Paleobiota of the Green River Formation

The Green River Formation is a geological formation located in the Intermountain West of the United States, in the states of Colorado, Utah, and Wyoming. It comprises sediments deposited during the Early Eocene in a series of large freshwater lakes: Lake Gosiute, Lake Uinta, and Fossil Lake (the last containing Fossil Butte National Monument). It preserves a high diversity of freshwater fish, birds, reptiles, and mammals, with some sections of the formation (including Fossil Lake and the Parachute Creek member of Lake Uinta) qualifying as Konservat-Lagerstätten due to their extremely well-preserved fossils.

Family	Genus	Species	Member	Abundance	Notes	Image
†Asterotrygonidae	†Asterotrygon	†A. maloneyi Carvalho, Maisey & Grande, 2004	Fossil Lake		An Asterotrygonidae stingray of uncertain higher affinities.^[1]^[2]
†Heliobatidae	†Heliobatis	†H. radians Marsh, 1877	Fossil Lake		A Heliobatid stingray of uncertain higher affinities. H. radians synonymy Dasyatis species Haseman, 1912 Palaeodasybatis discus Fowler, 1947 Xiphotrygon acutidens Cope, 1879 Xyphotrygus species Romer, 1971

Family	Genus	Species	Member	Notes
Lepisosteidae	Atractosteus	†A. atrox (Leidy, 1873)	Fossil Lake	An alligator gar relative. A. atrox is the largest Green River gar.
	Atractosteus	†A. simplex (Leidy 1873)	Fossil Lake Lake Gosiute?	An alligator gar relative.
	†Cuneatus	†C. cuneatus (Cope, 1884)	Fossil Lake? Lake Gosiute Lake Uinta	A Cuneatini tribe gar.
	†Cuneatus	†C. wileyi Grande, 2010	Lake Uinta	A Cuneatini tribe gar.
	Lepisosteus	†L. bemisi Grande, 2010	Fossil Lake	A Lepisosteini tribe gar.^[4]
	†Masillosteus	†M. janeae Grande, 2010	Fossil Lake	A Cuneatini tribe gar.

Family	Genus	Species	Member	Abundance	Notes	Image
Amiidae	Amia	†A. pattersoni Grande & Bemis, 1998	Fossil Lake Lake Gosiute		An amiid bowfin.
Amiidae	†Cyclurus	†C. gurleyi (Romer & Fryxell, 1928)	Fossil Lake	Only 8 specimens known	An amiid related to modern bowfins.

Family	Genus	Species	Member	Abundance	Notes	Image
Osteoglossidae	†Phareodus	†P. encaustus (Cope, 1871)	Fossil Lake Lake Gosiute		A phareodontine bonytongue.
Osteoglossidae	†Phareodus	†P. testis (Leidy, 1873)	Fossil Lake		A phareodontine bonytongue.

Family	Genus	Species	Member	Abundance	Notes
Clupeidae	†Gosiutichthys	†G. parvus Grande, 1982	Lake Gosiute		A clupeine clupeid herring relative.^[5]
	†Knightia	†K. alta (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		A pellonuline clupeid herring relative.^[6]
	†Knightia	†K. eocaena Jordan, 1907	Fossil Lake Lake Gosiute Lake Uinta	The most common articulated vertebrate fossil worldwide.^[7]	A pellonuline clupeid herring relative.

Family	Genus	Species	Member	Abundance	Notes	Image
†Hypsidoridae	†Hypsidoris	†H. farsonensis Lundberg & Case, 1970	Lake Gosiute		A hypsidorid catfish.
†Astephidae	†Astephus	†A. antiquus (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		An astephid catfish.^[9]

Family	Genus	Species	Member	Abundance	Notes	Image
Percopsidae	†Amphiplaga	†A. brachyptera Cope, 1877	Fossil Lake		A trout-perch relative. Includes the jr synonym Erismatopterus endlichi (1877)^[10]
Percopsidae	†Erismatopterus	†E. levatus (Cope, 1870)	Lake Gosiute Lake Uinta		A trout-perch relative. First described as Cyprinodon levatus (1870) includes the jr synonym Erismatopterus rickseckeri (1871)^[10]

Family	Genus	Species	Member	Abundance	Notes
Moronidae	†Cockerellites	†C. liops (Cope, 1877)	Fossil Lake	common	A planktivorous/insectivorous temperate bass.^[7]^[2] First described as Priscacara liops (1877) First moved to Cockerellites liops in 1923.^[2]
	†Hypsiprisca	†H. hypsacantha (Cope, 1886)	Fossil Lake	rare	A temperate bass.^[7] First named Priscacara hypsacantha 1886)^[2]
	†Hypsiprisca	†H. sp.	Fossil Lake	rare	An undescribed species of Hypsiprisca. A slenderer species with a more rounded tail fin outline.^[2]
	†Priscacara	†P. serrata Cope, 1877	Fossil Lake Lake Uinta		A temperate bass.^[7]
		†P. sp. a	Fossil Lake	1 specimen	An undescribed Priscacara Specimens have a rounder body and weak dorsal spines.^[2]
		†P. sp. b.	Fossil Lake	1 specimen	An undescribed Priscacara Possibly a new species or a variant specimen with 4 anal spines.^[2]

Family	Genus	Species	Member	Abundance	Notes	Image
†Asineopidae	†Asineops	†A. squamifrons Cope, 1870	Fossil Lake Lake Gosiute Lake Uinta		An acanthomorph of uncertain affinities Possibly a percopsiform, polymixiiform, or perciform.^[3]
†Asineopidae	†Asineops	A. sp.	Fossil Lake		An undescribed Asineops species Restricted to Fossil Lake.^[3]

Family	Genus	Species	Member	Abundance	Notes
Boidae	†Boavus	†B. idelmani Gilmore, 1938	Fossil Lake		A boa.
Dactyloidae	†Afairiguana	†A. avius Conrad et al., 2007	Fossil Lake		A stem-anole.^[13]
Shinisauridae	†Bahndwivici	†B. ammoskius Conrad, 2006	Fossil Lake		A relative of the Chinese crocodile lizard.
Tropidophiidae?	†Dunnophis	D. sp.	Fossil Lake		An undescribed possible dwarf boa.^[14]
Varanidae	†Saniwa	†S. ensidens Leidy, 1870	Fossil Lake		A stem-monitor lizard.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A possible gecko or iguanid Preserved with soft tissue and skin, but bones dissolved.^[15]
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A relative of Shinisaurids Preserved with soft tissue and skin, but bones dissolved.^[16]

Family	Genus	Species	Member	Abundance	Notes
Alligatoridae	†Tsoabichi	†T. greenriverensis Brochu, 2010	Fossil Lake		A caiman.^[17]
Incertae sedis	†Borealosuchus	†B. wilsoni (Mook, 1959)	Fossil Lake		A eusuchian crocodyliform of uncertain affinities. Moved from Leidyosuchus wilsoni (1959)
	†Borealosuchus	Cf. †B. sp.	Fossil Lake		Indeterminate eusuchian bones close to Borealosuchus.^[18]
	"Crocodylus"	†"Crocodylus" acer Cope, 1882	Lake Uinta (Manti beds)	1 specimen	A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.^[19]
	"Crocodylus"	Cf. †"Crocodylus" affinis Cope, 1871	Lake Gosiute		Bones close in morphology to †"Crocodylus" affinis A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.^[19]

Family	Genus	Species	Member	Abundance	Notes
Baenidae	†Chisternon	†C. undatum (Leidy, 1871)	Fossil Lake		A baenid.
Dermatemydidae	†Baptemys	†B. wyomingensis Leidy, 1873	Fossil Lake		A dermatemydid.
Dermatemydidae	†Baptemys	†B. Cf. wyomingensis Leidy, 1873	Lake Unita		A dermatemydid.^[18]
Geoemydidae	†Echmatemys	†E. haydeni (Cope, 1873)		1 specimen	A geoemydid tortoise.^[20]
		†E. naomi (Leidy, 1871)	Fossil Lake	1 specimen	A geoemydid tortoise.^[20]
		†E. wyomingensis Leidy, 1872	Fossil Lake		A geoemydid tortoise.^[21]
Testudinidae	†Hadrianus	†Cf. †H. corsoni (Leidy, 1871)	Fossil Lake		A tortoise species. A single complete hatchling^[20]
Trionychidae	†Axestemys	A. sp.	Fossil Lake		A softshell turtle.
	†Oliveremys	†O. uintaensis Cope, 1873	Fossil Lake		A softshell turtle.
	†Plastomenus	†P. thomasii (Cope, 1872	Fossil Lake		A softshell turtle.

Family	Genus	Species	Member	Abundance	Notes	Image
†Lithornithidae	†Lithornis	†L. grandei	Fossil Lake		A lithornithid.^[22] First described as Calciavis grandei.^[23]
†Lithornithidae	†Pseudocrypturus	†P. cercanaxius	Fossil Lake		A lithornithid.

Family	Genus	Species	Member	Abundance	Notes	Image
†Presbyornithidae	†Presbyornis	†P. pervetus	Fossil Lake		A presbyornithid anseriform.
Incertae sedis	†Paakniwatavis	†P. grandei	Fossil Lake		An anseriform of uncertain affinities.

Family	Genus	Species	Member	Abundance	Notes	Image
†Sandcoleidae	†Anneavis	†A. annae	Fossil Lake		A sandcoleid mousebird.
Incertae sedis	†Celericolius	†C. acriala	Fossil Lake		A stem-mousebird.^[24]

Family	Genus	Species	Member	Abundance	Notes
Eocypselidae	†Eocypselus	†E. rowei	Fossil Lake		An eocypselid apodiform.^[28]
Fluvioviridavidae	†Fluvioviridavis	†F. platyrhamphus Mayr & Daniels, 2001	Fossil Lake	2 specimens	A basal strisore, possibly a stem-frogmouth.^[29]^[30]
Incertae sedis	†Prefica	†P. nivea	Fossil Lake		A stem-oilbird.^[31]

Family	Genus	Species	Member	Abundance	Notes	Image
Fregatidae	†Limnofregata	L. azygosternon	Fossil Lake		An early frigatebird.
Fregatidae	†Limnofregata	L. hasegawai	Fossil Lake		An early frigatebird

Family	Genus	Species	Member	Abundance	Notes	Image
Threskiornithidae	†Rhynchaeites	†R. mcfaddeni^[34]		1 specimen	Known from a largely complete specimen.^[35]	R. messelensis, a related species
Threskiornithidae	†Vadaravis	†V. brownae	Fossil Lake		Potentially a stem-ibis, but may be an indeterminate aequornithine.^[35]^[36]

Family	Genus	Species	Member	Notes
†Avipedidae	†Avipeda	Cf. †A. phoenix Vyalov, 1960	Lake Unita	Avian trackways with three forward facing toes without inter-digit webbing.^[45]
†Ignotornidae	†Quadridigitus	†Q. semimembranus Lockley et al., 2021	Lake Unita	Avian trackways with three forward facing toes, a hallux, and inter-digit webbing.^[45]
†Jindongornipodidae	†Jindongornipes	†J. falkbuckleyi Lockley et al., 2021	Lake Unita	Large avian trackways with four toes, and wide spread between digits II and IV.^[45]
Incertae sedis	†Presbyornithiformipes	†P. feduccii Yang et al., 1995	Lake Uinta	Duck-like trackways with dabble marks attributed to Presbyornis.^[46]

Family	Genus	Species	Member	Abundance	Notes	Image
incertae sedis	†Plesiocathartes	†P. major Weidig, 2006	Fossil Lake		A stem-cuckoo roller.^[25]
incertae sedis	†Plesiocathartes	†P. wyomingensis Weidig, 2006	Fossil Lake		A stem-cuckoo roller

Family	Genus	Species	Member	Abundance	Notes	Image
Primobucconidae	†Primobucco	†P. mcgrewi	Fossil Lake		A primobucconid coraciiform.
Incertae sedis	†Paracoracias	†P. occidentalis			A coraciiform.^[26]

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Neanis	†N. schucherti	Fossil Lake		A possible piciform of uncertain affinities.
Incertae sedis	†Pristineanis	†P. kistneri	Fossil Lake (Fossil Butte Member) Lake Gosiute (Tipton tongue Member)	2 specimens	A possible piciform of uncertain affinities. First described as Primobucco kistneri (1973) Moved from Neanis kistneri 1976)^[27]

Family	Genus	Species	Member	Notes
†Halcyornithidae	†Cyrilavis	†C. colburnorum	Fossil Lake	A halcyornithid.
†Halcyornithidae	†Cyrilavis	†C. olsoni	Fossil Lake	A halcyornithid.
†Messelasturidae	†Tynskya	†T. eocaena	Fossil Lake	A messelasturid.
†Morsoravidae	†Consoravis	†C. turdirostris	Fossil Lake	A morsoravid.^[38]
†Psittacopedidae	†Eofringillirostrum	†E. boudreauxi	Fossil Lake	A psittacopedid.^[39]
†Vastanavidae	†Avolatavis	†A. tenens	Fossil Lake	A vastanavid.^[40]
†Zygodactylidae	†Eozygodactylus	†E. americanus	Fossil Lake	A zygodactylid.^[41]
†Zygodactylidae	†Zygodactylus	†Z. grandei	Fossil Lake	A zygodactylid.^[42]

Family	Genus	Species	Member	Abundance	Notes	Image
†Herpetotheriidae	†Herpetotherium	†H. innominatum	Lake Gosiute Lake Uinta		An opossum-like basal metatherian Originally described as Peratherium innominatum.	H. huntii, a related species
†Herpetotheriidae	†Herpetotherium	†H. knighti	Lake Gosiute		An opossum-like basal metatherian.	H. huntii, a related species

Family	Genus	Species	Member	Abundance	Notes
†Icaronycteridae	†Icaronycteris	†I. gunnelli	Fossil Lake	2 specimens	An icaronycterid bat.^[47]
†Icaronycteridae	†Icaronycteris	†I. index	Fossil Lake	4 specimens	An icaronycterid bat.^[47]
†Onychonycteridae	†Onychonycteris	†O. finneyi	Fossil Lake	2 specimens	An onychonycterid bat.^[47]

family	Genus	Species	Member	Abundance	Notes	Image
Nyctitheriidae	Nyctitherium	N. nitidum	Lake Gosiute		A nyctitheriid. Originally described as Talpavus nitidus.^{[clarification needed]}
Nyctitheriidae	Nyctitherium	N. sp.	Lake Uinta

Family	Genus	Species	Member	Notes	Image
Miacidae	Vulpavus	V. profectus	Lake Gosiute	A miacid.
Miacidae	Indeterminate	indeterminate	Lake Gosiute Lake Uinta	Indeterminate miacid bones.^[48]
Sinopinae	Sinopa	S. minor	Lake Uinta	A sinopid hyaenodont.	S. rapax, a related species
Viverravidae	Viverravus	V. minutus	Lake Uinta	A viverravid.
Incertae sedis	"Miacis"	"M." gracilis	Lake Uinta	A carnivoraform of uncertain affinities.

family	Genus	Species	Member	Abundance	Notes
†Brontotheriidae	†Lambdotherium	†L. popoagicum	Fossil Lake		A small-sized basal brontothere.
†Brontotheriidae	†L. sp.	Lake Unita		A small-sized basal brontothere. Reported from Powder Wash, Utah by Cope (1880)^[49]
Equidae	†Protorohippus	†Aff. P. venticolus	Fossil Lake	2 specimens	An equid closely affiliated with P. venticolus.^[50]
Equidae	Indeterminate	Indeterminate	Lake Gosiute Lake Unita<	Indeterminate equiid bones. First identified as Hyracotherium sp.
†Hyopsodontidae	†Hyopsodus	†H. minusculus	Lake Gosiute Lake Uinta		A hyopsodontid.
		†H. paulus (=H. vicarius)	Lake Uinta		A hyopsodontid. includes H. vicarius specimens.^{[citation needed]}
		†H. wortmani	Fossil Lake		A hyopsodontid.
↑Hyrachyidae	↑Hyrachyus	Indeterminate	Lake Unita		Undeterminate Hyrachyus bones.^[18]
Indeterminate	Indeterminate	Indeterminate	Fossil Lake	1 specimen	A juvenile tapiromorph, the largest mammal known from the formation. Initially described as Heptodon calciculus.^[51]

Family	Genus	Species	Member	Notes	Image
Ischyromyidae	Paramys	P. sp. ^{[citation needed]}	Lake Uinta	An ischyromyid rodent.	P. delicatus
Ischyromyidae	Thisbemys	T. sp. ^{[citation needed]}	Lake Gosiute	An undescribed ischyromyid rodent.^{[citation needed]}	T. plicatus
Sciuravidae	Sciuravus	S. eucristadens (Burke, 1937)	Lake Uinta (Douglas Creek Member)	A sciuravid rodent. Originally described as Viverravus eucristadens (1937).
Sciuravidae	Sciuravus	S. sp.^{[citation needed]}	Lake Uinta	An undescribed sciuravid.

Acipenseriformes

Lepisosteiformes

Amiiformes

Hiodontiformes

Osteoglossiformes

Ellimmichthyiformes

Clupeiformes

Gonorynchiformes

Cypriniformes

Siluriformes

Salmoniformes

Percopsiformes

Carangiformes

Acanthuriformes

Incertae sedis

Frogs

Salamanders

Squamates

Crocodilians

Turtles

Lithornithiformes

Anseriformes

Galliformes

Coliiformes

Leptosomiformes

Coraciiformes

Piciformes

Strisores

Musophagiformes

Mirandornithes

Suliformes

Pelecaniformes

Charadriiformes

Gruiformes

Eufalconimorphae

Neoaves incertae sedis

Avian ichnofossils

Metatheria

Cimolesta

Chiroptera

Eulipotyphla

Pan-Carnivora

Paraxonia

Pan-Perissodactyla

Apatotheria

Rodentia

Primatomorpha

Arachnida

Acariformes

Araneae

Parasitiformes

Scorpiones

Crustacea

Branchiopoda

Decapods

Ostracoda

Myriapoda

Insecta

Coleoptera

Adephaga

Elateriformia

Polyphaga - Bostrichiformia

Polyphaga - Cucujiformia

Polyphaga - Scarabaeiformia

Polyphaga - Staphyliniformia

Dictyoptera

Diptera

Hemiptera

Hymenoptera

Lepidoptera

Mantodea

Megaloptera

Neuroptera

Raphidioptera

Odonata

Orthoptera

Psocodea

Strepsiptera

Thysanoptera

Trichoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Microsyopidae	Microsyops	M. elegans	Lake Gosiute		A microsyopid stem-primate.	M. cardiorestes, a related species
Microsyopidae	Uintasorex	U. parvulus	Lake Gosiute		A microsyopid stem-primate.
Notharctidae	Notharctus	N. sp.	Lake Gosiute		A notharctid adapiform.	N. tenebrosus
Notharctidae	Smilodectes	S. gracilis	Lake Uinta		A notharctid adapiform. Formerly included in Notharctus matthewi.
Omomyidae	Anemorhysis	A. sp.	Lake Unita	1 specimen (lost?)	An omomyid primate. First named as Uintalacus nettingi (1958) Deemed Anemorhysis sp indeterminate (1976)^[53]
	Omomys	O. carteri	Lake Uinta		An omomyid primate. Includes O. pucillus^[54]
	Tetonius	T. sp.	Lake Uinta		An omomyid primate.	T. homunculus
	Washakius	W. insignis	Lake Gosiute		An omomyid primate.

Family	Genus	Species	Member	Abundance	Notes
Araneidae	Araneus	A. sp	Lake Gosiute	6 specimens	Possible orbweb-spiders, known from partial fossils First placed as Epeira sp.^[56]
Hersiliidae	Hersiliola (sensu lato)	H. sp.	Lake Uinta (Parachute Creek Member)	1 male	A tree trunk spider.^[57]
Selenopidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	1 specimen	A female or juvenile selenopid spider.^[57]
Thomisidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	2 males, 1 juv female	Crab spiders.^[57]
Uloboridae	Miagrammopes	M. sp.	Lake Uinta (Parachute Creek Member)	1 female, 2 males	A uloborid spider.^[57]
Uloboridae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)		A uloborid spider.^[57]
Unidentified	†Araneaovoius (Scudder, 1878)	†A. columbiae^[58]		1 specimen	An orb-web spider egg sack. First described as "Aranea" columbiae (1878)^[59]^[56]

Family	Genus	Species	Member	Abundance	Notes	Image
Cyclestheriidae	†Cyclestherioides	†C. wyomingensis	Lake Gosiute (Laney)		A cyclestheriid clam shrimp.^[62]
Lynceidae	†Prolynceus	†P. laneyensis	Lake Gosiute (Laney)		A clam shrimp.^[62]

Family	Genus	Species	Member	Abundance	Notes	Image
Palaemonidae	†Bechleja	†B. rostrata	Fossil Lake		A palaemonid shrimp.^[63]
Cambaridae	Procambarus	†P. primaevus	Fossil Lake		A cambarid crayfish.

Family	Genus	Species	Member	Notes
Candonidae	Candona	C. artesensis Swain, 1964	Fossil Lake Lake Unita	A candonid ostracod.^[64]
		C. rangliensis Swain, 1964	Fossil Lake Lake Unita	A candonid ostracod.^[64]
		C. whitei Scott & Smith, 1951	Fossil Lake Lake Gosiute Lake Unita	A candonid ostracod.^[64]
	Tuberocypris	T. centronotus Swain, 1964	Lake Unita	A candonid ostracod.^[64]
†Cyprideidae	†Cypridea	†C. bisulcata Swain, 1949	Lake Unita	A cyprideid ostracod.^[64]
Cyprididae	†Advenocypris	†A. umbonata Swain, 1964	Lake unita	A cypridid ostracod.^[64]
	Hemicyprinotus	†H. watsonensis Swain, 1964	Lake unita	A cypridid ostracod. One described subspecies H. watsonensis emaciata^[64]
	Heterocypris	†H. riflensis Swain, 1964	Lake unita	A cypridid ostracod.^[64]
	Heterocypris	†H. whiteriverensis Swain, 1964	Lake unita	A cypridid ostracod.^[64]
	Potamocypris	†P. williamsi Swain, 1964	Fossil Lake Lake Unita	A cypridid ostracod.^[64]
	†Procyprois	†P. ravenridgensis Swain, 1964	Lake Unita	A cypridid ostracod. Male and female specimens show shell sexual dimorphism^[64]
	Pseudoeucypris	†P. pagei (Swain) Swain, 1956	Lake Unita	A cypridid ostracod. First described as Candona pagei (1949) Moved from Cypris pagei (1956)^[64]
	Scottia	†S. subquadrata Swain, 1964	Lake Unita	A cypridid ostracod.^[64]
	Stenocypris	Cf. S. sinuata G.W. Müller, 1898	Lake Unita	A cypridid ostracod.^[64]
Limnocytheridae	Limnocythere	L. sp.	Lake Unita	Indistinct limnocytherid fossils.^[64]
Limnocytheridae	Metacypris	†M. paracordata Swain, 1964	Lake Unita	A limnocytherid ostracod.^[64]
Notodromadidae	Cyprois	†C. ephraimensis Swain, 1964	Fossil Lake Lake Gosiute Lake Unita	A notodromadid ostracod.^[64]

Family	Genus	Species	Member	Abundance	Notes
Carabidae	Bembidion	†B. exoletum Scudder, 1876	Lake Unita	1 specimen	A Bembidiini tribe trechine ground beetle.^[67]
	†Carabites	†C. eocenicus Cockerell, 1921	Lake Unita		A ground beetle of uncertain subfamily affiliation.^[68]
	†Carabites	†C. exanimus Scudder, 1892	Lake Uinta	1 specimen	A ground beetle of uncertain subfamily affiliation.^[69]
	Harpalus	†H. veterum Cockerell, 1921	Lake Uinta		A Harpalini tribe harpaline ground beetle.^[68]
	Lebia	†L. protospiloptera Cockerell, 1921	Lake Unita		A Lebiini tribe harpaline ground beetle.^[70]
	†Neothanes	†N. testeus (Scudder, 1878)	Lake Gosiute	2 specimens	A ground beetle of uncertain subfamily affiliation.^[56] First named as Cychrus testeus (1878)^[66]
	Platynus	†P. caesus Scudder, 1890	Lake Gosiute	2 specimens	A Platynini tribe platynine ground beetle.^[56]
	Platynus	†P. senex Scudder, 1878	Lake Gosiute	1 specimen	A Platynini tribe platynine ground beetle.^[66]
Dytiscidae	Laccophilus	Indeterminate	Lake Uinta	1 fragment	An indeterminate Laccophilini tribe predaceous diving beetle.^[67]

Family	Genus	Species	Member	Abundance	Notes
Byrrhidae	"Nosodendron"	†"N." tritavum Scudder, 1890	Lake Gosiute	1 specimen	A pill beetle in need of redescription.^[71] First named as a Nosodendrid beetle.^[56]
Elateridae	Cardiophorus	†C. exhumatus Cockerell, 1925	Lake Unita	1 specimen	A cardiophorine click beetle.^[72]
	Ctenicera	†C. velata (Scudder, 1876)	Lake Gosiute	1 specimen	A Prosternini tribe dendrometrine click beetle.^[73] First named as Corymbites velatus (1876).^[66]
	Oxygonus	†O. mortuus Scudder, 1876	Lake Uinta	1 specimen	A Prosternini tribe dendrometrine click beetle.^[67]
Eucnemidae	Epiphanis	†E. deletus Scudder, 1876	Lake Unita	1 specimen	An Epiphanini tribe melasine false click beetle.^[67]

Family	Genus	Species	Member	Abundance	Notes
Ptinidae	Anobium	†A. deceptum Scudder, 1878	Lake Gosiute	1 specimen	An Anobiini tribe anobiine death-watch beetle.^[66]^[74]
		†A. lignitum Scudder, 1878	Lake Gosiute	1 specimen	An Anobiini tribe anobiine death-watch beetle.^[66]^[74]
		†A. ovale Scudder, 1878	Lake Gosiute	2 specimens	An Anobiini tribe anobiine death-watch beetle.^[66]^[74]
	Stegobium	†S. defunctus (Scudder, 1876)	Lake Gosiute	1 specimen	A Stegobiini tribe anobiine death-watch beetle.^[75] First described as Sitodrepa defuncta (1876).^[67]^[66]

Family	Genus	Species	Member	Abundance	Notes
Anthribidae	Choragus	†C. fictilis Scudder, 1890	Lake Gosiute	1 specimen	A Choragini tribe choragine fungus weevil.^[56]^[76]
	Euparius	†E. elusus (Scudder, 1878)	Lake Gosiute	2 specimens	A Cratoparini tribe anthribine fungus weevil.^[76] First named as Cratoparis? elusus (1878)^[66]
	Euparius	†E. repertus (Scudder, 1878)	Lake Gosiute	1 specimen	A Cratoparini tribe anthribine fungus weevil.^[76] First named as Cratoparis repertus (1878)^[66]
	Ormiscus	†O. partitus (Scudder, 1890)	Lake Gosiute	1 specimen	A Zygaenodini tribe anthribine fungus weevil.^[76] First named as Hormiscus partitus (1890)^[56]
	†Pseudochirotenon	†P. eocaenicus Legalov, 2018	Lake Unita	1 specimen	An Ecelonerini tribe anthribine fungus weevil.^[77]
	Trigonorhinus	†T. pristinus (Scuddr, 1876)	Lake Gosiute	1 specimen	A Trigonorhinini tribe anthribine fungus weevil.^[76] First named as Brachytarsus pristinus (1876)^[66]
	Tropideres	†T. remotus Scudder, 1893	Lake Gosiute	1 specimen	A Tropiderini tribe anthribine fungus weevil.^[78]^[76]
Attelabidae	†Teretrum	†T. quiescitum Scudder, 1893	Lake Unita	1 specimen	An Auletini tribe rhynchitine leaf-rolling weevil.^[78]^[76]
Belidae	†Paltorhynchus	†P. bisulcatus Scudder, 1893	Lake Unita	2 specimens	A Metrioxenini tribe oxycorynite belid weevil.^[78]^[79]
Brentidae	Apionion	†A. evestigatum (Scudder, 1893)	Lake Unita	1 specimen	An Apionini tribe apionine primitive weevil.^[78]^[77]
	†Palaeonanophyes	†P. zherikhini Legalov, 2015	Lake Unita		A Nanophyini tribe nanophyine primitive weevil.^[80]
	Perapion	†P. rasnitsyni Legalov, 2018	Lake Uinta	1 specimen	An Aplemonini tribe apionine primitive weevil.^[77]
Cerambycidae	Clytus	†C. pervetustus (Cockerell, 1921)	Lake Uinta		A Clytini tribe cerambycine long-horn beetle.^[81] First named as Lema pervetusta (1921)^[68]
Cerambycidae	Saperda	†S. caroli Vitali, 2015	Lake Unita		A Saperdini tribe lamiine long-horn beetle.^[82]
Chrysomelidae	Bruchus	†B. anilis Scudder, 1876	Lake Unita	1 specimen	A Bruchini tribe bruchine leaf beetle.^[67]
	Cryptocephalus	†C. vetustus Scudder, 1878	Lake Gosiute	2 specimens	A Cryptocephalini tribe cryptocephaline leaf beetle.^[66]
	†Denaeaspis	†D. chelonopsis Chaboo & Engel, 2009	Lake Unita	1 specimen	An Imatidiini tribe Cassidine tortoise beetle.^[83]
	†Eosacantha	†E. delocranioides Chaboo & Engel, 2009	Lake Unita	1 specimen	A Notosacanthini tribe Cassidine tortoise beetle.^[83]
	†Palaeatalasis	†P. monrosi Legalov, 2021	Lake Unita	1 specimen	A Megamerini tribe sagrine leaf beetle.^[84]
	†Pulchritudo	†P. attenboroughi Krell & Vitali, 2021	Lake Uinta	1 specimen	A sagrine leaf beetle^[85]
Cryptophagidae	Antherophagus	†A. priscus Scudder, 1876	Lake Gosiute	4 specimens	A Cryptophagini tribe cryptophagine silken fungus beetle.^[67]^[66]
Curculionidae	Anthonomus	†A. revictus Scudder, 1890	Lake Gosiute	2 specimens	An Anthonomini tribe curculionine weevil.^[56]^[78]^[76]
	Anthonomus	†A. soporus Scudder, 1890	Lake Gosiute Lake Unita	7 specimens	An Anthonomini tribe curculionine weevil.^[56]^[78]^[76]
	†Archaeoheilus	†A. deleticius (Scudder, 1893)	Lake Uinta	1 specimen	A Molytini tribe molytine weevil. First named as Pachylobius deleticius (1893) Moved from Hylobius deleticius (2015)^[77]
		†A. ovalis Legalov, 2018	Lake Uinta	1 specimen	A Molytini tribe molytine weevil.^[77]
		†A. packardii (Scudder, 1893)	Lake Gosiute	1 specimen	A Molytini tribe molytine weevil. First named as Hylobius packardii (1893).^[77]
		†A. provectus (Scudder, 1876)	Lake Gosiute	2 specimens	A Molytini tribe molytine weevil. First named as Hylobius provectus (1876).^[77]
		†A. scudderi Legalov, 2018	Lake Uinta	1 specimen	A Molytini tribe molytine weevil.^[77]
	†Calandrites	†C. cineratius Scudder, 1893	Lake Gosiute Lake Unita	13 specimens	A weevil of uncertain subfamilial affiliation.^[78]^[76]
	†Calandrites	†C. defessus Scudder, 1893	Lake Gosiute Lake Unita	4 specimens	A weevil of uncertain subfamilial affiliation.^[78]^[76]
	Ceutorhynchus	†C. degravatus Scudder, 1893	Lake Unita	1 specimen	A Ceutorhynchini tribe ceutorhynchine weevil.^[78]^[76]
	"Coniatus"	†"C." refractus Scudder, 1893	Lake Uinta		A Tropiphorini tribe entimine weevil of uncertain genus affiliations.^[78]^[86]
	Cossonus	†C. rutus Scudder, 1893	Lake Unita	1 specimen	A Cossonini tribe cossonine weevil.^[78]^[76]
	Cryptorhynchus	†C. annosus Scudder, 1893)	Lake Gosiute Lake Unita	3 specimen	A Cryptorhynchini tribe cryptorhynchine weevil.^[67]^[76]
	Cryptorhynchus	†"C". durus Scudder, 1893)	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil of uncertain generic affiliations.^[78]^[76]
	"Dryocoetes"	†"D". carbonarius Scudder, 1876	Lake Gosiute	1 specimen	A scolytine bark beetle.^[66] Considered as genus and tribe incertae sedis by Legalov (2015).^[76]
	Entimus	†E. primordialis Scudder, 1876	Lake Uinta	1 specimen	An Entimini tribe entimine weevil.^[67]^[77]
	Epicaerus	†E. dilapsus (Scudder, 1893)	Lake Gosiute	1 specimen	A Geonemini tribe entimine weevil.^[78]^[76] First named as Phyxelis dilapsus (1893).^[78]
		†E. effossus (Scudder, 1876)	Lake Gosiute lake Uinta	numerous	A Geonemini tribe entimine weevil.^[66]^[78]^[76] First named as Eudiagogus effossus (1876).^[67]
		†E. eradicatus (Scudder, 1893)	Lake Unita	3 specimens	A Geonemini tribe entimine weevil.^[78]^[76] First named as Phyxelis eradicatus (1893).^[78]
		†E. evigoratus (Scudder, 1893)	Lake Unita	3 specimens	A Geonemini tribe entimine weevil.^[78]^[76] First named as Phyxelis evigoratus (1893).^[78]
		†E. exanimis (Scudder, 1876)	Lake Gosiute Lake Unita	numerous	A Geonemini tribe entimine weevil.^[66]^[78]^[76] First named as Eudiagogus exanimis (1876).^[67]
		†E. excissus (Scudder, 1893)	Lake Unita	1 specimen	A Geonemini tribe entimine weevil.^[78]^[76] First named as Phyxelis excissus (1893).^[78]
		†E. fodinarum (Scudder, 1876)	Lake Gosiute	3 specimens	A Geonemini tribe entimine weevil.^[66]^[78]^[76] First named as Sitona fodinarum (1893).^[67]
		†E. saxatilis (Scudder, 1876)	Lake Gosiute Lake Unita	numerous	A Geonemini tribe entimine weevil.^[66]^[78]^[76] First named as Eudiagogus saxatilis (1876).^[67]
		†E. subterraneus (Scudder, 1893)	Lake Gosiute Lake Unita	Numerous	A Geonemini tribe entimine weevil.^[78]^[76] First named as Scythropus subterraneus (1893).^[78]
		†E. terrosus (Scudder, 1878)	Lake Gosiute Lake Uinta	4 specimens	A Geonemini tribe entimine weevil.^[66]^[76] First named as Eudiagogus terrosus (1878).^[67]
	Eudiagogus	†E. vossi Legalov, 2018	Lake Uinta	1 specimen	A Eudiagogini tribe entimine weevil.^[77]
	†Extinctocentrinus	†E. brevirostris Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.^[79]
	Geraeus	†G. anvilis Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.^[79]
		†G. diruptus (Scudder, 1893)	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.^[79] First named as Centrinus diruptus (1893)^[78]
		†G. fossilis Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.^[79]
	"Gymnetron"	†G. lecontei Scudder, 1878	Lake Gosiute	1 specimen	A curculionine weevil of uncertain genus and tribal affiliations.^[66]^[76]
	"Lepyrus"	†L. evictus (Scudder, 1893)	Lake Gosiute	1 specimen	A Tropiphorini tribe entimine weevil of uncertain genus affiliation.^[78]^[76]
	†Limalophus	†L. compositus (Scudder, 1893)	Lake Gosiute Lake Unita	3 specimens	A Tropiphorini tribe entimine weevil.^[78]^[76]
		†L. contractus (Scudder, 1893)	Lake Gosiute	6 specimens	A Tropiphorini tribe entimine weevil.^[78]^[76]
		†L. receptus (Scudder, 1893)	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil.^[76] First named as Artipus receptus (1893).^[78]
	Listronotus	†L. muratus Scudder, 1890	Lake Gosiute	1 specimen	A Listroderini tribe cyclomine weevil.^[56]^[76]
	†Lithogeraeus	†L. anicilla (Scudder, 1893)	Lake Unita	5 specimens	An Apostasimerini tribe conoderine weevil.^[79] First described as Aulobaris ancilla (1893)^[78]
		†L. circumscripta (Scudder, 1893)	Lake Unita	5 specimens	An Apostasimerini tribe conoderine weevil.^[79] First described as Aulobaris circumscripta (1893)^[78]
		†L. comminuta (Scudder, 1893)	Lake Unita	4 specimens	An Apostasimerini tribe conoderine weevil.^[79] First described as Aulobaris comminuta (1893)^[78]
		†L. greenriverensis Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.^[79]
	Mitostylus	†M. abacus (Scudder, 1893)	Lake Unita	1 specimen	A Sciaphilini tribe entimine weevil.^[76] First named as Scythropus abacus (1893).^[78]
		†M. obdurefactus (Scudder, 1893)	Lake Unita	5 specimens	A Sciaphilini tribe entimine weevil.^[76] First named as Exomias obdurefactus (1893).^[78]
		†M. seculorum (Scudder, 1890)	Lake Gosiute	1 specimen	A Sciaphilini tribe entimine weevil.^[76] First named as Tanymecus seculorum (1890).^[56]
	Neoptochus	Indeteminate	Lake Unita	2 specimens	Indeterminate Cyphicerini tribe Entiminae weevil fossils.^[78]
	Nicentrus	†N. curvirostris Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.^[79]
	Ophryastes	†"O". compactus Scudder, 1878	Lake Gosiute	1 specimen	A Tropiphorini tribe entimine weevil.^[66] of uncertain genus affinities (2011, 2015).^[87]^[76]
		†O. grandis Scudder, 1893	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil.^[78] Considered nomen dubium (2015).^[76]
		†O. petrarum Scudder, 1893	Lake Unita	2 specimens	A Tropiphorini tribe entimine weevil.^[78] Considered nomen dubium (2011, 2015).^[87]^[76]
	†Ophryastites	†O. digressus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain tribal affiliation.^[76]
	†Ophryastites	†O. dispertitus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain tribal affiliation.^[76]
	†Otiorhynchites	†O. commutatus Scudder. 1893	Lake Unita	1 specimen	A weevil of uncertain subfamilial affiliations.^[78]^[76]
		†O. fossilis Scudder. 1893	Fossil Lake	1 specimen	A weevil of uncertain subfamilial affiliations.^[78]^[76]
		†O. tysoni Scudder. 1893	Lake Gosiute Lake Unita	2 specimens	A weevil of uncertain subfamilial affiliations.^[78]^[76]
	Otiorhynchus	†"O". flaccus Scudder, 1893	Lake Gosiute	1 specimen	An Otiorhynchini tribe entimine weevil.^[78] Considered genus and subfamily incertae sedis by Legalov (2015).^[76]
		†O. subteractus Scudder, 1893	Lake Unita	1 specimen	An Otiorhynchini tribe entimine weevil.^[78] Considered genus incertae sedis by Legalov (2015).^[76]
		†"O". tumbae (Scudder, 1890)	Lake Gosiute	1 specimen	An entimine weevil.^[56] Considered genus and tribus incertae sedis by Legalov (2015).^[76] First named Otiorhynchus dubius (1878)^[66]
	"Pachylobius"	†"P". compressus Scudder, 1893	Lake Gosiute Lake Unita	9 specimens	A weevil elytron of uncertain subfamily and genus affiliations.^[78]^[76]
	"Pachylobius"	†"P". depraedatus Scudder, 1893	Lake Unita	4 specimens	A weevil elytron of uncertain subfamily and genus affiliations.^[78]^[76]
	†Palaeomallerus	†P. longirostris Legalov, 2018	Lake Uinta	1 specimen	A Palaeomallerini tribe conoderine weevil.^[79]
	"Phyllobius"	†"P". antecessor Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain subfamily and genus affiliations.^[78]^[76]
		†"P". avus Scudder, 1893	Lake Gosiute Lake Unita	3 specimens	A weevil of uncertain subfamily and genus affiliations.^[78]^[76]
		†"P". carcerarius Scudder, 1893	Lake Unita	2 specimens	A weevil of uncertain subfamily and genus affiliations.^[78]^[76]
	"Polygraphus"	†"P". wortheni (Scudder, 1876)	Lake Gosiute	1 specimen	A scolytine bark beetle.^[78] Considered genus and tribus incertae sedis by Legalov (2015)^[76]
Procas	†P. vinculatus Scudder, 1893	Lake Unita	2 specimens	An Erirhinini tribe erirhinine weevil.^[78]
†Primocentron	†P. wickhami Legalov, 2018	Lake Uinta	1 specimen	A Eustylini tribe entimine weevil.^[77]
†Pseudophaops	†P. perditus (Scudder, 1876)	Lake Gosiute	2 specimens	A Eustylini tribe entimine weevil.^[77] First named Otiorhynchus perditus (1876)^[67]
†Pseudophaops	†P. wickhami Legalov, 2018	Lake Uinta		A Tropiphorini tribe entimine weevil.^[77]
"Rhyssomatus"	†"R". tabescens Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain generic affiliations.^[78]^[76]
†Sciabregma	†S. rugosa Scudder, 1893	Lake Unita	1 specimen	A Sciabregmini tribe molytine weevil.^[78]
†Sciabregma	†S. tenuicornis Cockerell, 1921	Lake Unita		A Sciabregmini tribe molytine weevil.^[70]
†Scythropus	†S. somniculosus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain subfamilial affiliations.^[78]^[76]
"Sitona"	†"S". paginarum Scudder, 1893	Lake Gosiute Lake Unita	4 specimens	A Tropiphorini tribe entimine weevil of uncertain genus affiliations.^[78]^[76]
†Steganus	†S. barrandei Scudder, 1893	Lake Unita	1 specimen	A conoderine weevil.^[78]^[76]
†Syntomostylus	†S. rudis Scudder, 1893	Lake Unita	3 specimens	A weevil of uncertain subfamilial affiliations.^[78]^[76]
"Trypodendron"	†"T". impressus (Scudder, 1876)	Lake Gosiute	4 specimens	A scolytine bark beetle. Considered genus and tribus incertae sedis by Legalov (2015)^[76] Jr synonym includes Dryocoetes impressus (1878 in paleontology)^[66]
Erotylidae	Tritoma	†T. binotata Scudder, 1878	Lake Gosiute	1 specimen	A Tritomini tribe erotyline pleasing fungus beetle.^[88] First named as Mycotretus binotata (1878)^[66]
Laemophloeidae	Parandrita	†P. vestita Scudder, 1890	Lake Gosiute	4 specimens	A lined flat bark beetle.^[56]
Melandryidae	†Cicindelopsis	†C. eophilus Cockerell, 1921	Lake Unita		A false darkling beetle.^[68]
Mordellidae	†Petrimordella	†P. priscula (Cockerell, 1924)		1 specimen	A tumbling flower beetle morphospecies. First named as Mordella priscula (1925).^[89]
Mordellidae	†Petrimordella	†P. rasnitsyni (Odnosum & Perkovsky, 2016)		1 specimen	A tumbling flower beetle morphospecies.^[89] First named as Isotrilophus rasnitsyni (2016).^[90]
Nemonychidae	†Eocaenonemonyx	†E. kuscheli Legalov, 2013	Lake Unita		An Eocaenonemonychini tribe cretonemonychine pine flower weevil.^[91]
	†"Eugnamptus"	†"E". decemsatus Scudder, 1878	Lake Gosiute	1 specimen	An Eocaenonemonychini tribe cretonemonychine pine flower weevil.^[66] In need of redescription per Legalov (2013).^[91]
	†"Eugnamptus"	†"E". grandaevus (Scudder, 1876)	Lake Gosiute	3 specimens	An Eocaenonemonychini tribe cretonemonychine pine flower weevil.^[66] First named as Sitona grandaevus (1876) In need of redescription per Legalov (2013).^[91]
Peltidae	"Phenolia"	†"P." incapax Scudder, 1876	Lake Gosiute	1 specimen	A possible shield beetle in need of redescription.^[92] First placed as a Nitidulid sap beetle.^[67]

Family	Genus	Species	Member	Abundance	Notes	Image
Scarabaeidae	Aegialia	†A. rupta Scudder, 1890	Lake Gosiute	1 specimen	An aegialiine scaraboid beetle.^[93]
Scarabaeidae	Phyllophaga	†P. avus (Cockerell), Wickham, 1927	Lake Unita	1 specimen	A melolonthine scaraboid beetle. First described as Melolonthites avus (1921)^[72]^[93]