)
- † Pupilla steinheimensis (K. Miller, 1900)
- Pupilla sterkiana (Pilsbry, 1890)
- Pupilla sterrii (Forster & Voith, 1840)[4]
- Pupilla striopolita Schileyko, 1984
- † Pupilla subquadridens (A. d'Orbigny, 1852)
- Pupilla syngenes (Pilsbry, 1890) – top-heavy column
- Pupilla tetrodus (Boettger, 1870)
- Pupilla triplicata (Studer, 1820)[4]
- † Pupilla triplicatoidea Steklov, 1966
- † Pupilla tschumakovi Prysjazhnjuk, 2017
- Pupilla turcmenica (O. Boettger)[5]
- † Pupilla zeisslerae Schlickum, 1975
- Pupilla ziaratana Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
Synonyms
- Pupilla barrackporensis Gude, 1914: synonym of Insulipupa malayana (Issel, 1874) (junior synonym)
- Pupilla cinghalensis Gude, 1914: synonym of Indopupa cinghalensis (Gude, 1914) (original combination)
- Pupilla cryptodon (Heude, 1880): synonym of Gibbulinopsis cryptodon (Heude, 1880) (unaccepted combination)
- Pupilla cupa (Jan 1832): synonym of Pupilla muscorum (Linnaeus, 1758) (junior synonym)
- Pupilla draparnaudi Leach [in Turton], 1831: synonym of Lauria cylindracea (Da Costa, 1778)
- Pupilla floridana Dall, 1885: synonym of Gastrocopta pentodon (Say, 1822) (junior synonym)
- Pupilla fontana (Krauss, 1848): synonym of Gibbulinopsis fontana (F. Krauss, 1848) (superseded combination)
- Pupilla gracilis Izzatullaev, 1970: synonym of Gibbulinopsis gracilis (Izzatullaev, 1970) (original combination)
- Pupilla hudsonianum [sic]: synonym of Pupilla hudsoniana Nekola & Coles, 2014 (wrong gender agreement of specific epithet)
- Pupilla muscerda (Benson, 1853): synonym of Microstele muscerda (Benson, 1853) (unaccepted combination)
- Pupilla pratensis (Clessin, 1871):[4] synonym of Pupilla alpicola (Charpentier, 1837) (unaccepted > junior subjective synonym)
- † Pupilla quadrigranata (F. Sandberger, 1858): synonym of † Pupilla selecta (Thomä, 1845) (junior synonym)
- Pupilla reibischi Dall & Ochsner, 1928: synonym of Gastrocopta reibischi (Dall & Ochsner, 1928) (original name)
- Pupilla salmensis (W. T. Blanford & H. F. Blanford, 1861): synonym of Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861) (misspelling of original name.)
- Pupilla signata (Mousson, 1873):[5] synonym of Gibbulinopsis signata (Mousson, 1873)
- Pupilla stoneri Chamberlin & Jones, 1929: synonym of Gastrocopta pilsbryana (Sterki, 1890)
- † Pupilla submuscorum Gottschick & Wenz, 1919: synonym of † Gibbulinopsis submuscorum (Gottschick & Wenz, 1919)
- Pupilla wolfae Tzvetkov, 1940: synonym of Pupilla sterrii (Voith, 1840) (junior synonym)
