Villar del Arzobispo Formation

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OverliesOolitic limestones of the Higuerueles Formation
ThicknessGreatly variable;108–1,383 m (354–4,537 ft)
Villar del Arzobispo Formation
Stratigraphic range: Kimmeridgian-early Berriasian?
An ornithopod trackway at the Las Cerradicas site, Villar del Arzobispo Formation. Photographed around 2013.[1]
TypeGeological formation
UnderliesAguilar del Alfambra Formation
OverliesOolitic limestones of the Higuerueles Formation
ThicknessGreatly variable;108–1,383 m (354–4,537 ft)
Lithology
PrimaryMudstone
OtherLimestone, sandstone, conglomerate
Location
Coordinates40°30′N 0°48′W / 40.5°N 0.8°W / 40.5; -0.8
Approximate paleocoordinates31°24′N 8°36′E / 31.4°N 8.6°E / 31.4; 8.6
RegionAragón, Teruel, Valencia
CountrySpain
ExtentMaestrazgo Basin
Type section
Named forVillar del Arzobispo
Villar del Arzobispo Formation is located in Spain
Villar del Arzobispo Formation
Villar del Arzobispo Formation (Spain)

The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Middle Berriasian, but more recent work suggests a Kimmeridigan-Late Tithonian, possibly dating to the Early Berriasian in some areas.[2][3][4] The Villar del Arzobispo Formation's age in the area of Riodeva in Spain has been dated based on stratigraphic correlations as middle-upper Tithonian, approximately 145-141 million years old. In the area of Galve, the formation potentially dates into the earliest Cretaceous.[2] Isotope dating gives the formation a Late Kimmeridgian date of 152.5-151 million years old.[5]

Most of the unit consists of siliciclastic mudstone, however the lower portion of the formation is dominated by bioclastic, oolitic and peloidal limestone, while channelized sandstone and conglomerate is found in the middle portion of the unit. While the lower part of the formation was deposited in an inner carbonate platform, the upward gradation into mudstone in the middle and upper portions of the formation represents a change in depositional environment to paralic and alluvial plain conditions.[3][6]

Dinosaur remains are found throughout the unit, but are more abundant and better preserved in the terrestrially deposited middle-upper sections. Remains of the stegosaurid Dacentrurus were recovered in the Barranco Conejero locality in this formation.[7] Over the years, other stegosaurian remains have been discovered in this formation, but none have yet been referred to a specific genus.[8] Turiasaurs and brachiosaurids are also known from the formation. The formation is also well known for its fossil footprints, representing many different taxa.

Dinosaurs

Ornithischians

Ornithopods
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Ornithischians of the Villar del Arzobispo Formation
Taxon Species Localities Material Notes Images
Ankylopollexia Indeterminate By Pass fossil site (Alpuente, Valencia) a left dentary tooth, an anterior caudal vertebra, a posterior caudal vertebra A small-sized ankylopollexian[9]
VS-1 site (Veguillas de la Sierra, Teruel) an anterior caudal vertebra A large-sized ankylopollexian[9]
aff. Camptosaurus[10] C. sp. Fuentecillas fragmentary remains: an anterior cervical centrum; an anterior dorsal centrum; a dorsosacral centrum; four sacral centra; a caudosacral centrum; two anterior caudal centra; three medial caudal centra; and a distal fragment from the left humerus An ornithopod. The Fuentecillas specimen is seen to be closer to Camptosaurus than to Draconyx, hence its tentative placement within the Camptosaurus genus.
Camptosaurus
Oblitosaurus
Dryosauridae[9] Indeterminate RD-61 site (Riodeva, Teruel) an anterior caudal vertebra
Oblitosaurus[11] O. bunnueli Barrihonda-El Humero "a dentary tooth, an ungual pollex of the manus, and an almost complete left hindlimb" Basalmost ankylopollexian, sister taxon to Draconyx.
Ornithopoda Indeterminate Cambrillas a pedal ungual phalanx[9]
Cerrito del Olmo a cervical centrum and two caudal vertebrae[12]
Stegosaurs
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Stegosaurs of the Villar del Arzobispo Formation
Taxon Species Localities Material Notes Images
Dacentrurus[13] D. armatus
  • Barrihonda–El Humero
  • San Cristóbal
  • Cerrito del Olmo
several different elements Subjective senior synonym of Miragaia.
Dacentrurus
Dacentrurinae[14] Indeterminate
  • Barranco del Curro
  • El Balsón
two partial skeletons Indeterminate as it cannot be compared with Miragaia, assuming Miragaia is distinct.
Deltapodus[15] D. ibericus El Castellar trackway on tidal limestone Belongs to a large stegosaur, possibly related to Dacentrurus.

Sauropods

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Sauropods of the Villar del Arzobispo Formation
Taxon Species Localities Material Notes Images
Aragosaurus[16] A. ischiaticus Las Zabacheras partial postcranial skeleton A basal macronarian previously thought to come from the El Castellar Formation.
Aragosaurus
Turiasaurus
Diplodocidae[17] Indeterminate Pino de Jarque 2 ilium Similar to Diplodocus and Barosaurus.
Diplodocinae[17] Indeterminate El Carrillejo anterior caudal vertebra
Galvesaurus[18] G. herreroi Cuesta Lonsal-1 partial skeleton Was once thought to be a turiasaur or basal macronarian, but now known to be a brachiosaurid. Possibly a junior synonym of Lusotitan.[19]
Losillasaurus[20] L. giganteus
  • La Cañada
  • San Lorenzo
caudal vertebrae, and two partial skeletons A large turiasaur closely related to Turiasaurus.
Macronaria[17] Indeterminate
  • La Quineta 1
  • Las Viñas
  • San Lorenzo
caudal vertebrae
Turiasaurus[21][20] T. riodevensis
  • Barrihonda–El Humero
  • Corral de la Cautiva
several postcranial and cranial elements A large turiasaur, similar to Losillasaurus.
Turiasauria[17] Indeterminate Puntal de Santa Cruz several postcranial elements Indeterminate turiasaur material originally referred to Turiasaurus, but does not share any unique features with the genus[22]

Theropods

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Theropods of the Villar del Arzobispo Formation
Taxon Species Localities Material Notes Images
Allosauridae[23] Indeterminate Barrihonda-El Humero 5 teeth Typical allosaurid teeth.
Dromaeosauridae[23] Indeterminate
  • Barrihonda-El Humero
  • Puntal de Santa Cruz
7 teeth Possibly belonging to a velociraptorine.
Iberosauripus[24] I. grandis El Castellar trackway Likely belonged to a megalosaurid.
Megalosauridae[23][24] Indeterminate
  • Carretera
  • La Fonseca
  • Masía de la Hoya Alta
  • RD-39
teeth Tentatively referred to the Megalosauridae. Some of the largest theropod teeth of Spain.

Reptiles

Turtles of the Villar del Arzobispo Formation
Taxon Species Localities Material Notes Images
Riodevemys R. inumbragigas Barrihonda–El Humero A pleurosternid freshwater turtle

Correlation

See also

References

Further reading

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