↑Hardy, Graham S(1985).“A new genus and species of pufferfish (Tetraodontidae) from Java”.Bulletin of marine science36(1): 145-149.
↑Yamanoue, Yusuke and Miya, Masaki and Doi, Hiroyuki and Mabuchi, Kohji and Sakai, Harumi and Nishida, Mutsumi(2011).“Multiple invasions into freshwater by pufferfishes (Teleostei: Tetraodontidae): a mitogenomic perspective”.PLoS One6(2): e17410.doi:10.1371/journal.pone.0017410.
