2019 in arthropod paleontology
From Wikipedia, the free encyclopedia
This list of fossil arthropods described in 2019 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that were described during the year 2019, as well as other significant discoveries and events related to arthropod paleontology that occurred in the year 2019.
General research
- A revision of higher-level arthropod taxonomy is published by Aria (2019), who proposes the name Cenocondyla for the least inclusive group containing both Mandibulata and Chelicerata.[1]
- A study on the molecular composition and microanatomy of the eyes of 54-million-year-old crane flies from the Fur Formation (Denmark), evaluating its implications for the knowledge of optical systems of other ancient arthropods (especially trilobites), is published by Lindgren et al. (2019).[2]
Arachnids
Research
- Two new specimens of laniatorid harvestmen (an adult specimen assignable to the species Proholoscotolemon nemastomoides and a smaller specimen belonging or related to the genus Proholoscotolemon) are described from the Eocene Baltic amber by Bartel & Dunlop (2019).[3]
- A review of the phylogenetic hypotheses of scorpion interrelationships, as well as of scorpion palaeontology, is published by Howard et al. (2019), who also provide a list of scorpion fossil calibrations for use in molecular dating.[4]
- A fossil spider, possibly a juvenile female golden silk orb-weaver, is described from the Lower Eocene Palana Formation of the Gurha opencast lignite mine (western Rajasthan, India) by Patel, Rana & Selden (2019), representing the first fossil spider from India which wasn't found in amber.[5]
- A review of the fossil record and evolutionary history of spiders, reevaluating the phylogenetic placement of key fossils and aiming to determine whether there has been a major turnover in the spider fauna between the Mesozoic and Cenozoic, is published online by Magalhaes et al. (2019).[6]
New taxa
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Caestaranea[7] |
Gen. et sp. nov |
Valid |
Selden, Huang & Garwood |
Middle–Late Jurassic |
A palpimanoid spider. Genus includes new species C. jurassica. |
|||
|
Chaerilobuthus hansgeorgmuelleri[8] |
Sp. nov |
Valid |
Lourenço in Lourenço & Velten |
Cretaceous |
Burmese amber |
A scorpion belonging to the family Chaerilobuthidae. |
||
|
Colossopantopodus nanus[9] |
Sp. nov |
Valid |
Sabroux et al. |
A sea spider. |
||||
|
Cosmobunus sagani[10] |
Sp. nov |
Valid |
Palencia et al. |
Rubielos de Mora Basin |
A member of Opiliones, a species of Cosmobunus. |
|||
|
Eatoniana crinita[11] |
Sp. nov |
Valid |
Sidorchuk et al. |
A mite belonging to the family Erythraeidae. |
||||
|
Eomesothele[12] |
Gen. et sp. nov |
Valid |
Wunderlich |
A spider belonging to group Mesothelae, assigned to the new family Eomesothelidae. The type species is E. noninclinata. |
||||
|
Eumesosoma abdelmawlai[13] |
Sp. nov |
Valid |
Elsaka, Mitov & Dunlop |
Europe (Baltic Sea region) |
A harvestman, a species of Eumesosoma. |
|||
|
?Eurycyde golem[9] |
Sp. nov |
Valid |
Sabroux et al. |
A sea spider belonging to the family Ascorhynchidae. |
||||
|
Intermesothele[12] |
Gen. et sp. nov |
Valid |
Wunderlich |
A spider belonging to group Mesothelae, possibly belonging to the family Eomesothelidae. The type species is I. pulcher. |
||||
|
Jinjumegops[14] |
Gen. et sp. nov |
Valid |
Park, Nam & Selden |
A spider belonging to the family Lagonomegopidae. The type species is J. dalingwateri. |
||||
|
Koreamegops[14] |
Gen. et sp. nov |
Valid |
Park, Nam & Selden |
A spider belonging to the family Lagonomegopidae. The type species is K. samsiki. |
||||
|
Lutetiana[15] |
Gen. et sp. nov |
Valid |
Selden & Wappler |
A spider, possibly a member of the family Cybaeidae. The type species is L. neli. |
||||
|
Metagonia esquincacanoi[16] |
Sp. nov |
García-Villafuerte |
A species of Metagonia. |
|||||
|
Neoliodes andreneli[17] |
Sp. nov |
Valid |
Arillo et al. |
Lebanese amber |
A mite belonging to the family Neoliodidae. Subsequently transferred to the separate genus Azarliodes.[18] |
|||
|
Onychopalpus[7] |
Gen. et sp. nov |
Valid |
Selden, Huang & Garwood |
Middle–Late Jurassic |
A palpimanoid spider. Genus includes new species O. thomisoides. |
|||
|
Paragilarovia[13] |
Gen. et sp. nov |
Valid |
Elsaka, Mitov & Dunlop |
Europe (Baltic Sea region) |
A harvestman belonging to the family Nemastomatidae. Genus includes new species P. hochae. |
|||
|
Procaeculus coineaui[19] |
Sp. nov |
Valid |
Porta et al. |
A mite belonging to the family Caeculidae. |
||||
|
Sp. nov |
Valid |
Elsaka, Mitov & Dunlop |
Europe (Baltic Sea region) |
A harvestman, a species of Protolophus. |
||||
|
Scolotydaeus vlaskini[20] |
Sp. nov |
Valid |
Klimov et al. |
Late Eocene |
A mite belonging to the family Paratydeidae. |
|||
|
Sinaranea brevicrus[7] |
Sp. nov |
Valid |
Selden, Huang & Garwood |
Middle–Late Jurassic |
||||
|
Tanytydeus pogrebnyaki[20] |
Sp. nov |
Valid |
Klimov et al. |
Late Eocene |
A mite belonging to the family Paratydeidae. |
|||
Crustaceans
Research
- A global dataset of exceptionally preserved muscles and muscle attachment scars in fossil malacostracan specimens is compiled by Klompmaker et al. (2019).[21]
- A three-dimensionally preserved tanaidacean specimen belonging or related to the species Jurapseudes friedericianus is described from the Middle Jurassic (Callovian) Ornatenton Formation (Germany) by Schädel et al. (2019).[22]
- A study on the evolutionary history of decapod crustaceans is published by Wolfe et al. (2019).[23]
- A study on the chemical composition of two specimens of fossilized shrimps from the Cretaceous Romualdo and Ipubi formations (Araripe Basin, Brazil), and on its implications for inferring the course of the processes of fossilization of these crustaceans, is published by Barros et al. (2019).[24]
- A study on the Middle Jurassic palaeoenvironment of La Voulte (France), as indicated by data from exceptionally preserved eyes of the polychelidan lobster Voulteryon parvulus and from epibiontic brachiopods associated with V. parvulus, is published by Audo et al. (2019).[25]
- A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.[26]
- An achelatan larval fossil with an intermediate type of morphology, interpreted as the oldest occurrence of an achelatan lobster larva reported so far, is described from the Toarcian Posidonia Shale (Germany) by Haug, Haug & Schweigert (2019).[27]
- Redescription of the isopod species Palaega sismondai Ristori (1891) is published by Hyžný, Pasini & Garassino (2019), who transfer this species to the genus Bathynomus.[28]
- A moult of a giant isopod is described from the early Oligocene Ranzano Formation by Pasini & Garassino (2019), representing the oldest fossil record of the genus Bathynomus from Italy.[29]
- A study on the Late Devonian ostracods from the Yangdi and Nandong sections in South China, and on their responses to the Frasnian-Famennian event, is published by Song, Huang & Gong (2019).[30]
- A diverse earliest Triassic ostracod fauna is reported from the Yangou section in South China by Qiu et al. (2019).[31]
- A revision of ostracods from the Upper Oligocene to Lower Miocene Pirabas Formation (Brazil) is published by Nogueira, Ramos & Hunt (2019).[32]
- A study on changes of diversity of Neogene fossil marine ostracods from Java, Indonesia is published by Shin et al. (2019).[33]
- A study on deep-sea benthic ostracod assemblages from the southern Sea of Japan covering the last 2 million years, evaluating their responses to the Mid-Brunhes Event, orbital-scale climatic cycles, and fluctuations of the Tsushima Warm Current, is published by Huang et al. (2019).[34]
- A three-dimensionally preserved specimen of a late growth stage of the phosphatocopid species Cyclotron angelini is described from the Furongian strata of northern Poland by Olempska et al. (2019).[35]
- A study on the ecology of clam shrimps from the Anjiagou and Hengdaozi beds of the Lower Cretaceous Yixian Formation (China) is published by Hethke et al. (2019).[36]
- A study on the oxygen isotope composition of whale barnacle shells from three Pleistocene localities along the eastern Pacific coast, and on their implications for the knowledge of the history of whale migrations, is published by Taylor et al. (2019).[37]
- A study on the internal soft-tissue anatomy of Hesslandona angustata is published by Liu et al. (2019).[38]
New taxa
Malacostracans
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Acanthocarpus osborni[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A crab, a species of Acanthocarpus. |
||||
|
Ahazianassa[40] |
Gen. et comb. nov |
Valid |
Karasawa et al. |
Izumi Group |
A member of the family Gourretiidae; a new genus for "Callianassa" masanorii Karasawa (1998). |
|||
|
Alontecarcinus[41] |
Gen. et sp. nov |
Valid |
De Angeli & Caporiondo |
A crab belonging to the family Potamidae. The type species is A. buratoi. |
||||
|
Amatukamius[42] |
Gen. et comb. nov |
Valid |
Karasawa & Ohara |
Arida Formation |
A member of Axiidea belonging to the family Micheleidae; a new genus for "Callianassa" sakakuraorum Karasawa (2000). |
|||
| Anapagurus dalessandroi[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) |  Italy |
A member of the family Paguridae. | ||
|
Annuntidiogenes elongatus[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Annuntidiogenidae. |
|||
|
Annuntidiogenes hoelderi[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Annuntidiogenidae. |
|||
|
Anoplocarcinus[45] |
Gen. et sp. nov |
Valid |
Schweitzer et al. |
A crab belonging to the group Raninoida and to the family Orithopsidae. The type species is A. hudspethi. |
||||
|
Aragolambrus[46] |
Gen. et sp. nov |
Valid |
Ferratges, Zamora & Aurell |
A crab belonging to the family Parthenopidae. Genus includes new species A. collinsi. |
||||
|
Archaeochiapasa[47] |
Gen. et sp. nov |
Valid |
Guinot, Carbot-Chanona & Vega |
A crab belonging to the group Heterotremata, assigned to the new family Archaeochiapasidae. The type species is A. mardoqueoi. |
||||
|
Archaeopus burgessi[48] |
Sp. nov |
Valid |
Nyborg et al. |
A crab belonging to the family Retroplumidae. |
||||
|
Archaeopus diabolus[49] |
Sp. nov |
Valid |
Schweitzer et al. |
|||||
|
Archaeopus hoploserratus[48] |
Sp. nov |
Valid |
Nyborg et al. |
A crab belonging to the family Retroplumidae. |
||||
|
Archaeopus morenoensis[48] |
Sp. nov |
Valid |
Nyborg et al. |
A crab belonging to the family Retroplumidae. |
||||
|
Bachmayerus gasparici[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. |
|||
|
Bachmayerus matushyznyi[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. |
|||
|
Bahiacaris[50] |
Gen. et comb. nov |
Valid |
Schweitzer et al. |
A caridean shrimp; a new genus for "Atyoida" roxoi Beurlen (1950). Announced in 2013; validated in 2023.[51] |
||||
|
Bathycalliax otsuka[52] |
Sp. nov |
Valid |
Ando, Kawano & Ugai |
Sakasegawa Formation |
A member of Axiidea belonging to the family Bathycalliacidae. |
|||
|
Bolcacalliax[53] |
Gen. et comb. nov |
Valid |
Pasini et al. |
A member of the family Callianassidae; a new genus for "Protaxius" eocenicus Secrétan (1975). |
||||
| Brachynotus multituberculatus[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A species of Brachynotus. | ||
|
Calappilia gableorum[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A species of Calappilia. |
||||
|
Calappilia granulosa[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A species of Calappilia. |
||||
|
Calappilia guttata[54] |
Sp. nov |
Valid |
Pasini, Garassino & Damarco |
A species of Calappilia. |
||||
|
Calappilia perreaulti[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A species of Calappilia. |
||||
|
Gen. et sp. nov |
Valid |
Luque et al. |
Churuvita Group |
A crab belonging to the new family Callichimaeridae. The type species is C. perplexa. |
 | |||
|
Caloxanthus kavanaghi[56] |
Sp. nov |
Valid |
Collins, Garvie & Mellish |
A crab belonging to the group Etyoidea and to the family Feldmannidae. |
||||
|
Cenocorystes libbyae[57] |
Sp. nov |
Valid |
Van Bakel et al. |
A palaeocorystoid crab. |
||||
| Corallomursia prominens[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Calappidae. | ||
|
Costadromia[58] |
Gen. et sp. nov |
Valid |
Feldmann & Schweitzer |
Late Cretaceous (late Campanian) |
A sponge crab. The type species is C. hajzeri. |
|||
|
Cretainermis[59] |
Gen. et sp. nov |
Valid |
Prado, Calado & Barreto |
A member of Penaeoidea. Genus includes new species C. pernambucensis. |
||||
|
Cretamaja klompmakeri[60] |
Sp. nov |
Valid |
González-León & Moreno-Bedmar in Vega, González-León & Moreno-Bedmar |
A crab. |
||||
|
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Pylochelidae. |
||||
|
Cymodoce (?) scopula[61] |
Sp. nov |
Valid |
Fraaije et al. |
Basque-Cantabrian Basin |
||||
| Daragrapsus setalis[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of Grapsoidea of uncertain phylogenetic placement. | ||
|
Dardanus caporiondoi[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A species of Dardanus. |
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|
Devonoaxius[63] |
Gen. et sp. nov |
Valid |
Feldmann & Schweitzer |
A member of Axiidea. Genus includes new species D. garlandi. |
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|
Dinocarcinus[64] |
Gen. et sp. nov |
Valid |
Van Bakel, Hyžný, Valentin & Robin in Robin et al. |
Late Cretaceous (late Campanian) |
A crab, a member of Portunoidea sensu lato. The type species is D. velauciensis. Announced in 2019; validated in 2023.[65] |
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|
Doraranina[66] |
Gen. et sp. nov |
Valid |
Gustafson, Nyborg & van Bakel |
Early Eocene |
A crab belonging to the group Raninoidea and to the family Lyreididae. Genus includes new species D. manleyi. |
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| Dromiopsis marginospinosa[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Dromiidae. | ||
|
Dromiopsis oscari[67] |
Sp. nov |
Valid |
Nyborg et al. |
|||||
| ?Dromiopsis rocchettii[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Dromiidae. | ||
|
Ebalia pumicosa[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A species of Ebalia. |
|||
|
Eobodotria[68] |
Gen. et sp. nov |
Valid |
Luque & Gerken |
A member of Cumacea. Genus includes new species E. muisca. |
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|
Eocarpilius vicetinus[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A crab belonging to the group Carpilioidea and the family Carpiliidae. |
|||
|
Eodorippe connori[67] |
Sp. nov |
Valid |
Nyborg et al. |
A crab belonging to the superfamily Dorippoidea and the family Telamonocarcinidae. Originally described as a species of Eodorippe, but subsequently transferred to the genus Personadorippe.[69] |
||||
|
Eopaguropsis grandis[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. |
|||
|
Eopaguropsis schindewolfi[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. |
|||
|
Eucalliax capsulasetaea[70] |
Sp. nov |
Valid |
Kiel et al. |
Early Oligocene |
Talara Basin |
A member of the family Callianassidae. |
||
| Eumunida veronensis[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Munididae. Originally described as a species of Eumunida, but subsequently transferred to the genus Muellermunida.[71] | ||
| Folguerolesia collinsi[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of Leucosioidea belonging to the family Folguerolesiidae. | ||
|
?Galathea genesis[72] |
Sp. nov |
Valid |
Robins & Klompmaker |
Ernstbrunn Limestone |
A member of the family Galatheidae. |
|||
|
Galatheites britmelanarum[72] |
Sp. nov |
Valid |
Robins & Klompmaker |
Ernstbrunn Limestone |
A member of Galatheoidea belonging to the family Catillogalatheidae. |
|||
|
Gastrosacus raboeufi[73] |
Sp. nov |
Valid |
Fraaije et al. |
A member of Galatheoidea. |
||||
|
Glaessnerella yamoori[67] |
Sp. nov |
Valid |
Nyborg et al. |
|||||
| Glabropilumnus bizzarinii[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Pilumnidae. | ||
|
Glabropilumnus tuberculatus[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A crab belonging to the group Pilumnoidea and the family Pilumnidae. |
|||
|
Gyroplanus[49] |
Gen. et sp. nov |
Valid |
Schweitzer et al. |
A cyclodorippoid crab. Genus includes new species G. sapphirinus. |
||||
| Hepatiscus meneghinii[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Aethridae. | ||
|
Hinecaris[40] |
Gen. et sp. nov |
Valid |
Karasawa et al. |
Izumi Group |
A member of the family Axiidae. The type species is H. simplex. |
|||
|
Kromtitis lluisprietoi[74] |
Sp. nov |
Valid |
Ossó |
A crab belonging to the family Dynomenidae. |
||||
|
Lindahomola[75] |
Gen. et sp. nov |
Valid |
Garassino et al. |
A crab belonging to the family Homolidae. Genus includes new species L. longispina. |
||||
|
Linuparus shacklefordae[49] |
Sp. nov |
Valid |
Schweitzer et al. |
A palinurid lobster. |
||||
|
Lophoranina lincki[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A crab. |
||||
|
Macroacaena hannya[52] |
Sp. nov |
Valid |
Ando, Kawano & Ugai |
Sakasegawa Formation |
A crab belonging to the family Lyreididae. |
|||
|
Maja prealpina[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A species of Maja. |
|||
|
Matutites periosus[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
A crab. |
||||
|
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Parapylochelidae. |
||||
|
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Parapylochelidae. |
||||
|
Mesostylus pervesleri[76] |
Sp. nov |
Valid |
Hyžný & Summesberger |
Piesting Formation |
A member of the family Callianassidae. |
|||
|
Montemagambrus[77] |
Gen. et sp. nov |
Valid |
Ceccon & De Angeli |
Early Eocene |
A crab belonging to the family Parthenopidae. Genus includes new species M. corallinus. |
|||
|
Munidopsis salinaria[78] |
Sp. nov |
Valid |
Gašparič et al. |
Tuzla Basin |
A species of Munidopsis. |
|||
|
Munitheites kuepperi[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Schobertellidae. |
|||
| Nachioides[43] | Gen. et sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Inachoididae. The type species is N. tuberculatus. | ||
|
Natatolana poblana[79] |
Sp. nov |
Valid |
Vega & Bruce in Vega et al. |
An isopod belonging to the family Cirolanidae. |
||||
| Nicolisia[43] | Gen. et sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of Xanthoidea belonging to the family Eumedonidae. The type species is N. antiqua. | ||
|
Notopus adinae[39] |
Sp. nov |
Valid |
Feldmann, Schweitzer & Phillips |
|||||
|
Ovalopus[72] |
Gen. et comb. nov |
Valid |
Robins & Klompmaker |
Ernstbrunn Limestone |
A crab belonging to the group Homolodromioidea and to the family Jurellanidae. The type species is O. hoheneggeri (Moericke, 1889); genus also includes Coelopus convexus Schweitzer & Feldmann (2010) and Coelopus repandus Schweitzer & Feldmann (2010). |
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| Pachycheles bazzanii[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Porcellanidae. | ||
|
Pagurus haasei[80] |
Sp. nov |
Valid |
Polkowsky & Fraaije |
A hermit crab, a species of Pagurus. |
||||
|
Palaemon aestuarius[81] |
Sp. nov |
Du et al. |
Early Miocene |
A species of Palaemon. |
||||
|
Palaeopalinurus serratus[61] |
Sp. nov |
Valid |
Fraaije et al. |
Basque-Cantabrian Basin |
||||
|
Palinurus? sauerweinorum[49] |
Sp. nov |
Valid |
Schweitzer et al. |
A palinurid lobster. |
||||
|
Panopeus santurbanensis[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A species of Panopeus. |
|||
| Paronachoides[43] | Gen. et sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Inachoididae. The type species is P. scaber. | ||
|
Percnon santurbanensis[82] |
Sp. nov |
Valid |
Ceccon & de Angeli |
|||||
| Petrolisthes punctatus[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A species of Petrolisthes. | ||
|
Pilumnopeus priscus[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A crab belonging to the group Pilumnoidea and the family Pilumnidae. |
|||
|
Pilumnus pedemontanus[62] |
Sp. nov |
Valid |
Ceccon & De Angeli |
Early Oligocene |
A species of Pilumnus. |
|||
|
Planoprosopon ogawaense[83] |
Sp. nov |
Valid |
Karasawa & Hirota |
Yatsuji Formation |
A crab belonging to the group Homolodromioidea and to the family Longodromitidae. |
|||
|
Planoprosopon sarumaru[83] |
Sp. nov |
Valid |
Karasawa & Hirota |
Yatsuji Formation |
A crab belonging to the group Homolodromioidea and to the family Longodromitidae. |
|||
|
Pretrizocheles[44] |
Gen. et sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Pylochelidae. The type species is P. cocullo. |
|||
|
Prolecythocaris rieberi[84] |
Sp. nov |
Valid |
Schweigert |
A crab belonging to the group Dromiacea and to the family Lecythocaridae. |
||||
|
Protopagurus[44] |
Gen. et sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Paguridae. The type species is P. janoscheki. |
|||
| Pseudocarinocarcinoides[43] | Gen. et sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of Goneplacoidea belonging to the family Carinocarcinoididae. The type species is P. karasawai. | ||
|
Retropluma minuta[78] |
Sp. nov |
Valid |
Gašparič et al. |
Tuzla Basin |
A species of Retropluma. |
|||
|
Ripleycarcinus[49] |
Gen. et sp. nov |
Valid |
Schweitzer et al. |
A homoloid crab. Genus includes new species R. primaevus. |
||||
|
Ripleycorystes[49] |
Gen. et sp. nov |
Valid |
Schweitzer et al. |
A palaeocorystoid crab. Genus includes new species R. cardwelli. |
||||
|
Symethoides danieli[49] |
Sp. nov |
Valid |
Schweitzer et al. |
A raninoid crab. |
||||
| Tethysmunida setifera[43] | Sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Munidopsidae. | ||
|
Tithopaguristes[44] |
Gen. et sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. The type species is T. porosus. |
|||
|
Tropidicarcinus[49] |
Gen. et 2 sp. nov |
Valid |
Schweitzer et al. |
A dakoticancroid crab. Genus includes new species T. mcanallyae and T. starnesi. |
||||
|
Ululapagurus oroszyi[44] |
Sp. nov |
Valid |
Fraaije et al. |
Ernstbrunn Limestone |
A hermit crab belonging to the family Diogenidae. |
|||
|
Vacuotheca[85] |
Gen. et sp. nov |
Valid |
Schädel, Perrichot & Haug |
A member of Isopoda belonging to the group Epicaridea. The type species is V. dupeorum. |
||||
|
Valamunida[86] |
Gen. et sp. nov |
Valid |
Klompmaker & Robins in Hryniewicz et al. |
Late Paleocene |
Basilika Formation |
A member of the family Munididae. The type species is V. haeggi. |
||
| Verarena[43] | Gen. et sp. nov | Valid | Beschin, Busulini & Tessier in Beschin et al. | Eocene (Priabonian) | Italy |
A member of the family Euryplacidae. The type species is V. katoi. | ||
|
Vibrissalana[72] |
Gen. et sp. nov |
Valid |
Robins & Klompmaker |
Ernstbrunn Limestone |
A porcelain crab. Genus includes new species V. jurassica. |
|||
|
Vicluvia[87] |
Gen. et sp. nov |
Valid |
Larghi et al. |
Meride Limestone |
A member of Lophogastrida. Genus includes new species V. lombardoae. |
|||
Ostracods
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Amicytheridea bilthanaensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Bairdiocypris fabiformis[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Bairdoppilata dorsoarcuata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Bairdiidae. |
|||
|
Bairdoppilata pintoi[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
La Rosa Formation |
A member of Podocopida belonging to the family Bairdiidae. |
|||
|
Bairdoppilata turonica[90] |
Sp. nov |
Valid |
Slipper |
|||||
|
Bairdoppilata vandenboldi[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Bairdiidae. |
|||
|
Bakunella anae[91] |
Sp. nov |
Valid |
Spadi et al. |
A member of the family Candonidae. |
||||
|
Bassleratia torquata[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Bolbozoe psittaca[92] |
Sp. nov |
Valid |
Perrier et al. |
A member of Myodocopa. |
||||
|
Boreobolbina morkokiana[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Bradleya majorani[93] |
Sp. nov |
Valid |
Bergue, Brandão & Anjos Zerfass |
A member of the family Thaerocytheridae. |
||||
|
Bradleya ybate[93] |
Sp. nov |
Valid |
Bergue, Brandão & Anjos Zerfass |
Late Miocene |
A member of the family Thaerocytheridae. |
|||
|
Bythocypris wangi[94] |
Sp. nov |
Valid |
Tanaka, Siveter & Williams |
Fukuji Formation |
||||
|
Bythoceratina? asulcata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Bythocytheridae. |
||
|
Bythoceratina? sinuocostata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Bythocytheridae. |
|||
|
Cardobairdia longitecta[90] |
Sp. nov |
Valid |
Slipper |
|||||
|
Carinobairdia cabralae[95] |
Sp. nov |
Forel in Forel, Thuy & Wisshak |
Maantang Formation |
A member of the family Bairdiidae. |
||||
|
Cativella primaveraensis[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Trachyleberididae. |
||
|
Clavofabellina fukujiensis[94] |
Sp. nov |
Valid |
Tanaka, Siveter & Williams |
Fukuji Formation |
||||
|
Clintiella antifrigga[96] |
Sp. nov |
Valid |
Siveter et al. |
Hitoegane Formation |
A beyrichioid palaeocopid ostracod. |
|||
|
Coelochilina reticulata[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Costa spinaventralis[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Trachyleberididae. |
||
|
Curfsina coarctata[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Curfsina hanumanpuraensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Cushmanidea? bragantinaensis[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Cushmanideidae. |
|||
|
Cyprideis? carmoi[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Cytherideidae. |
|||
|
Cytherella altacaelateralis[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of the family Cytherellidae. |
|||
|
Cytherella kempfi[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of the family Cytherellidae. |
|||
|
Cytherella notossinuosa[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of the family Cytherellidae. |
|||
|
Cytherella truncatoides[90] |
Sp. nov |
Valid |
Slipper |
|||||
|
Cytherella vulna[90] |
Sp. nov |
Valid |
Slipper |
|||||
|
Cytherella weaveri[90] |
Sp. nov |
Valid |
Slipper |
|||||
|
Cytherelloidea colini[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of the family Cytherellidae. |
||
|
Cytherelloidea cumpassorobusta[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of the family Cytherellidae. |
||
|
Cytherelloidea granulosa parca[90] |
Subsp. nov |
Valid |
Slipper |
|||||
|
Cytherelloidea mediocythara[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of the family Cytherellidae. |
||
|
Cytherelloidea quasilenisa[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of the family Cytherellidae. |
||
|
Cytheretta fortiscostata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytherettidae. |
||
|
Cytheretta petrii[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytherettidae. |
||
|
Cytheropteron hanumanpuraensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Cytheropteron ratitalaiensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Cytherura? ornatareticulum[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytheruridae. |
||
|
Cytherura? punctocentrata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytheruridae. |
||
|
Cytherura? quasilenisa[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Cytheruridae. |
|||
|
Eocytheropteron bilthanaensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Gibbosocythere[97] |
Gen. et 2 sp. nov |
Valid |
Karpuk, Whatley & Maybury |
Crimean Peninsula |
A member of Paradoxostomatidae. Genus includes new species G. ferelevis and G. cellulata. |
|||
|
Hemicytherura bradyisimilis[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytheruridae. |
||
|
Hiatobairdia senegasi[95] |
Sp. nov |
Forel in Forel, Thuy & Wisshak |
Late Triassic (Carnian and Norian) |
Kuahongdong Formation |
A member of the family Bairdiidae. |
|||
|
Hiatobairdia zhengshuyingi[95] |
Sp. nov |
Forel in Forel, Thuy & Wisshak |
Late Triassic (Carnian and Norian) |
Kuahongdong Formation |
A member of the family Bairdiidae. |
|||
|
Hollinella orienta[96] |
Sp. nov |
Valid |
Siveter et al. |
Gionyama Formation |
A hollinoid palaeocopid ostracod. |
|||
|
Hulingsina? lorenesmithae[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cushmanideidae. |
||
|
Hulingsina reticulorugosa[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Cushmanideidae. |
|||
|
Hungarella gommerii[95] |
Sp. nov |
Forel in Forel, Thuy & Wisshak |
Late Triassic (Carnian) |
Maantang Formation |
A member of the family Healdiidae. |
|||
|
Jatella ampla[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Krithe araucoensis[98] |
Sp. nov |
Valid |
Bergue, Coimbra & Finger |
Early Miocene |
||||
|
Krithe dawnpetersonae[98] |
Sp. nov |
Valid |
Bergue, Coimbra & Finger |
Early Miocene |
||||
|
Krithe nerudai[98] |
Sp. nov |
Valid |
Bergue, Coimbra & Finger |
Early Miocene |
||||
|
Leperditella paula[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Loxoconcha corpulenta[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Loxoconchidae. |
||
|
Microcythere acuminata[93] |
Sp. nov |
Valid |
Bergue, Brandão & Anjos Zerfass |
A member of the family Microcytheridae. |
||||
|
Moierina grata[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Mydionobairdia cauladelicata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Bairdiidae. |
||
|
Nigeroloxoconcha baghensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Nigeroloxoconcha diluta[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Nudator[92] |
Gen. et 4 sp. nov |
Valid |
Perrier et al. |
A member of Myodocopa belonging to the family Entomozoidae. The type species is N. inflatus; genus also includes new species N. angiportatus, N. artumatus and N. elegantulus.[99] |
||||
|
Ochesaarina aculeata[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Oculparva[92] |
Gen. et sp. nov |
Valid |
Perrier et al. |
A member of Myodocopa. Genus includes new species O. moncola. |
||||
|
Ovocytheridea baghensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Paijenborchella jeerabadensis[88] |
Sp. nov |
Valid |
Chaudhary & Nagori |
Bagh Formation |
||||
|
Pauproles[96] |
Gen. et sp. nov |
Valid |
Siveter et al. |
Hitoegane Formation |
A eurychilinoid palaeocopid ostracod. Genus includes new species P. supparata. |
|||
|
Piscarista[92] |
Gen. et sp. nov |
Valid |
Perrier et al. |
A member of Myodocopa. Genus includes new species P. sagenata. |
||||
|
Planiglandites[89] |
Gen. et sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
Genus includes new species P. mirabilis. |
||
|
Pontocyprella goussardi[95] |
Sp. nov |
Forel in Forel, Thuy & Wisshak |
Late Triassic (Carnian) |
Maantang Formation |
A member of the family Pontocyprididae. |
|||
|
Pontocyprella robusta cometa[90] |
Subsp. nov |
Valid |
Slipper |
|||||
|
Pontocypris aguilerai[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Pirabas Formation |
A member of Podocopida belonging to the family Pontocypridae. |
|||
|
Pseudotethysia[97] |
Gen. et sp. nov |
Valid |
Karpuk, Whatley & Maybury |
Crimean Peninsula |
A member of Cytheruridae. Genus includes new species P. reticulata. |
|||
|
Quadracythere adornata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Hemicytheridae. |
||
|
Quadracythere fortisrobusta[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Hemicytheridae. |
||
|
Quadracythere limbimodesta[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Hemicytheridae. |
||
|
Rectella cistelliformis[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Retiprimites alveolatus[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Retiprimites mirandus[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
|
Rostrocytheridea decurtata[100] |
Sp. nov |
Valid |
Chaudhary, Nagori & Khosla |
Bagh Formation |
A member of the family Cytherideidae. |
|||
|
Rostrocytheridea divergens[100] |
Sp. nov |
Valid |
Chaudhary, Nagori & Khosla |
Bagh Formation |
A member of the family Cytherideidae. |
|||
|
Semicytherura? fortisreticulata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Cytheruridae. |
||
|
Seminova[92] |
Gen. et comb. nov |
Valid |
Perrier et al. |
A member of Myodocopa belonging to the superfamily Bolbozoidea. The type species is "Entomis" depressa Jones (1884). |
||||
|
Tenedocythere? rugosocostata[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Hemicytheridae. |
||
|
Touroconcha ishizakii[32] |
Sp. nov |
Valid |
Nogueira, Ramos & Hunt |
Early Miocene |
Pirabas Formation |
A member of Podocopida belonging to the family Loxoconchidae. |
||
|
Uthoernia sibirica[89] |
Sp. nov |
Valid |
Melnikova |
Late Ordovician |
Delingde Formation |
|||
Other crustaceans
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Valid |
Collareta et al. |
Early Miocene |
A barnacle. |
||||
|
Calmanostraca hassbergella[102] |
Sp. nov |
Wagner, Hauh & Haug |
A member of Branchiopoda belonging to the group Calmanostraca. |
|||||
|
Cretiscalpellum matrioni[103] |
Sp. nov |
Valid |
Gale |
A barnacle. Originally described as a species of Cretiscalpellum; Gale (2020) transferred it to the genus Witherscalpellum.[104] |
||||
|
Cretiscalpellum mutterlosei[103] |
Sp. nov |
Valid |
Gale |
A barnacle. |
||||
|
Halicyne oosterinkorum[105] |
Sp. nov |
Valid |
Schweitzer, Feldmann & Schinker |
A member of Cyclida. |
||||
|
Jaegerscalpellum[103] |
Gen. et sp. et comb. nov |
Valid |
Gale |
A barnacle. Genus includes new species J. elegans, as well as J. comptum (Withers, 1910) and J. politum (Darwin, 1851). |
||||
|
Liapora[106] |
Gen. et sp. nov |
Valid |
Keupp |
Originally described as a scyphozoan polyp; genus includes new species L. neubigi. Gale, Keupp & Schweigert (2022) reinterpreted it as capitular valves of a phosphatic barnacle that had been described as Pollicipes(?) lotharingica by Méchin in 1901.[107] |
||||
|
Martillepas auriculum[103] |
Sp. nov |
Valid |
Gale |
A barnacle. |
||||
|
Martillepas decoratus[103] |
Sp. nov |
Valid |
Gale |
A barnacle. |
||||
|
Notostraca oleseni[102] |
Sp. nov |
Wagner, Hauh & Haug |
A member of Branchiopoda belonging to the group Notostraca. |
|||||
|
Paraleptestheria mitchelli[108] |
Sp. nov |
Valid |
Gallego et al. |
Late Eocene |
A clam shrimp. |
|||
|
Pedupycnolepas pulcher[103] |
Sp. nov |
Valid |
Gale |
A barnacle. |
||||
|
Pemphilimnadiopsis cheni[109] |
Sp. nov |
Valid |
Liao et al. |
Benxi Formation |
A clam shrimp. |
|||
|
Perumegabalanus[110] |
Gen. et sp. nov |
Valid |
Coletti et al. |
Chilcatay Formation |
A barnacle. Genus includes new species P. calziai. |
|||
|
Platylepas mediterranea[111] |
Sp. nov |
Valid |
Collareta et al. |
Early Pleistocene (Gelasian) |
A barnacle, a species of Platylepas. |
|||
|
Prolatcyclus[112] |
Gen. et comb. et sp. nov |
Valid |
Mychko et al. |
A member of Cyclida. Genus includes P. martinensis (Goldring, 1967) and a new species P. kindzadza. |
||||
|
Verruca alabamensis[113] |
Sp. nov |
Valid |
Perreault & Buckeridge |
A barnacle belonging to the family Verrucidae. |
||||
|
Verruca gailgoedertae[113] |
Sp. nov |
Valid |
Perreault & Buckeridge |
Middle Eocene |
A barnacle belonging to the family Verrucidae. |
|||
|
Verruca sorrellae[113] |
Sp. nov |
Valid |
Perreault & Buckeridge |
A barnacle belonging to the family Verrucidae. |
||||
Insects
Trilobites
Research
- A study on the rates of evolution of trilobites in the Cambrian is published by Paterson, Edgecombe & Lee (2019), who interpret their findings as indicative of Cambrian rather than Precambrian origin of trilobites, and as indicating that the Cambrian explosion had already largely concluded by the time the typical Cambrian fossil record begins (≈521 Ma).[114]
- A study on repaired injuries in trilobite specimens from the Cambrian Ruin Wash Lagerstätte (Nevada, United States) is published by Pates & Bicknell (2019), who interpret their findings as indicative of species specific predation on Cambrian trilobites, and of enlarged spines in some trilobite species serving as possible predation deterrents.[115]
- A study on the morphology and ontogeny of Changaspis elongata, based on data from specimens from the Lazizhai section of the Balang Formation (Cambrian Stage 4; Guizhou, China), is published by Du et al. (2019).[116]
- A cluster of over 100 juvenile trilobites belonging to the genus Aphelaspis is reported from the Cambrian (Paibian) Conasauga Formation (Georgia, United States) by Schwimmer & Montante (2019).[117]
- A study on patterns of occupancy and diversity trajectories in late Cambrian-early Ordovician trilobite communities from the Argentine Cordillera Oriental is published by Serra, Balseiro & Waisfeld (2019).[118]
- A study on the phylogenetic relationships of members of the family Olenidae is published by Monti & Confalonieri (2019).[119]
- A study on the phylogenetic relationships of members of the olenid subfamily Balnibarbiinae is published by Hopkins (2019).[120]
- Nine specimens of Symphysurus ebbestadi preserving all key stages of the exoskeleton moulting process are described from the Lower Ordovician Fezouata Formation (Morocco) by Drage et al. (2019).[121]
- A study on clusters of Ampyx priscus from the Ordovician Fezouata Shale of Morocco is published by Vannier et al. (2019), who interpret these clusters as evidence of a collective behaviour rather than a result of passive transportation and accumulation.[122]
- A study on the phylogenetic relationships of the species and subspecies assigned to the phacopid genus Austerops is published by Oudot et al. (2019).[123]
- A study on the morphological changes in the ontogeny of the Ordovician phacopid trilobite Calyptaulax strasburgensis is published by Jacobs & Carlucci (2019).[124]
- A well-preserved body cluster of 18 specimens of Arctinurus boltoni is described from the Silurian Rochester Shale (New York, United States) by Bicknell, Paterson & Hopkins (2019).[125]
- A study on development and evolution of the cephalon and the pygidium in phacopid trilobites is published by Oudot et al. (2019).[126]
- A study on the internal structures of eyes of trilobites belonging to the genera Asaphus and Archegonus is published by Scholtz, Staude & Dunlop (2019);[127] the study is subsequently criticized by Schoenemann & Clarkson (2021).[128][129]
- A study on the variability of trilobite moulting behaviour is published by Drage (2019).[130]
New taxa
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Anacaenaspis yanpingensis[131] |
Sp. nov |
Valid |
Wei & Zhan |
Niuchang Formation |
||||
|
Sp. nov |
Valid |
Ghobadi Pour |
||||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Astycoryphe rugocauda[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Couvin Formation |
A member of Proetida. |
|||
|
Astycoryphe tureki[134] |
Sp. nov |
Valid |
Mergl & Budil |
A member of Proetida. |
||||
|
Bigotina kelleri[135] |
Sp. nov |
Valid |
Geyer |
Igoudine Formation |
A member of Redlichiina belonging to the family Bigotinidae. |
|||
|
Bigotina monningeri[135] |
Sp. nov |
Valid |
Geyer |
A member of Redlichiina belonging to the family Bigotinidae. |
||||
|
Bigotinops chouberti[135] |
Sp. nov |
Valid |
Geyer |
A member of Redlichiina belonging to the family Bigotinidae. |
||||
|
Brachymetopus (Conimetopus) alekseevi[136] |
Sp. nov |
Valid |
Mychko in Mychko & Savchuk |
Early Permian |
||||
|
Brevibole minerva[137] |
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Achegoninae. |
||||
|
Bronteopsis tenuirhachis[138] |
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Povorotnaya Formation |
A member of the family Styginidae. |
||
|
Sp. nov |
Valid |
Ameri, Arjmandzadeh & Ghoorcgi |
Early Silurian |
Niur Formation |
Announced in 2019; the final version of the article naming was published in 2021. |
|||
|
Carbonoproetus bellona[137] |
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Bollandiinae. |
||||
|
Carbonoproetus volturnus[137] |
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Bollandiinae. |
||||
|
Conophrys multituberculatus[132] |
Sp. nov |
Valid |
Ghobadi Pour |
|||||
|
Cyphaspides ammari[140] |
Sp. nov |
Valid |
Chatterton, Gibb & McKellar |
Bou Tchrafine Group |
A member of Proetida belonging to the family Aulacopleuridae. |
|||
|
Cyphaspides nicoleae[140] |
Sp. nov |
Valid |
Chatterton, Gibb & McKellar |
Bou Tchrafine Group |
A member of Proetida belonging to the family Aulacopleuridae. |
|||
|
Cyphaspides pankowskiorum[140] |
Sp. nov |
Valid |
Chatterton, Gibb & McKellar |
Bou Tchrafine Group |
A member of Proetida belonging to the family Aulacopleuridae. |
|||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Alberti |
A member of the family Aulacopleuridae. |
|||||
|
Cyrtoproetus korni[137] |
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Achegoninae. |
||||
|
Cyrtoproetus roesenbeckensis[137] |
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Achegoninae. |
||||
|
Cyrtosymbolinella[142] |
Gen. et sp. nov |
Valid |
Basse & Lemke |
A member of the family Phillipsiidae belonging to the subfamily Cyrtosymbolinae. The type species is C. svojtkai. |
||||
|
Debrenella[135] |
Gen. et sp. nov |
Valid |
Geyer |
A member of Redlichiida belonging to the family Fallotaspididae. Genus includes new species D. larvalis. |
||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of Proetida. |
||||
|
Dionide trigintasegmentata[138] |
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Barkov Formation |
A member of Asaphida belonging to the family Dionididae. |
||
|
Eladiolinania[135] |
Gen. et 2 sp. et comb. nov |
Valid |
Geyer |
Igoudine Formation |
A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species E. castor and E. pollux, and possibly also "Serrania" palaciosi Liñán et al. (2008) and "Serrania" gordaensis Liñán et al. (2008). |
|||
|
Evenkaspis (Bilobaspis) mirabilis[138] |
Subgen. et sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Tolmachev Formation |
A member of the family Pterygometopidae. |
||
|
Failleana superba[138] |
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Povorotnaya Formation |
A member of the family Styginidae. |
||
|
Sp. nov |
Valid |
Geyer |
Igoudine Formation |
A member of Redlichiida belonging to the family Fallotaspididae. |
||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
|||||
|
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Heisdorf Formation |
|||||
|
Heliopeltis morrisoni[143] |
Sp. nov |
Valid |
Basse, Müller & Habenicht |
A member of the family Scutelluidae belonging to the subfamily Kolihapeltinae. |
||||
|
Hollandiella[144] |
Gen. et 3 sp. nov |
Valid |
Van Viersen & Lerouge |
Ihandar Formation |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. The type species is H. curvirostris; genus also includes new species H. lebruni and H. verecunda. |
|||
|
Interproetus mizobuchii[145] |
Sp. nov |
Valid |
Stocker et al. |
Paleozoic |
||||
|
Issendalenia[135] |
Gen. et sp. nov |
Valid |
Geyer |
Igoudine Formation |
A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species I. grandispina. |
|||
|
Kasachstania alperovichi[146] |
Sp. nov |
Valid |
Ghobadi Pour et al. |
Shakshagaily Formation |
||||
|
Kellerwaldops[142] |
Gen. et sp. nov |
Valid |
Basse & Lemke |
A member of the family Phacopidae belonging to the subfamily Phacopinae. The type species is K. kaufmanni. |
||||
|
Kettneraspis eftychia[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of the family Odontopleuridae. |
|||
|
Koneprusia chimaera[141] |
Sp. nov |
Valid |
Alberti |
A member of the family Odontopleuridae. |
||||
|
Koneprusites oresibios[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
||||
|
Lepidoproetus arenicolus[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
||||
|
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Achegoninae. |
|||||
|
Sp. nov |
Valid |
Müller & Hahn |
A member of the family Phillipsiidae belonging to the subfamily Achegoninae. |
|||||
|
Nephranops (Maternia)[147] |
Subgen. et sp. nov |
Valid |
Feist |
A member of Phacopida. The type species of the subgenus is "Nephranops incisus" dillanus Richter & Richter (1926) (raised to the rank of a separate species Nephranops (Maternia) dillanus); the subgenus also includes Nephranops franconicus Alberti (1970) and a new species Nephranops (Maternia) occitanicus. |
||||
|
Nephranops (Nephranops) angerae[147] |
Sp. nov |
Valid |
Feist |
A member of Phacopida. |
||||
|
Niobe (Niobella) obscura[148] |
Sp. nov |
Valid |
Zhou & Zhou |
Dawan Formation |
A member of the family Asaphidae belonging to the subfamily Isotelinae. Originally described as a species of Niobe, but subsequently transferred to the genus Zhiyia.[149] |
|||
|
Sp. nov |
Valid |
Alberti |
A member of the family Scutelluidae. |
|||||
|
Sp. nov |
Valid |
Alberti |
A member of the family Scutelluidae. |
|||||
|
Pararemopleurides ornatissimus[138] |
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Povorotnaya Formation |
A member of Asaphida belonging to the family Remopleurididae. |
||
|
Pedinopariops ceuthonymus[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of the family Phacopidae. |
|||
|
Pedinopariops richterianus geminus[133] |
Subsp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of the family Phacopidae. |
|||
|
Sp. nov |
In press |
Pereira & Gutiérrez-Marco in Gutiérrez-Marco et al. |
Announced in 2019; the final version of the article naming it is not published yet. |
|||||
|
Polypleuraspidella[152] |
Gen. et comb. nov |
Valid |
Peel |
A new genus for "Polypleuraspis" cooperi Robison & Babcock (2011). |
||||
|
Polypleuraspis glacialis[152] |
Sp. nov |
Valid |
Peel |
Telt Bugt Formation |
||||
|
Pseudobigotina[135] |
Gen. et sp. nov |
Valid |
Geyer |
A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species P. antiatlasensis. |
||||
|
Pulvinocephalus[147] |
Gen. et comb. et 2 sp. et subsp. nov |
Valid |
Feist |
A member of Phacopida. The type species is "Phacops (Trimerocephalus)" steinachensis Richter & Richter (1926) (including a new subspecies Pulvinocephalus steinachensis triangulus); genus also includes "Phacops (Cryphops)" latilimbatus Maximova (1955) and new species Pulvinocephalus ovatus and Pulvinocephalus walliseri. |
||||
|
Sp. nov |
Valid |
Van Viersen & Magrean |
Middle Devonian |
Hanonet Formation |
A member of the family Odontopleuridae. |
|||
|
Raymondella plastron[138] |
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Povorotnaya Formation |
A member of the family Raphiophoridae. |
||
|
Sp. nov |
Valid |
Holmes, Paterson & García-Bellido |
||||||
|
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Barkov Formation |
A member of Asaphida belonging to the family Remopleurididae. |
|||
|
Sp. nov |
Valid |
Ove, Ebbestad & Fortey |
Late Ordovician |
Povorotnaya Formation |
A member of Asaphida belonging to the family Remopleurididae. |
|||
|
Scabriscutellum archinalae[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of the family Scutelluidae. |
|||
|
Sculptoproetus raki[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
||||
|
Septimopeltis akatastasia[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of the family Scutelluidae. |
|||
|
Struveops wiedaensis[147] |
Sp. nov |
Valid |
Feist |
A member of Phacopida. |
||||
|
Suvorovaella[135] |
Gen. et comb. nov |
Valid |
Geyer |
El'gyan Formation |
A member of Redlichiina belonging to the family Bigotinidae. The type species is "Bigotinops" privus Suvorova (1960); genus might also include "Bigotinops" patrius Suvorova (1960). |
|||
|
Sp. nov |
Valid |
Gutierrez-Marco, Rabano & Garcia-Bellido |
A member of the family Nileidae. |
|||||
|
Tafilaltaspis rufa[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
A member of the family Proetidae belonging to the subfamily Eremiproetinae. |
||||
|
Tesselacauda kriegerae[156] |
Sp. nov |
Valid |
Adrain & Karim |
Early Ordovician |
||||
|
Tesselacauda morrisoni[156] |
Sp. nov |
Valid |
Adrain & Karim |
Early Ordovician |
||||
|
Timsaloproetus gibbae[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
Khebchia Formation |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
|||
|
Timsaloproetus pulchistriatus[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
Timrhanrhart Formation |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
|||
|
Timsaloproetus weddigei[144] |
Sp. nov |
Valid |
Van Viersen & Lerouge |
Bicken-Ense Formation |
A member of the family Proetidae belonging to the subfamily Cornuproetinae. |
|||
|
Tioutella[135] |
Gen. et sp. nov |
Valid |
Geyer |
Igoudine Formation |
A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species T. floccofavosa. |
|||
|
Trifoliops septimanicus[147] |
Sp. nov |
Valid |
Feist |
A member of Phacopida. |
||||
|
Tropidocoryphe insciens[133] |
Sp. nov |
Valid |
Van Viersen, Taghon & Magrean |
Jemelle Formation |
A member of Proetida. |
|||
Other arthropods
Research
- A study on the morphology of appendages of the euarthropod Ercaicunia multinodosa from the Early Cambrian Chengjiang biota (China) is published by Zhai et al. (2019).[157]
- Description of the appendicular morphology of Sinoburius lunaris is published by Chen et al. (2019).[158]
- Description of a new specimen of Emeraldella brutoni from the Cambrian Wheeler Formation (Drum Mountains, Utah, United States), providing new information on the appendicular anatomy of this species, is published by Lerosey-Aubril & Ortega-Hernández (2019).[159]
- Description of new bradoriid fossils from the Chengjiang biota, revealing hitherto unknown morphological differences in limb anatomy between bradoriid species, is published by Zhai et al. (2019), who evaluate the implications of this finding for the knowledge of the phylogenetic relationships of bradoriids.[160]
- A study on the appendage structure of Naraoia spinosa at both juvenile and adult stages is published by Zhai et al. (2019).[161]
- Description of exceptionally preserved soft tissues from mature individuals of Peronopsis and Ptychagnostus from the Cambrian Burgess Shale (British Columbia, Canada), and a study on their implications for inferring the phylogenetic placement of agnostinids within Arthropoda, is published by Moysiuk & Caron (2019).[162]
- Completely enrolled, phosphatized specimens of Tsunyidiscus yanjiazhaiensis are described from Cambrian Stage 3 of South China by Dai et al. (2019).[163]
- The ontogenetic series of Pagetia vinusta is documented from the Cambrian (Wuliuan) Kaili Formation (Guizhou, China) by Cui et al. (2019).[164]
- A study reevaluating crystals of a copper sulfide mineral found in association with specimens of Marrella splendens from the Burgess Shale, originally interpreted as evidence that the animal's blood contained the Cu-bearing protein hemocyanin, is published by Gaines et al. (2019).[165]
- A study on the anatomy of the nervous system of Alalcomenaeus is published by Ortega-Hernández, Lerosey-Aubril & Pates (2019).[166]
- Juvenile specimens of stylonurine eurypterids belonging to the family Hardieopteridae are described from the Famennian locality of Strud (Belgium) by Lamsdell et al. (2019), who interpret their findings as evidence indicating that eurypterids underwent a marine to freshwater transition during the Devonian, and that juvenile eurypterids inhabited sheltered nursery pools (and migrated to rivers upon reaching maturity).[167]
- A study on the structure of the eyes of eurypterids is published by Schoenemann, Poschmann & Clarkson (2019).[168]
- A study on ontogenetic changes in prosomal morphology in Carboniferous horseshoe crab Euproops danae is published by Tashman, Feldmann & Schweitzer (2019).[169]
- A study on the anatomy of the appendages of the Cretaceous xiphosurid Tachypleus syriacus and on the evolution of fossil horseshoe crab appendages is published by Bicknell et al. (2019).[170]
- A study re-evaluating the fossil evidence for lateral compound eyes in Paleozoic horseshoe crabs is published by Bicknell, Amati & Ortega-Hernández (2019).[171]
- A study on the morphology of the millipede Phryssonotus burmiticus from the Cretaceous amber from Myanmar is published by Su, Cai & Huang (2019).[172]
New taxa
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Albalimulus[173] |
Gen. et sp. nov |
Bicknell & Pates |
A member of Limuloidea. The type species is A. bottoni. |
 | ||||
|
Andrognathus burmiticus[174] |
Sp. nov |
Valid |
Moritz & Wesener |
A millipede, a species of Andrognathus. |
||||
|
Ankitokazocaris tatensis[175] |
Sp. nov |
Disputed |
Ehiro et al. |
Osawa Formation |
A thylacocephalan. Laville et al. (2021) considered this species to be a junior synonym of Ankitokazocaris chaohuensis.[176] |
|||
|
Sp. nov |
Valid |
Zhu, Lerosey-Aubril & Ortega-Hernández |
Sandu Formation |
A member of Aglaspidida. |
||||
|
Gen. et sp. nov |
Valid |
Stoev, Moritz & Wesener |
A millipede belonging to the group Callipodida and to the new family Burmanopetalidae. The type species is B. inexpectatum. |
 | ||||
|
Gen. et sp. nov |
Valid |
Moysiuk & Caron |
A member of Radiodonta belonging to the family Hurdiidae. |
 | ||||
| Concavicaris parva[175] |
Sp. nov |
Valid |
Ehiro et al. |
Osawa Formation |
A thylacocephalan. Originally described as a species of Concavicaris; Laville et al. (2021) transferred this species to the genus Ankitokazocaris.[176] |
|||
|
Duraznovis[180] |
Gen. et sp. nov |
Valid |
Lara et al. |
Potrerillos Formation |
An arthropod of uncertain phylogenetic placement. Genus includes new species D. gallegoi. It was argued to be an insect belonging to the group Cicadomorpha by Fu & Huang (2020).[181] |
|||
|
Electrosminthuridia[182] |
Gen. et sp. nov |
Robin, D'Haese & Barden |
A springtail belonging to the family Sminthurididae. The type species is E. helibionta. |
|||||
|
Falcatacaris[183] |
Gen. et sp. nov |
Valid |
Braig et al. |
Altmühltal Formation |
A thylacocephalan. The type species is F. bastelbergeri. |
|||
|
Gen. et sp. nov |
Valid |
Izquierdo-López & Caron |
A bivalved arthropod belonging to the group Mandibulata. The type species is F. nereidis. |
 | ||||
|
Gen. et sp. nov |
Valid |
Lamsdell, Gunderson & Meyer |
Late Ordovician |
A member of Chasmataspidida belonging to the family Diploaspididae. The type species is H. hiawathai. |
 | |||
|
Hyleoglomeris groehni[186] |
Sp. nov |
Valid |
Wesener |
Europe (Baltic Sea region) |
A millipede belonging to the family Glomeridae. |
|||
|
Ligulacaris[187] |
Gen. et sp. nov |
Valid |
Charbonnier et al. |
A thylacocephalan. Genus includes new species L. parisiana. |
||||
|
Sp. nov |
Valid |
Hou et al. |
A probable member of Artiopoda and the order Xandarellida. |
 | ||||
|
Sp. nov |
Valid |
Tortello & Esteban |
Santa Rosita Formation |
A member of Agnostida. Announced in 2019; the final version of the article naming it was published in 2020. |
||||
|
Miyagicaris[175] |
Gen. et sp. nov |
Disputed |
Ehiro et al. |
Osawa Formation |
A thylacocephalan. Genus includes new species M. costata. However, Laville et al. (2021) considered M. costata to be a junior synonym of Ankitokazocaris utatsuensis.[176] |
|||
|
Sp. nov |
Valid |
Aria & Caron |
A member of Chelicerata belonging to the family Mollisoniidae. |
 | ||||
|
Parisicaris[187] |
Gen. et sp. nov |
Disputed |
Charbonnier et al. |
A thylacocephalan. Genus includes new species P. triassica. Laville et al. (2021) transferred P. triassica to the genus Ankitokazocaris,[176] while Ehiro & Kano (2024) considered Parisicaris to be a genus distinct from Ankitokazocaris.[191] |
||||
|
Pickettia[192] |
Gen. et comb. nov |
Valid |
Bicknell, Lustri & Brougham |
Late Devonian |
A member of Xiphosura; a new genus for "Bellinurus" carteri Eller, 1940. |
|||
|
Pseudobeyrichona monile[193] |
Sp. nov |
Valid |
Streng & Geyer |
Tannenknock Formation |
A member of Bradoriida belonging to the family Hipponicharionidae. |
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|
Siphonophora hui[194] |
Sp. nov |
Valid |
Jiang et al. |
A millipede belonging to the group Siphonophorida. |
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|
Sloveniolimulus[195] |
Gen. et sp. nov |
Valid |
Bicknell et al. |
Strelovec Formation |
A horseshoe crab. Genus includes new species S. rudkini. |
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|
Stilpnocephalus[196] |
Gen. et sp. nov |
Valid |
Selden, Simonetto & Marsiglio |
A horseshoe crab belonging to the family Belinuridae. The type species is S. pontebbanus. |
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|
Gen. et sp. nov |
Valid |
Bicknell |
Jackey Shale |
A member of Xiphosurida belonging to the family Austrolimulidae. The type species is T. patersoni. |
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|
Sp. nov |
Valid |
Wen et al. |
||||||
|
Gen. et sp. nov |
Valid |
Du et al. |
Xiaoshiba Lagerstätte |
A non-trilobite artiopodan. Genus includes new species Z. coronata. |
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