Umbra (fish)
Genus of fishes
From Wikipedia, the free encyclopedia
Umbra is a genus of mudminnows native to Europe and North America. It is the only extant member of the family Umbridae.
| Umbra Temporal range: | |
|---|---|
| |
| European mudminnow, Umbra krameri | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Division: | Teleostei |
| Order: | Salmoniformes |
| Family: | Umbridae |
| Genus: | Umbra W. H. Kramer, 1777 |
| Type species | |
| Umbra krameri Walbaum, 1792 | |
Phylogenetic evidence suggests the North American and European clades split between the Late Cretaceous or early Paleogene, making this a rather ancient group.[1]
Species
The currently recognized species in this genus are:[2]
- Umbra krameri Walbaum, 1792 (European mudminnow)
- Umbra limi (Kirtland, 1840) (central mudminnow)
- Umbra pygmaea (DeKay, 1842) (eastern mudminnow)
The following fossil species are also known:
- †Umbra euronota Schwarzhans et al., 2024 - Middle Miocene of Ukraine [otolith][3]
- †Umbra irtyshensis Sytchevskaya, 1968[4]
- †Umbra longidorsalis Böhme, 2004 - Early Miocene of the Czech Republic (Most Formation)[5]
- †Umbra oderiensis Oberhlová, 1978 - Late Oligocene/Early Miocene of the Czech Republic[5]
- †Umbra perpusilla (Agassiz, 1839) - Middle Miocene of Germany[6]
- †Umbra praekrameri Weinfurter, 1950 - Late Miocene of Austria [otolith][3]
- †Umbra prochazkai Oberhlová, 1978 - Late Oligocene of the Czech Republic[1][5]
Indeterminate Umbra scales are known from the Late Eocene/Early Oligocene of the Czech Republic.[7]
