Khasurty locality

Fossil locality in Russia From Wikipedia, the free encyclopedia

The Khasurty locality is an Early Cretaceous (Neocomian) lagerstätte in Russia, within the Sangin Formation. This lagerstätte is located outside the village of Tsakir in the region of Zakamensk. Over 6,000 insect specimens have been found from the site, belonging to around 130 families. Its insect biota resembles both Jurassic and Cretaceous ones, most heavily the Jehol biota of China.^[1]

Unit ofGusinoe Ozero Group

RegionZakamensky District

CountryRussia

Named forKhasurty River

Quick facts Unit of, Location ...

Khasurty locality
Stratigraphic range: Neocomian
Unit of	Gusinoe Ozero Group
Location
Region	Zakamensky District
Country	Russia
Type section
Named for	Khasurty River

Paleobiota

Non-Insecta

More information Non-insect organisms, Genus ...

Non-insect organisms
Genus	Species	Higher taxon	Notes	Images
Palaeoeorak	P. scherbakovi	Prochydorusidae (Anomopoda)^[2]	Only named crustacean from Khasurty^[1]	The extant anomopod Daphnia magna
Bryokhutuliinia	B. obtusifolia	Hookeriales?	Indeterminate family^[3]
Khasurtya	K. ginkgoides	Marchantiidae	Specific name comes from the similarity in leaf shape to the ginkgo^[1]
Khasurtythallus	K. monosolenioides	Marchantiidae	Likely an extinct clade of liverworts^[4]
Palaeodichelyma	P. sinitzae	Hypnanae (pleurocarpous mosses)	Also known from the Jurassic of Russia^[3]
Tricostium	T. longifolium	Bryophyta incertae sedis	Similar to a species found ~2000 km to the east^[3]

Insects

Palaeopterans

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Palaeopterans
Genus	Species	Higher taxon	Notes	Images
Baikalogenites	B. firmus	Hexagenitidae (Ephemeroptera)	Bears a longer seventh gill pair^[5]
Epiproctophora spp.	Unapplicable	Odonata	Known from several unnamed species, most seemingly within Isophlebiidae or Campterophlebiidae^[1]	Isophlebia aspasia specimen from the Jurassic of Germany
Furvoneta	F. khasurtensis	Mesonetidae (Ephemeroptera)	Largest member of the genus^[5]
Mesobaetis	M. crispa	Siphlonuridae	Similar to the species M. amplectus, but has a larger rib on its gill plates^[5]
Mesogenesia	M. sp	Mickoleitiidae	Only known from a poorly preserved specimen^[5]	Larva of the closely related Mickoleitia
Proameletus	P. branchiatus	Siphlonuridae	Bears larger gills than the type species^[5]

Polyneopterans

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Polyneopterans
Genus	Species	Higher taxon	Notes	Images
Aboilus?	A?. khasurty	Prophalangopsidae	Differs from Aboilus in wing venation, but still fairly close to it^[1]	Pycnophlebia fossil from Jurassic Germany
Archisusumania	A. contacta	Susumanioidea	Differs from other genera in wing venation^[1]	Three specimens of the closely related Cretophasmomima from China
Dimoula	D. khasurtensis	Nemouridae	One of the few extinct nemourids known^[6]	Nemoura cinerea, an extant nemourid
Jurataenionema	J. rohdendorfi	Taeniopterygidae	Known from a complete female specimen^[6]	Taeniopteryx nebulosa, an extant taeniopterygid
Mongoloxya	M. fossor	Tridactylidae	Differs from the other species in the genus by having digging-adapted forelimbs^[1]	An unidentified extant tridactylid from India
Rasnitsyrina	R. culonga, R. desiliens	Perlariopseidae (Plecoptera)	Most abundant stonefly from the site^[6]
Shurabia	S. creta	Geinitziidae	Latest known winged notopteran besides those known from Burmese amber^[1]
Siberoperla	S. bashkuevi	Peltoperlidae	Oldest peltoperlid, alongside the first fossil of a nymph from the family^[7]	An indeterminate peltoperlid nymph

Paraneopterans

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Paraneopterans
Genus	Species	Higher taxon	Notes	Images
Buryatocera	B. beta	Kobdocoridae (Pentatomoidea)	Has relatively short antennae^[8]
Creocanadaphis	C. hirtus	Canadaphididae	Abdomen bears numerous small spines on sclerites^[1]
Extralapis	E. carens, E. breviscutum	Kobdocoridae	Intermediate between the two other kobdocorid genera from Khasurty in pronotum (top-front section of thorax) proportions^[8]
Khasuaphis	K. serotinus	Juraphididae	Bears longer antennae than other members of its family^[1]
Khasoris	K. yuripopovi	Kobdocoridae (Pentatomoidea)	Bears a very long ovipositor^[8]
Liadopsylla	L. (Basicella) lauteri, L. (Basicella) loginovae	Liadopsyllidae	Type genus of the family^[9]
Occidoscelis	O. glaber	Anthocorinae	One of the only fossil anthocorids known^[1]	Anthocoris nemorum, an extant anthocorine bug
Reticycla	R. drospoulosi	Hylicellidae (Cicadomorpha)	Has a smaller ovipositor and a different foreleg structure to most other hylicellids^[9]
Stigmapsylla	S. klimaszewskii	Liadopsyllidae	Bears a dark pterostigma^[9]	Holotype of Amecephala pusilla, a different liadopsyllid from Burmese amber
Tobleronothrips	T. deparis	Aeolothripidae	Named for Toblerone chocolate, which "noticeably helps when identifying complex imprints of thrips"^[1]	The modern Erythrothrips, another aeolothripid species

Holometabola

Hymenopterans

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Hymenopterans
Genus	Species	Higher taxon	Notes	Images
Amplicella	A. mininae, A. bashkuyevi, A. shcherbakovi, A. minor^[10]	Tanychorinae (Ichneumonidae)	Has a dark body with lighter legs/wing veins^[11]	Tanychora petiolata fossil from the Zaza Formation
Ghilarella	G. kopylovi^[12]	Sepulcidae	Originally placed within G. mercurialis, as it only differed from the type of that species in wing colouration^[13]
Hymenoptera spp.	Unapplicable	Holometabola	Includes numerous undescribed species from within various families like Angarosphecidae, Megalyridae and Proctotrupidae^[1]
Khasips	K. alisecta, K. sculpta, K. kovalevi	Archaeocynipidae (Cynipoidea)	K. kovalevi has clubbed antennae, K. sculpta has an unusual pattern on its thorax^[14]
Khasurtella	K. buryatica	Tanychorinae (Ichneumonidae)	Genus also contains a former Tanychora species^[11]
Micramphilius	M. mirabilipennis	Sepulcidae	First member of its genus which preserves the hind wings^[15]
Onokhoius	O. venustus	Sepulcidae	Bears a dark band on its pterostigma^[15]
Pamparaphilius	P. khasurtensis	Sepulcidae	Holotype wing has numerous unusual veins, likely as a result of mutation^[15]
Parasyntexis	P. khasurtensis	Anaxyelidae^[16]
Praeichneumon	P. dzhidensis, P. khamardabanicus, P. zakhaaminicus	Praeichneumonidae	Known from several mostly complete specimens^[17]
Sclerostigma	S. trimaculata	Dolichostigmatinae (Anaxyelidae)	Species name means "three-spotted", as its abdominal tergites have patterns of three spots^[1]
Trematothorax	T. zhangi, T. brachyurus, T. extravenosus	Sepulcidae	Another distinct species is known, however the material is too poor to name a new taxon^[13]

Neuropteridans

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Neuropteridans
Genus	Species	Higher taxon	Notes	Images
Aberrantochrysa	A. buryatica, A. pulchella	Chrysopidae^[18]
Cretohondelagia	C. viridis	Priscaenigmatidae (Raphidioptera)	Youngest priscaenigmatid fossil known^[19]
Dzhidosmylus	D. solus	Osmylidae	Uncertain subfamily^[1]
Khasurtoberotha	K. bellissima	Berothidae	Similar to the modern Protobiella^[1]	A specimen of the extant berothid Protobiella
Mesypochrysa	M. cannabina, M. naranica	Chrysopidae^[18]
Osmylochrysa	O. navasia	Osmylidae	Previously only known from England, resembles chrysopid lacewings^[1]
Zakamnosmylus	Z. elongatus	Osmylidae	Similar to the Jurassic Ensiosmylus^[1]

Coleopterans

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Coleopterans
Genus	Species	Higher taxon	Notes	Images
Coleoptera spp.	Unapplicable	Coleoptera	Contains many undescribed species in families such as Staphylinidae and Scarabaeidae^[1]
Coptoclava	C. longipoda	Coptoclavidae	May not actually belong to the species^[1]	Coptoclava larva from the Jinju Formation
Distenorrhinus	D. (Buryatnemonyx) ovatus, D. (Buryatnemonyx) gratshevi	Nemonychidae	Another undescribed species bears very large eyes^[1]
Gracilicupes	G. minimus	Cupedidae	Much smaller than other species of the genus^[20]
Jurodes	J. shef	Jurodidae	Similar to Jurodes from Daohugou^[21]	Fossil of Jurodes
Notocupes	N. khasurtyiensis	Archostemata incertae sedis	Has larger eyes and a shorter first abdominal segment than other species in the genus^[20]	Several Notocupes specimens from Jurassic China alongside mid-Cretaceous Burmese amber
Palaeonecrophilus	P. buryaticus	Agyrtidae	Earliest known agyrtid beetle^[22]	The extant agyrtid Necrophilus subterraneus
Ponohydrochus	P. buryaticus	Hydrochidae	First fossil hydrochid genus^[23]	An extant Hydrochus beetle
Unda	U. popovi	Trachypachidae	Also known from the Jurassic^[24]	The extant trachypachid Trachypachus gibbsii

Mecopterans

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Mecopterans
Genus	Species	Higher taxon	Notes	Images
Antiquanabittacus	A. dispar, A. robustus, A. saltator	Bittacidae	Similar to extant Orobittacus^[25]	The extant bittacid Bittacus selysi
Itaphlebia	I. spp	Nannochoristidae	Known from very well-preserved material, however the difficult diagnosis of Itaphlebia species means the Khasurty species have not been named^[1]	The extant nannochoristid Nannochorista philpotti
Lichnomesopsyche	L. sp	Mesopsychidae	Only known from one wing^[1]	Complete specimen of L. daohugouensis from Jurassic China
Meropeidae spp.	Unapplicable	Mecoptera	Known from two different species each with a complete specimen, unusual for meropeids (which rarely fly)^[1]	The extant meropeid Merope tuber
Mesochorista	M. sp	Permochoristidae (related to Nannochoristidae)	Formerly known as "Yanorthophlebia", which is a synonym of "Liassochorista" and in turn Mesochorista^[26]

Dipterans

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Dipterans
Genus	Species	Higher taxon	Notes	Images
Amorimyia	A. robusta	Anisopodidae	Differs from related genera in its hairy head and larger size^[27]
Collessomma	C. sibirica, C. gnoma	Perissommatidae	Unlike modern perissommatids, this genus has fused (holoptic) eyes^[28]	The extant perissommatid Perissomma mcalpinei
Eoptychoptera	E. fasbenderi	Ptychopteridae	Distinguished from other species in its genus by an additional cross-vein on the wing^[27]	The extant ptychopterid Ptychoptera contaminata
Gilkania	G. transbaikalica	Podonominae (Chironomidae)	Only known from pupal exuvia^[27]
Juraxymyia	J. krzeminskii	Axymyiidae	Also known from Daohugou^[29]	The extant axymyiid Axymyia furcata
Kaluginamyia	K. enigmatica	Kaluginamyiidae (Chironomoidea)	Placed in its own family^[30]
Kovalevimyia	K. sp	Simuliidae	One of the earliest hematophagous fly fossils, likely fed on birds^[1]
Mangas	M. brevisubcosta, M. kovalevi	Bolitophilidae	Also known from Mongolia^[31]	The extant bolitophilid Bolitophila cinerea
Praearchitipula	P. kaluginae, P. podenasi, P. ribeiroi^[27]	Pediciidae	Relatively large for pediciids^[1]	The extant pediciid Pedicia margarita
Protanyderus?	P. madrizi	Tanyderidae	May not belong to the genus^[27]
Sinaxymyia	S. szadziewskii	Axymyiidae	Also known from Daohugou^[29]
Zhiganka	Z. longialata	Ptychopteridae	Known from complete specimens^[32]	The extant ptychopterid Bittacomorpha clavipes

Amphiesmenopterans

More information Amphiesmenopterans, Genus ...

Amphiesmenopterans
Genus	Species	Higher taxon	Notes	Images
Axiomaldia	A. flinti	Trichoptera	Uncertain family^[1]
Buranima	B. gorhon	Psychomyiidae	Differs from other genera in having a maxillary palp without a longer second segment^[1]	The extant psychomyiid Tinodes waeneri
Burimodus	B. novus	Vitimotauliidae (Trichoptera)^[33]
Buryatocentrus	B. suspiciosus	Yantarocentridae (Limnephiloidea)	Has two-segmented gonopods^[1]
Cretocrania	C. stekolnikovi, C. glossancestralis, C. inconcessa	Cretocraniinae (Eriocraniidae)	Differs from modern eriocraniids in having round wing tips and very long galeae^[1]	The extant eriocraniid Eriocrania semipurpurella
Dauroglos	D. hohlomak	Glossosomatidae	Similar to the genus Dajella^[1]	The extant Glossosoma intermedium
Duamodus	D. anichkovi, D. necessarius	Vitimotauliidae (Integripalpia)	Similar to Multimodus^[1]
Dysoneura	D. subbota	Dysoneuridae	Specific name derives from the Russian word for "Saturday"^[1]
Khasurtia	K. kopylovi, K. alexeii, K. lukashevichae, K. novissima^[1]	Dysoneuridae	A fairly diverse caddisfly genus, almost entirely known from Khasurty^[34]
Juraphilopotamus	J. callidus	Philopotamidae	Has small dark spots on forewing^[33]	The extant philopotamid Philopotamus montanus
Meloclada	M. diuturna, M. frequentatoria	Cladochoristidae (Trichoptera)	Similar to the related Cladochoristella^[1]
Mesocolepus	M. deflectum	Ptilocolepidae (Hydroptiloidea)	Bears large setal warts, with their shape used to distinguish the genus^[1]
Multimodus	M. amplus	Vitimotauliidae (Trichoptera)	Largest species of the genus^[33]
Palaeosiberomiya	P. zugzwanga	Dysoneuridae	Bears longitudinal stripes of dark scales on its wings^[1]
Protobaikalopsyche	P. rossica	Protobaikalopsychidae (Hydropsychoidea)	Likely basal to a clade including Polycentropodidae^[1]
Protorodinia	P. khasurtensis	Leptoceridae	Bears a large mesoscutum and unusual abdominal tip^[1]	The extant leptocerid Leptocerus tineiformis
Protosiberopteryx	P. antitheton, P. equesaeneus, P. praevernalis	Eolepidopterigidae	Known from one male and two female specimens, similar to Undopterix^[1]
Purbimodus	P. parvulus, P. khramovi	Vitimotauliidae (Trichoptera)	Redescribed in a 2021 paper^[35]
Sententimiya	S. wichardi	Psychomyiidae	Bears a plate on the fifth sternite similar to those on psychomyiids^[1]
Shadareniskia	S. inambularia	Hydrobiosidae	Known from a female specimen, as shown by the genitalia and a suture on the sixth sternite^[1]	The (male) holotype of the extant Hydrobiosis falcis
Siberocretopetus	S. ekaterinae	Glossosomatidae	Specific name honours Ekaterina Sidorchuk, a palaeoacarologist (researcher who studies fossil mites) who died in 2019^[1]	The extant agapetine glossosomatid Agapetus fuscipes
Siberoprostoria	S. archaeoscriptor	Hydrobiosidae	Known from a female specimen, as shown by the genitalia and a suture on the sixth sternite^[1]
Styxowia	S. predponimania	Polycentropodidae	Has large, spiny gonopods^[1]	The extant polycentropodid Polycentropus flavomaculatus
Susurimiya	S. transbaikalica	Psychomyiidae	Bears a swollen abdomen tip like modern psychomyiids, but differs in the structure of its maxillary palps^[1]
Terminoptysma	T. contradictoria	Terminoptysmatidae (Trichoptera)	Placed in its own monotypic family^[1]
Terrindusia	T. khasurtica, T. buriatica	Trichoptera	Known only from larval casings^[34]

References

[1]
Kopylov, D. S.; Rasnitsyn, A. P.; Aristov, D. S.; Bashkuev, A. S.; Bazhenova, N. V.; Dmitriev, V. Yu.; Gorochov, A. V.; Ignatov, M. S.; Ivanov, V. D.; Khramov, A. V.; Legalov, A. A.; Lukashevich, E. D.; Mamontov, Yu. S.; Melnitsky, S. I.; Ogłaza, B.; Ponomarenko, A. G.; Prokin, A. A.; Ryzhkova, O. V.; Shmakov, A. S.; Sinitshenkova, N. D.; Solodovnikov, A. Yu.; Strelnikova, O. D.; Sukacheva, I. D.; Uliakhin, A. V.; Vasilenko, D. V.; Wegierek, P.; Yan, E. V.; Zmarzły, M. (December 2020). "The Khasurty Fossil Insect Lagerstätte". Paleontological Journal. 54 (11): 1221–1394. Bibcode:2020PalJ...54.1221K. doi:10.1134/S0031030120110027.
[2]
Kotov, Alexey A. (February 2009). "A revision of the extinct Mesozoic family Prochydoridae Smirnov, 1992 (Crustacea: Cladocera) with a discussion of its phylogenetic position". Zoological Journal of the Linnean Society. 155 (2): 253–265. doi:10.1111/j.1096-3642.2008.00412.x.
[3]
Ignatov, Michael S.; Schherbakov, Dmitry E. (December 2011). "Lower Cretaceous mosses from Khasurty (Baikal Area of South Siberia)". Arctoa. 20 (1): 19–42. doi:10.15298/arctoa.20.02.
[4]
Mamontov, Yuriy S.; Ignatov, Michael S. (July 2019). "How to rely on the unreliable: Examples from Mesozoic bryophytes of Transbaikalia". Journal of Systematics and Evolution. 57 (4): 339–360. doi:10.1111/jse.12483.
[5]
Sinitshenkova, N. D. (May 2017). "New mayflies (Insecta: Ephemerida = Ephemeroptera) from the Upper Mesozoic locality of Khasurty, Western Transbaikalia". Paleontological Journal. 51 (3): 280–287. Bibcode:2017PalJ...51..280S. doi:10.1134/S003103011703011X.
[6]
Sinitshenkova, N. D. (September 2011). "New stoneflies (Insecta: Perlida = Plecoptera) from the late Mesozoic locality Khasurty, Transbaikalia". Paleontological Journal. 45 (5): 552–558. doi:10.1134/S0031030111050133.
[7]
Sinitshenkova, Nina; Yan, Evgeny (15 October 2024). "The most ancient roachfly (Insecta: Plecoptera, Peltoperlidae) from the Lower Cretaceous of the West Transbaikalia, Russia". Zootaxa. 5523 (2): 284–290. doi:10.11646/zootaxa.5523.2.9.
[8]
Ryzhkova, Olga V.; Yao, Yunzhi; Kopylov, Dmitry S. (July 2023). "New fossil Kobdocoridae (Heteroptera: Pentatomomorpha) from the Lower Cretaceous of Transbaikalia, Eastern Russia". Cretaceous Research. 147 105509. doi:10.1016/j.cretres.2023.105509.
[9]
Shcherbakov, D. E. (December 2020). "New Homoptera from the Early Cretaceous of Buryatia with notes on the insect fauna of Khasurty". Russian Entomological Journal. 29 (1): 127–138. doi:10.15298/rusentj.29.2.02.
[10]
Kopylov, D. S. (May 2018). "New Ichneumonids (Hymenoptera: Ichneumonidae) from the Lower Cretaceous Khasurty Locality". Paleontological Journal. 52 (3): 284–289. doi:10.1134/S003103011803005X.
[11]
Kopylov, D. S. (July 2011). "Ichneumon wasps of the Khasurty locality in Transbaikalia (Insecta, Hymenoptera, Ichneumonidae)". Paleontological Journal. 45 (4): 406–413. doi:10.1134/S0031030111040058.
[12]
Li, Yi; Wang, Mei; Rasnitsyn, Alexandr P.; Shih, Chungkun; Zhuang, Jialiang; Ren, Dong (July 2024). "Two new species of Ghilarellinae (Hymenoptera, Cephoidea, Sepulcidae) from the Lower Cretaceous". Cretaceous Research. 159 105875. doi:10.1016/j.cretres.2024.105875.
[13]
Kopylov, D. S.; Rasnitsyn, A. P. (May 2017). "New sepulcids (Hymenoptera: Sepulcidae) from the Lower Cretaceous of Asia: II. Ghilarellinae and Trematothoracinae". Paleontological Journal. 51 (3): 291–303. doi:10.1134/S0031030117030029.
[14]
Kopylov, D. S. (February 2014). "New archaeocynipids (Hymenoptera, Cynipoidea, Archaeocynipidae) from the Lower Cretaceous of Transbaikalia". Paleontological Journal. 48 (1): 58–64. doi:10.1134/S0031030114010067.
[15]
Kopylov, D. S.; Rasnitsyn, A. P. (January 2017). "New sepulcids (Hymenoptera: Sepulcidae) from the Lower Cretaceous of Asia. I. Parapamphiliinae and Xyelulinae". Paleontological Journal. 51 (1): 69–76. doi:10.1134/S0031030117010087.
[16]
Kopylov, Dmitry S. (9 May 2019). "New anaxyelids (Hymenoptera: Anaxyelidae) from the Cretaceous of Asia". Zootaxa. 4603 (2). doi:10.11646/ZOOTAXA.4603.2.7.
[17]
Kopylov, D. S. (January 2012). "New species of Praeichneumonidae (Hymenoptera, Ichneumonoidea) from the Lower Cretaceous of Transbaikalia". Paleontological Journal. 46 (1): 66–72. doi:10.1134/S0031030112010078.
[18]
Khramov, Alexander V. (June 2016). "A new assemblage of Early Cretaceous green lacewings (Chrysopidae: Neuroptera) from Transbaikalia". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 107 (2–3): 195–202. doi:10.1017/S1755691017000342.
[19]
Khramov, Alexander V. (3 October 2021). "The youngest record of Priscaenigmatidae (Insecta: Neuropterida) from the Mesozoic of Kazakhstan and Russia". Historical Biology. 33 (10): 2432–2437. doi:10.1080/08912963.2020.1800683.
[20]
Strelnikova, O. D. (May 2019). "New Cupedids (Insecta: Coleoptera, Cupedidae) from the Lower Cretaceous of Buryatia". Paleontological Journal. 53 (3): 292–299. doi:10.1134/S0031030119030146.
[21]
Yan, Evgeny V.; Strelnikova, Olesya D.; Ponomarenko, Alexander G. (14 December 2021). "Jurodidae (Coleoptera: Archostemata) from Transbaikalia: new findings and redescription of type material". Palaeoentomology. 4 (6). doi:10.11646/palaeoentomology.4.6.9.
[22]
Strelnikova, O. D.; Yan, E. V.; Vasilenko, D. V. (November 2020). "A New Agyrtid Beetle (Coleoptera, Agyrtidae) from the Lower Cretaceous Khasurty Locality". Paleontological Journal. 54 (6): 627–631. doi:10.1134/S003103012006009X.
[23]
Prokin, Alexander A.; Strelnikova, Olesya D. (July 2021). "The first remarkable fossil Hydrochidae (Coleoptera) from the Lower Cretaceous (Hauterivian) of the Buryatia Republic, Russia". Cretaceous Research. 123 104795. doi:10.1016/j.cretres.2021.104795.
[24]
Ponomarenko, A. G. (April 2023). "A New Species of Ground Beetle (Insecta, Coleoptera, Trachypachidae) from the Lower Cretaceous Khasurty Locality, Western Transbaikalia". Paleontological Journal. 57 (2): 196–198. doi:10.1134/S0031030123010112.
[25]
Bashkuev, Alexey S. (August 2023). "Morphology and relationships of the bittacid genus Antiquanabittacus Petrulevičius et Jarzembowski, 2004 (Mecoptera), with description of three new species from the Lower Cretaceous of south-east Siberia". Cretaceous Research. 148 105549. doi:10.1016/j.cretres.2023.105549.
[26]
Ansorge, Jörg (1995). "Revision der liassischen "Permochoristidae" (Stammgruppenvertreter der Antliophora = Mecoptera + Diptera)" (PDF). Studia dipterologica. 2 (1): 21–26.
[27]
Lukashevich, E. D. (November 2020). "Rare Nematoceran Dipterans (Insecta: Diptera) from the Khasurty Locality, Transbaikalia". Paleontological Journal. 54 (6): 600–612. doi:10.1134/S0031030120060040.
[28]
Lukashevich, Elena D.; Blagoderov, Vladimir A. (8 January 2020). "Review of Mesozoic Perissommatidae (Insecta: Diptera)". Zootaxa. 4718 (4). doi:10.11646/zootaxa.4718.4.3.
[29]
Blagoderov, Vladimir A.; Lukashevich, Elena D. (1 December 2013). "New Axymyiidae (Insecta: Diptera) from the Mesozoic of East Siberia". Polish Journal of Entomology / Polskie Pismo Entomologiczne. 82 (4): 257–271. doi:10.2478/v10200-012-0040-9.
[30]
Lukashevich, Elena D.; Pepinelli, Mateus; Currie, Douglas C. (1 February 2020). "A new family and genus of Mesozoic Simulioidea (Insecta: Diptera)". Journal of Systematic Palaeontology. 18 (3): 213–231. doi:10.1080/14772019.2019.1588796.
[31]
Greenwalt, Dale E.; Blagoderov, Vladimir A. (18 March 2019). "Review of the fossil record of Bolitophilidae, with description of new taxa and discussion of position of Mangas Kovalev (Diptera: Sciaroidea)". Zootaxa. 4567 (3). doi:10.11646/zootaxa.4567.3.6.
[32]
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