Paleobiota of Burmese amber

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up to the end of 2023 can be found in Ross (2024).^[1]

[1]

Family	Genus	Species	Authority	Year described	Notes
Calymperaceae	†Calymperites	†Calymperites burmensis^[3]	Heinrichs 'et al.	2014	A Calymperaceous dicranale moss
		†Calymperites chenianus^[4]	Li in Tan et al.	2025	A Calymperaceous dicranale moss
		†Calymperites heinrichsianus^[5]	Li & Wang in Li et al.	2025	A Calymperaceous dicranale moss
		†Calymperites marginatus^[5]	Li et al.	2025	A Calymperaceous dicranale moss
		†Calymperites proboscideus^[4]	Li in Tan et al.	2025	A Calymperaceous dicranale moss
		†Calymperites striatus^[5]	Li et al.	2025	A Calymperaceous dicranale moss
Incertae sedis	†Ditrichites	†Ditrichites aristatus^[4]	Li in Tan et al.	2025	A member of Dicranales sensu lato.

Family	Genus	Species	Authority	Year described	Notes
Incertae sedis	†Vetiplanaxis	†Vetiplanaxis espinosus^[6]	Hedenäs, Heinrichs, & Schmidt	2014	A hypnodendralean moss
		†Vetiplanaxis longiacuminatus^[6]	Hedenäs, Heinrichs, & Schmidt	2014	A hypnodendralean moss
		†Vetiplanaxis oblongus^[6]	Hedenäs, Heinrichs, & Schmidt	2014	A hypnodendralean moss
		†Vetiplanaxis obtusus^[7]	Li et al.	2022	A hypnodendralean moss
		†Vetiplanaxis pyrrhobryoides^[6]	Bell	2007	A hypnodendralean moss

Family	Genus	Species	Authority	Year described	Notes	image
Selaginellaceae	Selaginella	†Selaginella alata^[8]	Li & Wang	2022	A clubmoss
		†Selaginella amplexicaulis^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella amplexicaulis
		†Selaginella aurita^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella aurita
		†Selaginella ciliifera^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella ciliifera
		†Selaginella cretacea^[10]	Li et al.	2022	A Selaginella subg. Stachygynandrum clubmoss^[9]	Selaginella cretacea
		†Selaginella grimaldii^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella grimaldii
		†Selaginella heinrichsii^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella heinrichsii
		†Selaginella heterosporangiata^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella heterosporangiata
		†Selaginella isophylla^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Ericetorum clubmoss	Selaginella isophylla
		†Selaginella konijnenburgiae^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella konijnenburgiae
		†Selaginella longifimbriata^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella longifimbriata
		†Selaginella minutissima^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella minutissima
		†Selaginella obscura^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella obscura
		†Selaginella ohlhoffiorum^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella ohlhoffiorum
		†Selaginella ovoidea^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella ovoidea
		†Selaginella patrickmuelleri^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella patrickmuelleri
		†Selaginella pellucida^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella pellucida
		†Selaginella tomescui^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation
		†Selaginella villosa^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella villosa
		†Selaginella wangboi^[9]	A. R. Schmidt & L. Regalado	2022	A clubmoss of uncertain subgenus affiliation	Selaginella wangboi
		†Selaginella wangxinii^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Stachygynandrum clubmoss	Selaginella wangxinii
		†Selaginella wunderlichiana^[9]	A. R. Schmidt & L. Regalado	2022	A Selaginella subg. Ericetorum clubmoss	Selaginella wunderlichiana

Family	Genus	Species	Authority	Year described	Notes	image
Frullaniaceae	Frullania	†Frullania baerlocheri^[11]	Heinrichs et al.	2011	A frullaniacious liverwort. Frullania pinnata (2017) deemed a jr synonym.^[12]
		Frullania cretacea^[13]	Hentschel, Schmidt & Heinrichs	2009	A frullaniacious liverwort
		Frullania kachinensis^[14]	Li et al.	2021	A frullaniacious liverwort
		Frullania partita^[15]	Li et al.	2020	A frullaniacious liverwort
	†Protofrullania	†Protofrullania cornigera^[16]	Heinrichs	2017	A frullaniacious liverwort
Lepidolaenaceae	†Gackstroemia	†Gackstroemia cretacea^[17]	Heinrichs, Schäfer-Verwimp, Feldberg, & A.R. Schmidt	2017	A lepidolaenacious liverwort
Radulaceae	Radula	†Radula cretacea^[18]	Bechteler, Renner, Schäfer-Verwimp, & Heinrichs	2017	A Radulaceae liverwort	Radula cretacea
		†Radula heinrichsii^[19]	Feldberg et al.	2021	A radulaceous liverwort
		†Radula patrickmuelleri^[20]	Feldberg, Schäfer-Verwimp & Renner	2022	A radulaceous liverwort
		†Radula tanaiensis^[20]	Feldberg, Schäfer-Verwimp & Renner	2022	A radulaceous liverwort

Family	Genus	Species	Authority	Year described	Notes	image
Thyrsopteridaceae	Thyrsopteris	†Thyrsopteris cretacea^[21]	Li et al.	2020	A thyrsopteridacious tree-fern
Thyrsopteridaceae	Thyrsopteris	†Thyrsopteris cyathindusia^[22]	Zhang et al.	2021	A thyrsopteridacious tree-fern

Family	Genus	Species	Authority	Year described	Notes	image
Cystodiaceae	Cystodium	†Cystodium sorbifolioides^[23]	Regalado et al.	2017	A Cystodiacious fern	Cystodium sorbifolioides
Dennstaedtiaceae	†Cladarastega	†Cladarastega burmanica^[24]	Poinar	2021	A dennstaedtiaceous fern
	†Krameropteris	Krameropteris calophyllum^[25]	Li in Li & Meng	2025	A dennstaedtiacious fern
	†Krameropteris	Krameropteris resinatus^[26]	Schneider, Smith, & Heinrichs	2016	A dennstaedtiacious fern
	Microlepia	†Microlepia burmasia^[27]	Long, Wang & Shi in Long et al.	2022	A species of Microlepia.
Hymenophyllaceae	†Hymenophyllites	†Hymenophyllites angustus^[8]	Li & Wang	2022	A filmy fern
		†Hymenophyllites kachinensis^[8]	Li & Wang	2022	A filmy fern
		†Hymenophyllites setosus^[8]	Li & Wang	2022	A filmy fern
Lindsaeaceae	†Proodontosoria	†Proodontosoria myanmarensis^[28]	Li et al.	2020	A lindsaeacious fern
Marsileaceae	†Marsileaceaephyllum	†Marsileaceaephyllum ciliatum^[29]	Wang et al.	2021	A marsileaceous fern.
Pteridaceae	†Heinrichsia	†Heinrichsia cheilanthoides^[30]	Regalado, Schmidt, Krings & Schneider	2019	A pteridacious fern
Incertae sedis	†Cretacifilix	†Cretacifilix fungiformis^[31]	Poinar & Buckley	2008	A eupolypod fern
Incertae sedis	†Holttumopteris	†Holttumopteris burmensis^[32]	Regalado, Schneider, Krings, & Heinrichs	2018	A fern of uncertain placement

Family	Genus	Species	Authority	Year described	Notes	image
Cornaceae	†Eoëpigynia	Eoëpigynia burmensis^[33]	Poinar, Chamber & Buckley	2007	A cornalean flower
Nyssaceae	†Nyssapollenites	Sp.^[34]	N/A	2019	A nyssaceous pollen morphotype

Family	Genus	Species	Authority	Year described	Notes
Atherospermataceae	†Dasykothon	Dasykothon leptomiscus^[36]	Poinar & Chambers	2020	A lauralean flowering plant of uncertain phylogenetic placement.
Hernandiaceae	†Zygadelphus	†Zygadelphus aetheus^[37]	Poinar & Chambers	2019	A possible lauralian of uncertain phylogenetic placement
Lauraceae	†Cascolaurus	†Cascolaurus burmitis^[38]	Poinar	2017	A lauracious angiosperm
Monimiaceae	†Palaeoanthella	†Palaeoanthella huangii^[39]	Poinar & Chambers	2005	A possible monimiacious angiosperm
	†Setitheca	†Setitheca lativalva^[40]	Poinar & Chambers	2018	A possible monimiacious angiosperm
	†Thymolepis	†Thymolepis toxandra^[41]	Chambers & Poinar	2020	A possible monimiacious angiosperm
	†Valviloculus	†Valviloculus pleristaminis^[42]	Poinar et al.	2020	A possible monimiacious angiosperm
Incertae sedis	†Cyathitepala	†Cyathitepala papillosa^[43]	Poinar & Chambers	2020	A lauralean angiosperm of uncertain affinity
	†Dispariflora	†Dispariflora robertae^[44]	Poinar & Chambers	2019	A possible lauralean flower
	†Jamesrosea	Jamesrosea burmensis^[45]	Crepet et al.	2016	A mosaic laralean genus

Family	Genus	Species	Authority	Year described	Notes
Cunoniaceae	†Tropidogyne	†Tropidogyne euthystyla^[48]	Poinar, Chambers & Vega	2021	A small flowered cunoniacious angiosperm.
		†Tropidogyne lobodisca^[49]	Poinar & Chambers	2019	A cunoniacious angiosperm.
		†Tropidogyne pentaptera^[50]	Poinar & Chambers	2017	A cunoniacious angiosperm.
		†Tropidogyne pikei^[51]	Chambers, Poinar & Buckley	2010	A cunoniacious angiosperm.
Incertae sedis	†Lachnociona	†Lachnociona camptostylus^[52]	Poinar & Chambers	2018	An oxalidalean angiosperm of uncertain affinity
Incertae sedis	†Lachnociona	†Lachnociona terriae^[53]	Poinar, Chambers & Buckley	2008	An oxalidalean angiosperm of uncertain affinity

family	Genus	Species	Authority	Year described	Notes	image
Poaceae	†Programinis	†Programinis burmitis^[54]	Poinar	2004	A poalean of uncertain placement
Poaceae	†Programinis	Programinis laminatus^[54]	Poinar	2004	A poalean of uncertain placement

Family	Genus	Species	Authority	Year described	Notes	image
Rhamnaceae	†Eophylica	†Eophylica priscastellata^[55]	Shi et al.	2022	A rhamnaceous floral morphotaxon Electrophycus astroplethus^[56] Deemed a jr synonym^[57]
Rhamnaceae	Phylica	†Phylica piloburmensis^[55]	Shi et al.	2022	A rhamnaceous floral morphotaxon.

Genus	Species	Authority	Year described	Notes	image
†Antiquifloris	†Antiquifloris latifibris^[58]	Poinar, Buckley & Chen	2016	An angiosperm of uncertain placement	Antiquifloris latifibris
†Chainandra	†Chainandra zeugostylus^[59]	Poinar & Chambers	2020	An angiosperm of uncertain placement
†Chenocybus	†Chenocybus allodapus^[60]	Poinar	2018	An angiosperm of uncertain placement
†Exalloanthum	↑Exalloanthum burmense^[60]	Poinar	2018	An angiosperm of uncertain placement, originally named Diaphoranthus burmensis^[61]
†Lijinganthus	†Lijinganthus revoluta^[62]	Liu et al.	2018	A core eudicot of uncertain placement	Lijinganthus revoluta
†Micropetasos	†Micropetasos burmensis^[63]	Poinar, Chambers, & Wunderlich	2004	An angiosperm of uncertain placement
†Rasenganus	†Rasenganus auricularus^[64]	Xing & Gu	2020	A possible epizoochorous fruit.
†Strombothelya	†Strombothelya monostyla^[65]	Poinar & Chambers	2019	An angiosperm of uncertain phylogenetic placement
†Strombothelya	†Strombothelya grammogyna^[65]	Poinar & Chambers	2019	An angiosperm of uncertain phylogenetic placement

Family	Genus	Species	Authority	Year described	Notes
Clavicipitaceae	†Palaeoclaviceps	†Palaeoclaviceps parasiticus^[66]	Poinar, Alderman & Wunderlich	2015	An ergot
Ophiocordycipitaceae	†Paleoophiocordyceps	†Paleoophiocordyceps coccophagus^[67]	Sung, Poinar & Spatafora	2008	A parasite of a scale insect
Incertae sedis	†Entropezites	†Entropezites patricii^[68]	Poinar & Buckley	2007	A hypermycoparasitic fungus
Incertae sedis	†Mycetophagites	†Mycetophagites atrebora^[68]	Poinar & Buckley	2007	A hypermycoparasitic fungus

Family	Genus	Species	Authority	Year described	Notes	image
Spheciophilaceae	†Philothysanus	†Philothysanus burmanicus^[71]	Poinar	2021	An ectoparasitic fungus
Spheciophilaceae	†Spheciophila	†Spheciophila adercia^[72]	Poinar	2016	An ectoparasitic fungus	Spheciophila adercia
Incertae sedis	†Palaeomycus	†Palaeomycus epallelus^[73]	Poinar	2018	A fungus belonging to the Pezizomycotina

Family	Genus	Species	Authority	Year described	Notes	image
Incertae sedis	†Burmacharon	†Burmacharon dunlopi^[77]	Hu et al.	2020		Burmacharon dunlopi
	†Kronocharon	†Kronocharon engeli^[78]	Wunderlich	2015	A tailless whip scorpion of uncertain placement.	Kronocharon engeli
		†Kronocharon longicalcaris^[78]	Wunderlich	2015	A tailless whip scorpion of uncertain placement.	Kronocharon longicalcaris
		†Kronocharon prendinii^[79]	Rossi & Merendino	2016	A tailless whip scorpion of uncertain placement. One subspecies described K. p. adornellae^[80]	Kronocharon prendinii

Family	Genus	Species	Authority	Year described	Notes
†Aliendiguetidae	Aliendiguetia	†Aliendiguetia praecursor^[81]	Wunderlich & Müller	2020	An aliendiguetid spider of uncertain affinities Possibly related to the Ochyroceratoidea.
†Alteruloboridae	†Alteruloborus	†Alteruloborus araneoides^[82]	Wunderlich in Wunderlich & Müller	2018	An alteruloborid spider.
Archaeidae	†Burmesarchaea	†Burmesarchaea alissa^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea bilongapophyses^[81]	Wunderlich & Müller	2020	An archaeid spider.
		†Burmesarchaea caudata^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea crassicaput ^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea crassichelae^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea gibber^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea gibberoides^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea gibbosa^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea grimaldii^[83]	(Penny)	2003	An archaeid spider, originally described as Afrarchaea grimaldii
		†Burmesarchaea longicollum^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea propinqua^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea pseudogibber^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea pustulata^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea quadrata^[83]	Wunderlich	2017	An archaeid spider
		†Burmesarchaea speciosa^[83]	(Wunderlich)	2008	An archaeid spider, Originally described as Lacunauchenius speciosus.^[84]
	†Eomysmauchenius	†Eomysmauchenius cretaceominimus^[85]	Peng et al.	2023	A planarchaeid^[86] palpimanoid spider.
		†Eomysmauchenius dubius^[83]	Wunderlich	2017	A planarchaeid^[86] palpimanoid spider.
		†Eomysmauchenius ovata^[82]^[86]	(Wunderlich)	2017	A planarchaeid^[86] palpimanoid spider, originally Planarchaea ovata
		Eomysmauchenius septentrionalis^[83]	Wunderlich	2008	A planarchaeid^[86] palpimanoid spider
	†Spiniarchaea	†Spiniarchaea aberrans^[86]	Wunderlich in Wunderlich & Müller	2021	An archaeid spider.
†Burmascutidae	†Burmascutum	†Burmascutum aenigma^[84]	Wunderlich	2008	A burmascutid spider.
†Burmascutidae	†Burmascutum	†Burmascutum brevis^[82]	Wunderlich	2018	A burmascutid spider.
†Burmathelidae	†Burmathele	†Burmathele biseriata^[83]	Wunderlich	2017	A burmathelid mesothele spider.
†Burmorsolidae	†Burmorsolus	Burmorsolus crassus^[83]^[81]	Wunderlich	2015	Transferred to the separate genus Pseudorsolus in 2017, but moved back to the genus Burmorsolus in 2020.
		Burmorsolus curvatus^[83]^[81]	(Wunderlich)	2017	Originally described as "Loxoderces" curvatus.
		Burmorsolus globosus^[81]	Wunderlich & Müller	2020
		Burmorsolus longembolus^[81]	Wunderlich & Müller	2020
		Burmorsolus longibulbus^[81]	Wunderlich & Müller	2020
		Burmorsolus longicymbium^[83]^[81]	(Wunderlich)	2017	Originally described as "Loxoderces" longicymbium.
		Burmorsolus longitibia^[86]	Wunderlich in Wunderlich & Müller	2021
		Burmorsolus nonplumosus^[87]	Wunderlich	2015	Probably a relative of trogloraptorids.^[81]
		Burmorsolus rectus^[83]^[81]	(Wunderlich)	2017	Originally described as "Loxoderces" rectus.
†Crassicephalidae	†Crassicephalus	†Crassicephalus parvibulbus^[86]	Wunderlich in Wunderlich & Müller	2021	A Deinopoidean spider.
†Cretaceothelidae	†Cretaceothele	†Cretaceothele lata^[87]	Wunderlich	2015	A cretaceothelid Mesothelaen spider.^[83]
†Cretamysmenidae	†Cretamysmena	†Cretamysmena fontana^[82]	Wunderlich in Wunderlich & Müller	2018	A cretamysmenid spider.
Ctenizidae	†Parvocteniza	†Parvocteniza parvula^[81]	Wunderlich & Müller	2020	A ctenizid spider.
Dipluridae?	†Alterphyxioschemoides	†Alterphyxioschemoides spicula^[86]	Wunderlich in Wunderlich & Müller	2021	A possible curtain-web spider.
Dipluridae?	†Phyxioschemoides	†Phyxioschemoides collembola^[87]	Wunderlich	2015	A spider of uncertain affinities,^[88] originally described as a curtain-web spider.

Family	Genus	Species	Authority	Year described	Notes	image
Ixodidae	Amblyomma	†Amblyomma birmitum^[113]	Chitimia-Dobler et al.	2017	An ixodid hard tick.
	Archaeocroton	†Archaeocroton kaufmani^[114]	Chitimia-Dobler, Mans & Dunlop	2022	An ixodid hard tick.
	Bothriocroton	†Bothriocroton muelleri^[114]	Chitimia-Dobler, Mans & Dunlop	2022	An ixodid hard tick.
	†Compluriscutula	†Compluriscutula vetulum^[115]	Poinar & Buckley	2008	An ixodid hard tick.
	†Cornupalpatum	†Cornupalpatum burmanicum^[116]	Poinar & Brown	2003	An ixodid tick	Cornupalpatum burmanicum
	Haemaphysalis	†Haemaphysalis cretacea^[117]	Chitimia-Dobler, Pfeffer & Dunlop	2018	A Haemaphysaline hard tick.
	Ixodes	†Ixodes antiquorum^[118]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2022	A tick, a species of Ixodes.
†Khimairidae	†Khimaira	†Khimaira fossus^[118]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2022	A tick combining a body resembling that of a soft tick with a basis capitulum more like that of a hard tick, assigned to a new family Khimairidae as a possible transitional form between soft and hard ticks.
Nuttalliellidae	†Deinocroton	Deinocroton bicornis^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick
		Deinocroton copia^[118]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2022	A nuttalliellid tick
		Deinocroton draculi^[120]	Peñalver, Arillo, Anderson & Pérez-de la Fuente	2017	A nuttalliellid^[119] tick.	Deinocroton draculi
		Deinocroton lacrimus^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick
	†Legionaris	†Legionaris robustus^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.	Legionaris robustus
	Nuttalliella	Nuttalliella gratae^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.
		Nuttalliella odyssea^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.
		Nuttalliella placaventrala^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.
		Nuttalliella tropicasylvae^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.
		Nuttalliella tuberculata^[119]	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	2024	A nuttalliellid tick.

Taxon	Authority	Year described	Notes
Biungulus xiai^[122]	Bartel et al.	2020	An epedanid armoured harvestman.
Burmalomanius circularis^[123]	Bartel et al.	2022	A podoctid harvestman.
Ellenbergellus tuberculatus^[122]	Bartel et al.	2020	A tithaeid harvestman.
Foveacorpus cretaceus^[124]	Bartel, Dunlop & Giribet	2023	A member of Cyphophthalmi.
Foveacorpus parvus^[124]	Bartel, Dunlop & Giribet	2023	A member of Cyphophthalmi.
Gigantocheles nilsi^[122]	Bartel et al.	2020	An epedanid armoured harvestman.
Halitherses grimaldii^[125]	Giribet & Dunlop	2005	An ortholasmatine nemastomatid harvestman
Leptopsalis breyeri^[124]	Bartel, Dunlop & Giribet	2023	A stylocellid harvestman.
Mesodibunus tourinhoae^[123]	Bartel et al.	2022	An epedanid harvestman.
Mesokanus oehmkuehnlei^[122]	Bartel et al.	2020	A mesokanid laniatorean harvestman.
Mesopsalis oblongus^[124]	Bartel, Dunlop & Giribet	2023	A member of Cyphophthalmi.
Monooculricinuleus incisus^[126]	Wunderlich	2017	A monooculricinuleid harvestman.^[89]
Monooculricinuleus semiglobosus^[126]	Wunderlich	2017	A monooculricinuleid harvestman.^[89]
Palaeobeloniscus thilolebi^[122]	Bartel et al.	2020	A beloniscid harvestman.
Palaeosiro burmanicum^[127]	Poinar	2008	A stylocellid harvestmant
Petrobunoides sharmai^[128]	Selden et al.	2016	An epedanid armoured harvestman.
Petroburma tarsomeria^[123]	Bartel et al.	2022	A petrobunid harvestman.
Protopyramidops nalae^[122]	Bartel et al.	2020	Possibly a pyramidopid harvestman.
Sirocellus iunctus^[124]	Bartel, Dunlop & Giribet	2023	A member of Cyphophthalmi.
Tyrannobunus aculeus^[129]	Bartel & Dunlop	2023	A eupnoid harvestman.

Taxon	Authority	Year described	Notes	image
Amblyolpium burmiticum^[131]	(Cockerell)	1920	A garypinid pseudoscorpion. Originally described as "Garypus" burmiticus.
Burmeochthonius kachinae^[132]	Johnson et al.	2022	A chthoniid pseudoscorpion.
Burmeochthonius muelleri^[132]	Johnson et al.	2022	A chthoniid pseudoscorpion.
Electrobisium acutum^[133]	Cockerell	1917	A neobisiid pseudoscorpion
Hya fynni^[134]	Röschmann et al.	2024	A hyid pseudoscorpion.
Prionochthonius burmiticus^[135]	Wriedt et al.	2021	A chthoniid pseudoscorpion
Proalbiorix compactus^[136]	Geißler et al.	2021	A pseudoscorpion belonging to the family Ideoroncidae.
Proalbiorix gracilis^[136]	Geißler et al.	2021	A pseudoscorpion belonging to the family Ideoroncidae.
Procheiridium judsoni^[137]	Porta et al.	2020	A cheiridiid pseudoscorpion.	Procheiridium judsoni
Protofeaella peetersae^[138]	Henderickx in Henderickx & Boone	2016	A pseudoscorpion of uncertain placement.^[139]
Weygoldtiella plausus^[140]	Harvey et al.	2018	A chthoniid pseudoscorpion.

Taxon	Authority	Year described	Notes
Hirsutisoma acutiformis^[126]	Wunderlich	2017	A Hirsutisomid hooded tickspider.
Hirsutisoma bruckschi^[126]	Wunderlich	2017	A Hirsutisomid hooded tickspider.
Hirsutisoma dentata^[126]	Wunderlich	2017	A Hirsutisomid hooded tickspider.
Hirsutisoma grimaldii^[141]	Botero-Trujillo et al.	2022	A hirsutisomid hooded tickspider.
?Poliochera cretacea^[142]	Wunderlich	2012	A poliocherid hooded tickspider of uncertain generic placement.
Primoricinuleus pugio^[78]	Wunderlich	2015	A primoricinuleid hooded tickspider.
Sigillaricinuleus tripares^[143]	Wunderlich	2022

Family	Genus	Species	Authority	Year described	Notes	image
Achipteriidae	†Cerachipteria	†Cerachipteria ahsokatanoae^[144]	Arillo, Subías & Huang	2023	An achipteriid mite.
Pediculochelidae	†Paralycus	†Paralycus primus^[145]	Kolesnikov et al.	2025	A pediculochelid mite.

Taxon	Authority	Year described	Notes
Annazomus parvulus^[146]	De Francesco Magnussen in De Francesco Magnussen et al.	2022	A member of the family Hubbardiidae.
Cretaceozomus angustocaudatus^[146]	De Francesco Magnussen in De Francesco Magnussen et al.	2022
Cretaceozomus robustus^[146]	De Francesco Magnussen in De Francesco Magnussen et al.	2022
Groehnizomus oculiferans^[146]	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	2022
Groehnizomus rodrigoi^[146]	Müller in De Francesco Magnussen et al.	2022
Mesozomus groehni^[147]	Müller et al.	2020	A short-tailed whip scorpion
Muellerizomus amandae^[146]	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	2022
Muellerizomus palicaudatus^[146]	De Francesco Magnussen in De Francesco Magnussen et al.	2022

Taxon	Authority	Year described	Notes	image
Archaeoananteroides carusoi^[148]	Lourenço in Lourenço & Velten	2025	A buthoid scorpion.
Archaeoananteroides maderai^[149]	Lourenço & Velten	2016	A buthoid scorpion.
Archaeoscorpiops cretacicus^[150]	Lourenço	2015	A palaeoeuscorpiid scorpion.
Archaeoscorpiops grossei^[151]	Lourenço in Lourenço & Velten	2023	A palaeoeuscorpiid scorpion.
Betaburmesebuthus bellus^[152]	Lourenço	2016	A palaeoburmesebuthid scorpion.	Betaburmesebuthus bellus
Betaburmesebuthus bidentatus^[153]	Lourenço	2015	A palaeoburmesebuthid scorpion.^[152]
Betaburmesebuthus fleissneri^[154]	Lourenço & Velten	2016	A palaeoburmesebuthid scorpion.^[152]
Betaburmesebuthus fuscus^[155]	Xuan, Cai & Huang	2023	A palaeoburmesebuthid scorpion.
Betaburmesebuthus joergi^[156]	Lourenço & Rossi	2017	A palaeoburmesebuthid scorpion.
Betaburmesebuthus kobberti^[157]	Lourenço	2015	A palaeoburmesebuthid scorpion.
Betaburmesebuthus larafleissnerae^[158]	Lourenço & Velten	2016	A palaeoburmesebuthid scorpion.
Betaburmesebuthus muelleri^[153]	Lourenço	2015	A palaeoburmesebuthid scorpion.
Betaburmesebuthus spinipedis^[159]	Xuan, Cai & Huang	2022	A palaeoburmesebuthid scorpion.
Betaburmesebuthus thomasvelteni^[160]	Lourenço in Lourenço & Velten	2024	A palaeoburmesebuthid scorpion.
Betaburmesebuthus villosus^[155]	Xuan, Cai & Huang	2023	A palaeoburmesebuthid scorpion.
Burmesescorpiops groehni^[161]	Lourenço	2016	An Archaeoscorpiopinae palaeoeuscorpiid scorpion.
Burmesescorpiops velteni^[162]	Lourenço	2024	An Archaeoscorpiopinae palaeoeuscorpiid scorpion.
Chaerilobuthus birmanicus^[163]	Lourenço	2015	A chaerilobuthid or pseudochactid^[164] scorpion.
Chaerilobuthus brandti^[165]	Lourenço	2022	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus bruckschi^[163]	Lourenço	2015	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus complexus^[166]	Lourenço & Beigel	2011	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus enigmaticus^[167]	Lourenço	2015	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus gigantosternum^[161]	Lourenço	2016	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus hansgeorgmuelleri^[168]	Lourenço	2019	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus knodelorum^[169]	Lourenço	2018	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus longiaculeus^[170]	Lourenço	2013	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus meggeri^[171]	Lourenço	2021	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus petersi^[172]	Lourenço in Lourenço & Velten	2024	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus serratus^[161]	Lourenço	2016	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus schwarzi^[173]	Lourenço	2015	A chaerilobuthid or pseudochactid scorpion.
Chaerilobuthus staxi^[174]	Lourenço in Lourenço & Velten	2024	A chaerilobuthid or pseudochactid scorpion.
Chaeriloiurus brigittemuellerae^[175]	Lourenço	2020	A chaerilobuthid or pseudochactid scorpion. Possibly a species of Chaerilobuthus.^[164]
Chaeriloscorpiops bautschi^[175]	Lourenço	2020	A palaeoeuscorpiid scorpion.
Cretaceousbuthus fraaijeorum^[176]	Lourenço	2022	A buthid scorpion.
Cretaceousbuthus petersi^[177]	Lourenço in Lourenço & Velten	2023	A buthid scorpion.
Cretaceoushormiops elegans^[178]	Xuan et al.	2023	A protoischnurid scorpion.
Cretaceoushormiops knodeli^[179]	Lourenço	2018	A protoischnurid scorpion.
Cretaceoushormiops petersi^[180]	Lourenço	2025	A protoischnurid scorpion.
Cretaceoushormiops staxi^[181]	Lourenço in Lourenço & Velten	2022	A protoischnurid scorpion.
Cretaceousopisthacanthus smeelei^[182]	Lourenço in Lourenço & Velten	2021	A protoischnurid scorpion.
Electrochaerilus buckleyi^[183]	Santiago-Blay, Fet, Solegrad & Anderson	2004	A chaerilid scorpion
Palaeoburmesebuthus andrewrossi^[184]	Santiago-Blay et al.	2022	A palaeoburmesebuthid scorpion.
Palaeoburmesebuthus grimaldii^[185]	Lourenço	2002	A palaeoburmesebuthid scorpion.
Palaeoburmesebuthus knodeli^[186]	Lourenço	2018	A palaeoburmesebuthid scorpion.
Palaeoburmesebuthus longipalpis^[156]	Lourenço & Rossi	2017	A palaeoburmesebuthid scorpion.
Palaeoburmesebuthus ohlhoffi^[163]	Lourenço	2015	A palaeoburmesebuthid scorpion.
Palaeoburmesebuthus smithi^[184]	Santiago-Blay, Soleglad, Craig & Fet in Santiago-Blay et al.	2022	A palaeoburmesebuthid scorpion.
Palaeotrilineatus ellenbergeri^[187]	Lourenço	2012	A palaeotrilineatid scorpion.
Paranotaburmesebuthus schmidti^[188]	Lourenço in Lourenço & Velten	2024	A palaeoburmesebuthid scorpion.
Serratochaerilobuthus schmidti^[189]	Lourenço in Lourenço & Velten	2024	A chaerilobuthid or pseudochactid scorpion. Possibly a species of Chaerilobuthus.^[164]
Spinoburmesebuthus knodelorum^[190]	Lourenço	2021	A palaeoburmesebuthid buthoidean scorpion.
Spinoburmesebuthus pohli^[191]	Lourenço & Velten	2017	A palaeoburmesebuthid buthoidean scorpion.
Sucinlourencous adrianae^[192]	Rossi	2015	A sucinlourencoid scorpion.

Taxon	Authority	Year described	Notes
Burmathelyphonia prima^[78]	Wunderlich	2015	A whip scorpion of uncertain placement.
Crethypoctonus kachinus^[194]	Zhou et al.	2023	A whip scorpion.
Mesothelyphonus parvus^[195]	Cai & Huang	2017	A thelyphonine whip scorpion.

Family	Genus	Species	Authority	Year described	Notes	image
Bdellidae	†Odontoscirus	†Odontoscirus cretacico^[196]	Porta et al.	2020	A bdellid mite.
Caeculidae	†Procaeculus	†Procaeculus coineaui^[197]	Porta et al.	2019	A caeculid mite.
Cheyletidae	†Cheyletus	†Cheyletus burmiticus^[198]	Cockerell	1917	A cheyletid mite
Erythraeidae	†Burerythrites	†Burerythrites pankowskii^[199]	Konikiewicz & Mąkol	2018	An erythraeid mite.
Erythraeidae	†Burphanolophus	†Burphanolophus joergwunderlichi^[199]	Konikiewicz & Mąkol	2018	An erythraeid mite.
Resinacaridae	†Protoresinacarus	†Protoresinacarus brevipedis^[200]	Khaustov & Poinar	2011	A resinacarid pyemotoid mite.
Smarididae	†Burfessonia	†Burfessonia maryae^[199]	Konikiewicz & Mąkol	2018	A Smaridid mite.
Smarididae	†Immensmaris	†Immensmaris chewbaccei^[201]	Dunlop, Frahnert & Mąkol	2018	A smaridid mite.	Immensmaris chewbaccei
Trochometridiidae	†Rhombometridium	†Rhombometridium pankowskiorum^[202]	Khaustov, Vorontsov & Lindquist	2024	A trochometridiid mite.
Trombellidae	†Nothrotrombidium	†Nothrotrombidium myanmarum^[199]	Konikiewicz & Mąkol	2018	A trombellid mite.

Taxon	Authority	Year described	Notes	image
Chimerarachne alexbeigel^[101]	Wunderlich	2023
Chimerarachne longiflagellum^[101]	(Wunderlich)	2022	Originally described as Parachimerarachne longiflagellum.^[108]
Chimerarachne patrickmueller^[101]	Wunderlich	2023
Chimerarachne spiniflagellum^[101]	Wunderlich	2023
Chimerarachne yingi^[203]	Wang et al.	2018	A stem-group spider	Chimerarachne yingi (artists reconstruction)

Taxon	Authority	Year described	Notes	image
Kachinophilus pereirai^[204]	Bonato, Edgecombe & Minelli	2013	A geophilid centipede
Lithopendra anjafliessae^[205]	Haug, Haug & Haug	2023	A centipede belonging to the group Pleurostigmophora

Taxon	Authority	Year described	Notes	image
Monstrodesmus grimaldii^[209]	Su, Cai & Huang	2022	A trichopolydesmid millipede
Propolydesmus cretaceus^[210]	Su, Cai & Huang	2023	A polydesmid millipede

Taxon	Authority	Year described	Notes
Pauropsxenus extraneus^[211]	Su, Cai & Huang	2020	A polyxenid millipede.
Pauropsxenus ordinatus^[211]	Su, Cai & Huang	2020	A polyxenid millipede.
Phryssonotus burmiticus^[198]	Cockerell	1917	A synxenid millipede

Chlorophyte green algaes

Bryophyte true mosses

Lycopods and spike mosses

Marchantiophyte liverworts

Polypodiopsid ferns

Angiosperm flowering plants

incertae sedis

Rozellomycota

"Zygomycetes"

Priscadvenales

Crinoidea

Arachnida

Uropygi (Thelyphonida)

Chilopoda

Diplopoda

"Entognatha"

Insecta

Malacostraca

Ostracoda

Symphyla

Cephalopoda

Gastropoda

"Architaenioglossa"

Chromadorea

Enoplea

Secernentea

Gordioidea

Amphibia

Reptilia

Archosauria

Insecta

Dictyoptera

Mollusca

Bivalvia

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Taxon	Authority	Year described	Notes	image
Siphoniulus muelleri^[212]	Liu, Rühr & Wesener	2017	A siphoniulid millipede.
Siphoniulus preciosus^[212]	Liu, Rühr & Wesener	2017	A siphoniulid millipede.

Taxon	Authority	Year described	Notes
Siphonophora hui^[213]	Jiang et al.	2019	A siphonophorid sucking millipede.
Siphonorhinus globosus^[214]	Su, Cai & Huang	2024	A siphonorhinid millipede.
Siphonorhinus peculiaris^[214]	Su, Cai & Huang	2024	A siphonorhinid millipede.

Taxon	Authority	Year described	Notes
Electrocambala cretacea^[215]	Moritz & Wesener	2021	A member of Cambalidea.
Electrocambala ornata^[215]	Moritz & Wesener	2021	A member of Cambalidea.
Kachincambala distorta^[215]	Moritz & Wesener	2021	A member of Cambalidea.
Kachincambala muelleri^[215]	Moritz & Wesener	2021	A member of Cambalidea.

Taxon	Authority	Year described	Notes
Burmisotoma lamellifera^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Cretacentomobrya burma^[216]	Christiansen & Nascimbene	2006	A praentomobryid springtail
Praentomobrya avita^[216]	Christiansen & Nascimbene	2006	A praentomobryid springtail
Proisotoma pettersonae^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Propachyotoma conica^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Protodesoria granda^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Protoisotoma burma^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Villusisotoma brevis^[216]	Christiansen & Nascimbene	2006	An isotomid springtail
Villusisotoma longa^[216]	Christiansen & Nascimbene	2006	An isotomid springtail

Taxon	Authority	Year described	Notes
Grinnellia ventis^[216]	Christiansen & Nascimbene	2006	A sminthurid springtail
Mucrovirga incompleta^[216]	Christiansen & Nascimbene	2006	A sminthurid springtail
Sminthurconus grimaldi^[216]	Christiansen & Nascimbene	2006	A sminthurid springtail
Sminthuricinus deceptus^[216]	Christiansen & Nascimbene	2006	A sminthurid springtail

Taxon	Authority	Year described	Notes	image
Cretapsara athanata^[217]	Luque in Luque et al.	2021	A crab belonging to the group Eubrachyura and to the family Cretapsaridae.
Xiaopenaeus electrinus^[218]	Xing et al.	2021	A member of the family Penaeidae.

Taxon	Authority	Year described	Notes
Cryptolacruma nidis^[219]	Schädel et al.	2021	A member of Epicaridea.
Electrolana madelineae^[220]	Schädel, Hyžný & Haug	2021	A member of Cymothoida, possibly a species of Cirolana.^[221]
Metacirolana jimlowryi^[221]	Bruce & Rodcharoen	2023	A member of the family Cirolanidae.
Myanmariscus deboiseae^[222]	Broly, Maillet & Ross	2015	A possible styloniscid woodlouse.^[223]
Palaeoarmadillo microsoma^[223]	Poinar	2018	An armadillid woodlouse.

Taxon	Authority	Year described	Notes
Electrocypria burmitei^[225]	Wang, Matzke-Karasz & Horne	2022	A candonid ostracod.
Myanmarcypris hui^[226]	Wang et al.	2020	A candonid ostracod.
Sanyuania sp.^[226]	Wang et al.	2020	A loxoconchid cytheroid.
Thalassocypria sp.^[226]	Wang et al.	2020	A candonid ostracod.

Taxon	Authority	Year described	Notes	image
Archaeocyclotus brevivillosus^[229]	Bichain et al.	2022	A cyclophorid land snail
Archaeocyclotus plicatula^[230]	Asato & Hirano in Hirano et al.	2019	A cyclophorid land snail	Archaeocyclotus plicatula
Burminella artiukhini^[231]	Balashov, Perkovsky & Vasilenko	2021	A land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea.
Coptocheilus electrothauma^[230]^[232]	(Asato & Hirano in Hirano et al.)	2019	A species of Coptocheilus. Originally described as Schistoloma electrothauma.	Coptocheilus electrothauma
Coptocheilus kachinensis^[233]	Yu, Zhuo & Páll-Gergely	2022	A species of Coptocheilus.
Cretadiostoma caperatum^[233]	Yu, Zhuo & Páll-Gergely	2022	A member of the family Pupinidae.
Cretadiostoma umbilicarinatum^[233]	Yu, Zhuo & Páll-Gergely	2022	A member of the family Pupinidae.
Cretatortulosa cretakachinensis^[234]	Yu, Salvador & Jarzembowski	2021	A pupinid land snail
Cretatortulosa gignens^[235]	Jochum, Yu & Neubauer	2021	A pupinid land snail
Eotrichophorus kachin^[236]	Bullis et al.	2020	A cyclophorid land snail
Euthema annae^[237]	Balashov	2020	A diplommatinid land snail
Euthema hesoana^[230]	Asato & Hirano in Hirano et al.	2019	A diplommatinid land snail	Euthema hesoana
Euthema truncatellina^[238]	Balashov, Perkovsky & Vasilenko	2020	A diplommatinid land snail
Hirsuticyclus canaliculatus^[239]	Yu	2021	A cyclophorid land snail
Hirsuticyclus electrum^[240]	Neubauer, Xing & Jochum	2019	A cyclophorid land snail
Lagocheilus cretaspira^[230]	Asato & Hirano in Hirano et al.	2019	A cyclophorid land snail	Lagocheilus cretaspira
Lagocheilus electrospira^[230]	Asato & Hirano in Hirano et al.	2019	A cyclophorid land snail. Might be a species of Eotrichophorus.^[237]	Lagocheilus electrospira
Macropupina electricus^[236]	Bullis et al.	2020	A pupinid land snail.
Paleodiplommatina spelomphalos^[236]	Bullis et al.	2020	A diplommatinid land snail. Might be a species of Euthema.^[237]
Perissocyclos kyrtostoma^[236]	Bullis et al.	2020	A cyclophorid land snail.
Pseudopomatias lyui^[241]^[242]	(Yu, Wang & Jarzembowski)	2019	A cyclophorid land snail, first as "Epitonium (Papyriscala) lyui"
Pseudopomatias zhuoi^[241]^[242]	(Yu, Wang & Jarzembowski)	2019	A cyclophorid land snail, first as "Epitonium (Epitonium) zhuoi"
Pulchraspira teneristoma^[234]	Yu, Salvador & Jarzembowski	2021	A diplommatinid land snail
Rhaphaulus zhuoi^[234]	Yu, Salvador & Jarzembowski	2021	A pupinid land snail
Xenostoma lophopleura^[236]	Bullis et al.	2020	Originally described as a pupinid land snail. Might be a species of Euthema instead.^[243]

Taxon	Authority	Year described	Notes
Galba prima^[244]	Yu, Neubauer & Jochum	2021	A species of Galba.
Mathilda^[228]	N/A	"2019"	2 dead shells likely entombed as shoreline debris
Truncatellina dilatatus^[245]	Yu	2020	Originally described as a species of Truncatellina. Might be a species of Euthema instead.^[243]