↑ Peter R. Teske, Colin L. McLay, Jonathan Sandoval-Castillo, Isabelle Papadopoulos, Brent K. Newman, Charles L. Griffiths, Christopher D. McQuaid, Nigel P. Barker, Gaetan Borgonie, Luciano B. Beheregaray(2009).“Tri-locus sequence data reject a "Gondwanan origin hypothesis" for the African/South Pacific crab genus Hymenosoma”.Mol. Phyl. Evol.53(1): 23–33.doi:10.1016/j.ympev.2009.05.031.
↑ Takeda,Masatune (武田正倫);Miyake,Sadayoshi (三宅貞祥)(1971).“A new freshwater crab of the family Hymenosomatidae from the Palau Islands.(パラオ諸島産ヤワラガニ科の1新種)”.甲殻類の研究4,5(157-163).NAID110002698416.