Dengying Formation
Geologic formation in China
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The Dengying Formation is an upper Ediacaran (551-541 Ma[2]) fossiliferous geologic formation found in South China. It was deposited on a shallow marine carbonate platform.[3] It also contains a rare example of Burgess Shale-type Preservation within the Ediacaran, in the form of the Tongshan Lagerstätte.[4]
| Dengying Formation | |
|---|---|
| Stratigraphic range: Ediacaran ~ | |
Transition between uppermost Doushantuo Formation and lowermost Dengying Formation | |
| Type | Formation |
| Sub-units | See: Members |
| Underlies | Yanjiahe Formation[1] |
| Overlies | Doushantuo Formation |
| Lithology | |
| Primary | Dolomite |
| Other | Chert, Limestone |
| Location | |
| Region | Yangtze Gorges |
| Country | China |
Members
Listed by ascending age:
- The Hamajing Member is characterized by light grey, medium- to thick-bedded dolomite, intercalated with thin layers of chert.[3]
- The Shibantan Member is interpreted to represent a subtidal environment and is composed of dark grey, thin-bedded bituminous limestone. It is the most fossiliferous of the Dengying Formation members and has yielded several trace fossils, such as Lamonte,[5] as well as possible lobopodian trackways.[6]
- The Baimatuo Member consists of light grey, thick-bedded dolomite.[3]
Traditionally, the Tianzhushan Member was considered to be the uppermost unit of the Dengying Formation. However, its small shelly fossils and Micrhystridium-like acritarchs are a shared characteristic with the Cambrian Yanjiahe Formation.[3] The Kuanchuanpu Formation also is occasionally placed as a member of the Dengying Formation,[7] however it is likely separate for the same reasons.
Paleobiota
The Dengying formation is unique from other Ediacaran formations in that its top is almost right up against the Ediacaran - Cambrian boundary, with more recent organisms being described that have striking similarities with Cambrian fauna, like Alienum. There are also a lot of ichnogenera in the form of burrows, something that is common in younger Ediacaran rocks the closer to the Cambrian boundary they are.
A recent paper has also found a rare example of Burgess Shale-type Preservation outside of the known range of all members, in what is known as the Tongshan Lagerstätte. This preserves a wide range of fossil forms, including the first rangeomorph fronds with Burgess Shale-type preservation.[4]
Color key
|
Notes Uncertain or tentative taxa are in small text; |
Proarticulata
| Genus | Species | Notes | Images |
|---|---|---|---|
| Dickinsonia[8] |
|
Oval to elongated motile organism. Only one incomplete specimen is known. | |
Petalonamae
| Genus | Species | Notes | Images |
|---|---|---|---|
| Charniodiscus[9] |
|
Frondose organism. | |
| Paracharnia[10] |
|
Frondose organism. | |
| Pteridinium[9] |
|
Frondose organism. | |
| Rangea[9] |
|
Frondose organism. | |
Trilobozoa
| Genus | Species | Notes | Images |
|---|---|---|---|
| Lobodiscus[11] |
|
Tri-radial organism. | |
Porifera
| Genus | Species | Notes | Images |
|---|---|---|---|
| Helicolocellus[12] |
|
Conical sponge, possible relations to hexactinelliid sponges. | |
incertae sedis
| Genus | Species | Notes | Images |
|---|---|---|---|
| Alienum[13] |
|
Vetulicolian-like organism. | |
| Cloudina[14] |
|
Tubular organism. | |
| Curviacus[15] |
|
Palaeopascichnid organism. | |
| Conotubus[16] |
|
Tubular organism. | |
| Cycliomedusa[17] |
|
Discoid organism. | |
| Eoandromeda[4] |
|
Discoid organism. | |
| Gaojiashania[18] |
|
Tubular organism. | |
| Hiemalora[9] |
|
Discoid organism, possibly holdfasts of petalonamids. | |
| Palaeorhopalon[14] |
|
Tubular organism. | |
| Palaeopascichnus[4] |
|
Palaeopascichnid organism. | |
| Protoconites[4] |
|
Curved conical organism, possible relations to Cnidaria. | |
| Shufangtubulus[14] |
|
Tubular organism. | |
| Sinotubulites[19][14] |
|
Tubular organism. | |
| Wutubus[9] |
|
Tubular organism. | |
| Yangtziramulus[20] |
|
Enigmatic organism, affinities with other forms is unknown, although possibly related to the Petalonamae, like Charnia. | |
| Yilingia[21] |
|
Bilterian animal of unknown affinities, with possible relations to Panarthopoda or Annelida. | |
Flora
| Genus | Species | Notes | Images |
|---|---|---|---|
| Longfengshania[17] |
|
Thallus-like macroalgae. | |
| Protocodium[22] |
|
Codiacean alga. | |
| Baculiphyca[4] |
|
Thallus-like macroalgae. | |
| Zhongbaodaophyton[4] |
|
Branching macroalgae. | |
| Doushantuophyton[4] |
|
Branching macroalgae. | |
| Chuaria[4] |
|
Enigmatic fossil, possibly synonymous with Tawuia. | |
| Beltanelliformis[4] |
|
Cyanobacterial colony. | |
| Longifuniculum[4] |
|
Whip-like macroalgae. | |
Microorganisms
| Genus | Species | Locality | Notes | Images |
|---|---|---|---|---|
| Protolagena[23] |
|
Lower Shibantan Member (Gaojiashan and Lijiagou Sections) | Vase-like microorganisms. | |
| Sicylagena[23] |
|
Lower Shibantan Member (Gaojiashan and Lijiagou Sections) | Vase-like microorganisms. | |
Ichnogenera
| Genus | Species | Locality | Notes | Images |
|---|---|---|---|---|
| Lamonte[24] |
|
Middle Shibantan Member (South Quarry) | Burrows. | |
| Helminthoidichnites |
|
Burrows. | ||
| Shaanxilithes |
|
Junior synonym of Nenoxites. | ||
| Nenoxites |
|
Movement traces. | ||
| Torrowangea |
|
Burrows. | ||
Undescribed
| Genus | Species | Locality | Notes | Images |
|---|---|---|---|---|
| ESEN 0013[4] |
|
Tongan Lagerstätte (Wanjia section) | Tubular organism, possible affinities to Metazoa. | |
| Undescribed rangeomorph "ESEN 0016"[4] |
|
Tongan Lagerstätte (Wanjia section) | Frondose organism, 3 distinct lobe-like regions, consisting of a thick stalk and two petaloids, which are made up of tightly packed branches. | |
| Fan-shaped rangeomorphs[4] |
|
Tongan Lagerstätte (Wanjia section) | Two frondose organisms with a similar appearance to Bradgatia, with one of the specimens, ESEN 0018, preserving with a possible holdfast. | |
| Undescribed rangeomorph "ESEN 0019"[4] |
|
Tongan Lagerstätte (Wanjia section) | Frondose organism, with a similar appearance to Charniodiscus, consisting of a thick stalk and two petaloids, which are made up of tightly packed branches, also preserves a notable holdfast. | |
| Undescribed rangeomorph "ESEN 0020"[4] |
|
Tongan Lagerstätte (Wanjia section) | Elongated frondose organism, with a similar appearance to Paracharnia, consisting of a thin stalk and two petaloids, which are made up of tightly packed branches, alongside a possible holdfast at its base. | |
| Zhenba columns[25] |
|
Tongan Lagerstätte (Wanjia section) | Possible affinities with coralline algae. | |
| Baseball bat-shaped form[4] |
|
Tongan Lagerstätte (Wanjia section) | Baseball bat-shaped form, possibly macroalgae. | |
| Irregular circular form[4] |
|
Tongan Lagerstätte (Wanjia section) | Oblong-shaped form, possibly macroalgae. | |
| Net-like form[4] |
|
Tongan Lagerstätte (Wanjia section) | Netted form, possibly macroalgae. | |
| ribbon-like form[4] |
|
Tongan Lagerstätte (Wanjia section) | Ribbon like form, possibly macroalgae. | |
| Bilateria indet. |
|
Tongan Lagerstätte (Wanjia section) | Known to have several pairs of appendages, but its exact affinity is unknown[26] | |












