Splenial

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Hemimandible of Trimerorhachis; the splenial (labeled "sp") is visible on the interior surface
Skull and jaws diagram of the primitive synapsid Dimetrodon, showing location of the splenial

The splenial is a small bone in the lower jaw of reptiles, amphibians and birds, usually located on the lingual side (closest to the tongue) between the angular and surangular.[1][2] The splenial and coronoid are together known as the paradental elements.[3] In non-tetrapod vertebrates, the splenial is known as the first infradentary and forms part of the infradentary series along with the postsplenial and angular.[4][5] The earliest known taxon with infradentary bones is the placoderm Entelognathus, while they were absent in the earlier-diverging Qilinyu.[6] Splenials are present in non-mammalian synapsids, but lost in mammals, though there are some claims of a vestigial splenial in some early mammals.[3]

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