↑ N. W. Thornhill, ed(1993).Inbreeding and outbreeding in captive populations of wild animal species In The Natural History of Inbreeding and Outbreeding.Chicago:The University of Chicago Press.pp.pp. 352-374
↑ “Estimates of lethal equivalents and the cost of inbreeding in mammals.”.Conservation Biology2: 185-198.(1988).
↑ D. E. Wildt;M. Bush, K. L. Goodrowe, C. Packer, A. E. Pusey, J. L. Brown, P. Joslin, S. J. O. Brien(1987).“Reproductive and genetic consequences of founding isolated lion populations.”.Nature329: 328-331.
↑ C. Packer;A. E. Pussey(1993).N. W. Thornhill.ed.Dispersal, kinship, and inbreeding in African lions In The Natural History of Inbreeding and Outbreeding.Chicago:The University of Chicago Press.pp.pp. 352-374