2020 in paleomalacology

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This list 2020 in paleo malacology is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that were described during the year 2020, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2020.

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acrimeroceras hoppeckense^[1]	Sp. nov	Valid	Korn & Buchwald in Korn et al.	Late Devonian		Germany
Acrimeroceras ropicense^[1]	Sp. nov	Valid	Korn, Bartzsch & Weyer in Korn et al.	Late Devonian
Acrimeroceras saalense^[1]	Sp. nov	Valid	Korn, Bartzsch & Weyer in Korn et al.	Late Devonian
Acutimitoceras alabasense^[2]	Sp. nov	Valid	Nikolaeva	Devonian-Carboniferous boundary		Kazakhstan
Acutimitoceras dzhanganense^[2]	Sp. nov	Valid	Nikolaeva	Devonian-Carboniferous boundary		Kazakhstan
Almites hayasakai^[3]	Sp. nov	Valid	Ehiro & Ozawa	Permian (Asselian)	Taishaku Limestone	Japan
Angolaites elegans^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Aspidoceras schweigertense^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Aulacostephanus camericensis^[6]	Sp. nov	Valid	Cope & Etches	Late Jurassic (Kimmeridgian)		United Kingdom
Baidites obesus^[7]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Boesites biconcavus^[3]	Sp. nov	Valid	Ehiro & Ozawa	Permian (Asselian)	Taishaku Limestone	Japan
Borissjakoceras falcatum^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Tunisia	A member of the family Binneyitidae.
?Bouleiceras betetensis^[9]	Sp. nov	Valid	Martínez & Joral	Early Jurassic (Toarcian)		Spain	A member of the family Hildoceratidae.
Bouleiceras ibericum^[9]	Sp. nov	Valid	Martínez & Joral	Early Jurassic (Toarcian)		Spain	A member of the family Hildoceratidae.
Cagliceras baldii^[10]	Sp. nov	Valid	Kovács, Dunai & Evanics	Early Jurassic (Toarcian)		Hungary	A member of the family Erycitidae.
Cagliceras wegeneri^[10]	Sp. nov	Valid	Kovács, Dunai & Evanics	Early Jurassic (Toarcian)		Hungary	A member of the family Erycitidae.
Carthaginites elegans^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Turrilitidae.
Churkites warei^[7]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Collignoniceras badilleti^[11]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Collignoniceras hourqueigi^[11]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Collignoniceras vigennum^[11]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Conlinites evolutum^[8]	Sp. nov	Valid	Kennedy	Early Cretaceous (Albian)		Algeria	A member of the family Brancoceratidae.
Coquandiceras^[8]	Gen. et comb. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria Tunisia	A member of the family Acanthoceratidae belonging to the subfamily Mantelliceratinae. The type species is "Ammonites" jubae Coquand (1880); genus also includes "Ammonites" villei Coquand (1862).
Cryptoturrilites^[8]	Gen. et sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Tunisia	A member of the family Turrilitidae. The type species is C. tenuicostatus.
Elobiceras (Craginites) simplex^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Elobiceras (Craginites) sparcicostatum^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Flemingites lidakensis^[7]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Goniatites abaiensis^[12]	Sp. nov	Valid	Korn et al.	Carboniferous (Viséan)	Baktysai Formation	Kazakhstan
Goniatites zhankurganensis^[12]	Sp. nov	Valid	Korn et al.	Carboniferous (Viséan)	Baktysai Formation	Kazakhstan
Graysonites elegans^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Acanthoceratidae.
Gregoryceras boreale^[13]	Sp. nov	Valid	Mitta & Stupachenko	Late Jurassic (Oxfordian)		Russia	A member of the family Aspidoceratidae.
Hoevelia megalomanica^[14]	Sp. nov	Valid	Korn, Hairapetian & Gholamalian	Late Devonian		Iran
Homeomorphites^[15]	Gen. et sp. et comb. nov	Valid	Bert, Bersac & Canut	Early Cretaceous (Hauterivian to Barremian)		Argentina Chile	A member of the family Neocomitidae. The type species is H. aguirreurretae; genus also includes H. ploszkiewiczi (Riccardi and Aguirre-Urreta, 1989) and H. varicostatus (Riccardi and Aguirre-Urreta, 1989). Announced in 2019; the final version of the article naming was published in 2020.
Homoceratoides wangchengi^[16]	Sp. nov	Valid	Korn & Wang in Korn et al.	Carboniferous (Bashkirian)	Jingyuan Formation	China	A member of the family Bisatoceratidae. Announced in 2020; the final version of the article naming was published in 2021.
Hoploscaphites peterseni^[17]	Sp. nov	Valid	Landman et al.	Late Cretaceous	Bearpaw Shale Pierre Shale	United States ( Montana Wyoming)
Howarthiceras^[18]	Gen. et comb. nov	Valid	Zakharov & Rogov	Upper Jurassic (Tithonian)		United States ( California)	A new genus for "Groebericeras" baylei Imlay & Jones.
Hybonoticeras pseudoharpephorum^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Hypowaagenia^[19]	Gen. et sp. nov	Valid	Schweigert & Schlampp	Late Jurassic (Kimmeridgian)	Arzberg Formation	Germany Pakistan?	A member of the family Aspidoceratidae. The type species is H. endressi; genus might also include "Aspidoceras" acanthomphalum Zittel (1870).
Kosmoclymenia ebbighauseni^[20]	Sp. nov	Valid	Korn	Devonian (Famennian)		Morocco
Lasswitzia^[4]	Gen. et comb. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek Formation	United States ( Texas)	A member of the family Brancoceratidae. The type species is "Schloenbachia austinensis" Roemer var. minima Lasswitz (1904) (raised to the rank of a separate species Lasswitzia minima).
Lenicostites^[15]	Gen. et comb. nov	Valid	Bert, Bersac & Canut	Early Cretaceous (Barremian)		France	A member of Ancyloceratoidea belonging to the new family Lenicostitidae. The type species is "Hemihoplites" rusticus Vermeulen (1996). Announced in 2019; the final version of the article naming was published in 2020.
Lithacoceras főzyi^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Litophragmatoceras curiosus^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Flickiidae.
Lytoceras pseudorsinii^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Marcouxia chaiburiensis^[21]	Sp. nov	Valid	Tongtherm et al.	Early Triassic	Chaiburi	Thailand
Margoceras^[22]	Gen. et sp. nov	Valid	Borissenkov	Permian (Cisuralian)		Russia ( Nenets Autonomous Okrug)	A member of the family Paragastrioceratidae. Genus includes new species M. boreale.
Mittaites^[23]	Gen. et comb. nov	Valid	Frau et al.	Early Cretaceous (Berriasian)		Russia	The type species is "Mazenoticeras" ceccai. Announced in 2020; the final version of the article naming it was published in 2021.
Mortoniceratoides elegans^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Kiamichi	United States ( Texas)	A member of the family Brancoceratidae.
Muessenbiaergia bockwinkeli^[20]	Sp. nov	Valid	Korn	Devonian (Famennian)		Morocco	A member of the family Kosmoclymeniidae.
Neophlycticeras algeriense^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Lyelliceratidae.
Neophyllites lavernockensis^[24]	Sp. nov	Valid	Hodges	Early Jurassic (Hettangian)	White Lias Formation	United Kingdom	Announced in 2020; the final version of the article naming it was published in 2021.
Neotelodactylites^[25]	Gen. et comb. nov	Valid	Jattiot, Gris & Trincal	Early Jurassic (Toarcian)		France Italy	A new genus for "Peronoceras" eucosmum Lippi Boncambi (1947).
Neouddenites echiensis^[26]	Sp. nov	Valid	Kutygin	Permian		Russia ( Sakha)	A member of the family Medlicottiidae.
Paracraginites^[4]	Gen. et sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae. The type species is P. conlini.
Paracrioceras giraudae^[27]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous		France
Paragastrioceras cantabrignianum^[22]	Sp. nov	Valid	Borissenkov	Permian (Cisuralian)		Russia ( Nenets Autonomous Okrug)	A member of the family Paragastrioceratidae.
Paraspidites bicarinatus^[7]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Paratornoceras ayense^[1]	Sp. nov	Valid	Korn & Buchwald in Korn et al.	Late Devonian
Paratornoceras harounense^[1]	Sp. nov	Valid	Korn & Ebbighausen in Korn et al.	Late Devonian
Paratornoceras peterseni^[1]	Sp. nov	Valid	Korn & Buchwald in Korn et al.	Late Devonian
Paratornoceras thuringense^[1]	Sp. nov	Valid	Korn, Bartzsch & Weyer in Korn et al.	Late Devonian
Pernaceras leroyi^[28]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous		France
Pervinquieria (Deiradoceras) serpentiforme^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Pervinquieria (Pervinquieria) asper^[4]	Sp. nov	Valid	Kennedy in Gale & Kennedy	Early Cretaceous (Albian)	Lower Duck Creek	United States ( Texas)	A member of the family Brancoceratidae.
Procrioceratites^[27]	Gen. et sp. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous		France	A member of the family Crioceratitidae. The type species is P. etiennei; genus also includes P. primitivus (Reboulet, 1996) and P. barrabei (Sarkar, 1955).
Protacanthoceras sottaraense^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria	A member of the family Acanthoceratidae.
Protacanthodiscus guerrai^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Prouddenites evolutus^[26]	Sp. nov	Valid	Kutygin	Permian		Russia ( Sakha)	A member of the family Medlicottiidae.
Pseudodiscosphinctes pseudorhodanicus^[5]	Sp. nov	Valid	Caracuel & Sarti in Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Pseudolissoceras atesinum^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Pseudolongaeviceras^[29]	Gen. et sp. nov	Valid	Alifirov & Knyazev	Middle Jurassic (Callovian)		Russia	A member of the family Cardioceratidae. Genus includes new species P. densicostatum.
Pseudoxybeloceras simplex^[30]	Sp. nov	Valid	Kennedy	Late Cretaceous		United Kingdom
Ptychites embreei^[31]	Sp. nov	Valid	Bischof & Lehmann	Middle Triassic (Anisian)	Favret Formation	United States ( Nevada)
Reymenticoceras ornatum^[32]	Sp. nov	Valid	Kennedy	Late Cretaceous (Turonian)		Colombia	A member of the family Acanthoceratidae belonging to the subfamily Mammitinae. Announced in 2020; the final version of the article naming it was published in 2021.
Romaniceras (Romaniceras) marigniacum^[11]	Sp. nov	Valid	Amédro & Châtelier in Amédro et al.	Late Cretaceous (Turonian)		France
Roopnarinites^[7]	Gen. et comb. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia	Genus includes "Meekoceras" paucesculptatum Welter (1922).
Scaphites occlusus^[8]	Sp. nov	Valid	Kennedy	Late Cretaceous (Cenomanian)		Algeria Tunisia	A member of the family Scaphitidae.
Schaireria (Schaireria) vetera^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Shumardites umbilicatus^[3]	Sp. nov	Valid	Ehiro & Ozawa	Permian (Asselian)	Taishaku Limestone	Japan
Sormaites^[33]	Gen. et sp. nov	Valid	Muramiya & Shigeta	Late Cretaceous (Turonian)		Japan	A member of the family Diplomoceratidae. Genus includes new species S. teshioensis.
Stoliczkaia (Stoliczkaia) djerissaensis^[8]	Sp. nov	Valid	Kennedy	Early Cretaceous (Albian)		Tunisia	A member of the family Lyelliceratidae.
Subflemingites bihatiense^[7]	Sp. nov	Valid	Jattiot, Bucher & Brayard	Early Triassic		Indonesia
Tescheniceras^[34]	Gen. et comb. nov	Valid	Vašíček	Early Cretaceous (Valanginian to Hauterivian)		Austria France Italy Morocco Romania Serbia Slovakia Spain Switzerland	A member of the family Neocomitidae. The type species is "Neocomites (Teschenites)" flucticulus Thieuloy (1977); genus also includes "Neocomites (Teschenites)" callidiscus Thieuloy (1971), "Teschenites" subflucticulus Reboulet (1996), "Neocomites (Teschenites)" pachydicranus Thieuloy (1977) and "Teschenites" subpachydicranus Reboulet (1996).
Tumaroceras paveli^[22]	Sp. nov	Valid	Borissenkov	Permian (Cisuralian)		Russia ( Nenets Autonomous Okrug)	A member of the family Paragastrioceratidae.
Vidrioceras ellipticum^[3]	Sp. nov	Valid	Ehiro & Ozawa	Permian (Asselian)	Taishaku Limestone	Japan
Virgatosimoceras propecostatum^[5]	Sp. nov	Valid	Sarti	Late Jurassic (Tithonian)	Rosso Ammonitico Veronese Formation	Italy
Yezoceras elegans^[35]	Sp. nov	Valid	Aiba, Karasawa & Iwasaki	Late Cretaceous (Coniacian)		Japan	A member of the family Nostoceratidae.

General research

A study aiming to determine the drag force experienced by a wide range of ammonite shell shapes is published by Hebdon, Ritterbush & Choi (2020).^[36]
A study evaluating how different parameters affected the ammonite shell's response to water pressure, aiming to determine whether the ammonite septa strengthened the shell against pressure at increasing water depths, is published by Lemanis (2020).^[37]^[38]^[39]
A study on global patterns of ammonite diversification and extinction from the Early Devonian (Emsian) to the Early Triassic (Induan) is published by Whalen, Hull & Briggs (2020).^[40]
A study on the abundance and size distribution of small specimens of Baculites and Hoploscaphites at eight cold methane seep sites in the Upper Cretaceous Pierre Shale (South Dakota, United States) is published by Rowe et al. (2020), who interpret their findings as indicating that newly hatched ammonites lived in close proximity to seep fluids emerging at the sediment-water interface and the associated microbial food web.^[41]
A study on changes of hydrostatic properties of three species of Didymoceras during their ontogeny is published by Peterman et al. (2020).^[42]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Alexoceras^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid related to Temnocheilus. Genus includes new species A. mazaevi.
Ancistroceras ristnensis^[44]	Sp. nov	Valid	Aubrechtová & Meidla	Ordovician		Estonia
Ancistroceras vahikuelaensis^[44]	Sp. nov	Valid	Aubrechtová & Meidla	Ordovician		Estonia
Bairstowius amaliae^[45]	Sp. nov	Valid	Rita et al.	Early Jurassic (Pliensbachian)	Lemede	Portugal	A belemnite belonging to the family Hastitidae.
Barskoceras^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A probable member of Oncocerida. Genus includes new species B. mirum.
Chromatoceras^[46]	Gen. et comb. nov	Valid	Turek & Manda	Silurian (Pridoli)	Kopanina	Czech Republic	A member of Oncocerida belonging to the family Oonoceratidae. The type species is "Cyrtoceras" veteranum Barrande (1866); genus also includes "Cyrtoceras" constringens Barrande (1866).
Domatoceras bashkiricum^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)
Domatoceras sterlitamakense^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)
Enigmabelus^[47]	Gen. et 3 sp. nov	Valid	Keupp & Fuchs	Early Jurassic (Pliensbachian)	Amaltheenton	Germany	A member of Belemnitina. Genus includes new species E. doppelsteini, E. micros and possibly ? E. leptos.
Houdongoceras^[48]	Gen. et sp. nov	Valid	Yang et al.	Permian Guadalupian	Gufeng Formation	China	A member of Orthocerida. Genus includes new species H. chaohuensis. Announced in 2020; the final version of the article naming it was published in 2021.
Leniceras^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid belonging to the family Liroceratidae. Genus includes new species L. ovale.
Liroceras shakhtauense^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid belonging to the family Liroceratidae.
Lituites nehatuensis^[44]	Sp. nov	Valid	Aubrechtová & Meidla	Ordovician		Estonia
Megaglossoceras barskovi^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid belonging to the family Ephippioceratidae.
Mosquoceras planum^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)
Shatoceras^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid related to Domatoceras. Genus includes new species S. umbilicatum.
Shikhanonautilus^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid belonging to the family Liroceratidae. Genus includes new species S. siphonoventralis.
Sholakoceras formosum^[43]	Sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)
Thyoceras^[43]	Gen. et sp. nov	Valid	Leonova & Shchedukhin	Early Permian		Russia ( Bashkortostan)	A nautiloid related to Permonautilus. Genus includes new species T. involutum.
Trematoceras watanabei^[49]	Sp. nov	Valid	Niko & Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan

General research

A study aiming to determine whether the development of chamber volume in fossil cephalopod phragmocones conveys information about their physiology is published by Tajika et al. (2020), who evaluate the implications of their findings for the knowledge of the ecology and extinction selectivity of fossil cephalopods.^[50]
A study on the spatial distribution of Givetian oncocerid fossils from the Hamar Laghdad ridge (eastern Anti-Atlas, Morocco), evaluating its implications for the knowledge of the life cycle of these cephalopods, is published by Pohle et al. (2020).^[51]
Jurassic nautiloid Somalinautilus antiquus, previously known only from the Kimmeridgian strata of Ethiopia, is reported from the Kimmeridgian of Southern Germany by Schweigert (2020), who also reports the first finding of the species Somalinautilus clavifer the Middle Jurassic (Bathonian) of Southern Germany.^[52]
A study on the reproductive strategies of fossil coleoids, based on data from shells of belemnoids, spirulids and sepiids, is published by Fuchs et al. (2020).^[53]
A specimen of Plesioteuthis preserved with an associated tooth of a pterosaur is reported from the Upper Jurassic Altmühltal Formation (Germany) by Hoffmann et al. (2020), who evaluate the implications of this finding for the knowledge of the likely habitat of Plesioteuthis.^[54]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acanthinula karsdorfensis^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A species of Acanthinula sensu lato.
Acroloxus aspis^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A species of Acroloxus.
Acroloxus korys^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A species of Acroloxus.
"Acteon" distinctus^[56]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France
Afrodontops^[55]	Gen. et 2 sp. nov	Valid	Kadolsky	Early Eocene		Germany	A member of the family Charopidae. The type species is A. europaea; genus also includes A. comes.
Alanstukella^[57]	Gen. et comb. nov	Valid	Mazaev	Carboniferous (Kasimovian) to Permian (Sakmarian)		Russia	A member of the family Trochonematidae. The type species is A. rossica (Stuckenberg, 1905).
Albinulopsis^[55]	Gen. et sp. et comb. nov	Valid	Kadolsky	Eocene		France Germany	A member of the family Gastrocoptidae. The type species is A. gibba; genus also includes "Pupa" bonneti Cossmann (1907).
Altotomaria^[58]	Gen. et sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand	Genus includes new species A. reticulata.
Ammonicera oliveri^[56]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Ammonicera.
Andonia delgadoi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France
Andonia wilhelminamariae^[60]	Sp. nov	Valid	Landau & Mulder	Pliocene		Italy Spain
Anomphalus murankaensis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Apachella noinskyi^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Apachella thailandensis^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Aphanitoma roesti^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Aphanitoma.
Aquitanobursa javana^[62]	Sp. nov	Valid	Landau et al.	Miocene		Indonesia	A member of the family Bursidae.
Assiminea striatura^[63]	Sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Assiminea.
Auristomia insulsa^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Auristomia.
Bactrocythara pascaleae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bactrocythara.
Balticopta^[65]	Gen. et sp. nov	Valid	Balashov & Perkovsky	Eocene (Priabonian)	Baltic amber	Russia ( Kaliningrad Oblast)	A member of the family Gastrocoptidae. The type species is B. gusakovi.
Bania goehlichae^[66]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Bela henkmulderi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela keukelaari^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela pseudomegastoma^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela pseudovulpecula^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela redoniana^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela scarponii^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Bela sceauxensis^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Bela.
Belgrandiella? nichterli^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Possibly a species of Belgrandiella.
Bellerophon erawanensis^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Biarmeaspira mazaevi^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Buccinaria minuscula^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Buccinaria.
Calculocochlea^[68]	Gen. et sp. nov	Valid	Juárez-Ruiz, Matamales-Andreu & Kadolsky	Early Oligocene		Spain	A member of the family Vidaliellidae. Genus includes new species C. oliveri.
Callitrochilia^[69]	Gen. et comb. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A member of the family Calliostomatidae. The type species is "Calliostoma (Eutrochus)" ditropis Cossmann & Pissarro (1902).
Calogoniodiscus pyrenaicus^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Campanile madagasikara^[70]	Sp. nov		Pacaud	Late Miocene		Madagascar	A species of Campanile.
Careliopsis gallica^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Careliopsis.
Carychiopsina^[55]	Gen. et comb. nov	Valid	Kadolsky	Late Oligocene and early Miocene		Czech Republic Poland	A member of the family Ellobiidae belonging to the subfamily Carychiinae. The type species is "Pupa" schwageri Reuss (1868); genus also includes C. costulata (Sandberger, 1862), C. prisyazhnyuki (Stworzewicz, 1999) and possibly C.? surai (Stworzewicz, 1999).
Carychium bachi^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A species of Carychium.
Carychium vagum^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A species of Carychium.
Casmaria primitiva^[62]	Sp. nov	Valid	Landau et al.	Miocene		Indonesia	A species of Casmaria.
Cassis robusta^[62]	Sp. nov	Valid	Landau et al.	Miocene		Indonesia	A species of Cassis.
Cassitriton^[71]	Gen. et sp. nov	Valid	Craig & Tracey in Craig et al.	Eocene (Lutetian)		France	A member of the family Cymatiidae. The type species is C. magnificus Craig, Tracey & Gain.
Cataegis ramosi^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A species of Cataegis.
Cheilospicata cedrus^[73]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Chemnitzia miogallica^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Chemnitzia.
Chemnitzia robusticostata^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Chemnitzia.
Chilodonta tricostatula^[74]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Chilodonta.
Clanculus (Clanculopsis) krachi^[75]	Sp. nov	Valid	Nosowska	Middle Miocene		Poland Ukraine	A species of Clanculus.
Clathrella semilaeve^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France
Clathromangelia daisyae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia densecostata^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia hakkennesi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia helwerdae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia karinneae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia pereirae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia vannieulandei^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clathromangelia wopkeae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clathromangelia.
Clavatula sceauxensis^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Clavatula.
Clio gailae^[73]	Sp. nov	Valid	Goedert & Janssen in Garvie, Goedert & Janssen			United States ( Washington)	A species of Clio.
Cochleochilus? spiralocostatus^[74]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	Possibly a member of the family Ataphridae.
Coenaculum boucheti^[56]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Coenaculum.
Cryptochorda altavesna^[76]	Sp. nov	Valid	Pacaud & Sautereau	Eocene (Lutetian)		France	A member of the family Harpidae.
Cryptochorda cosediensis^[76]	Sp. nov	Valid	Pacaud & Sautereau	Eocene (Lutetian))		France	A member of the family Harpidae.
Cryptochorda neptis^[76]	Sp. nov	Valid	Pacaud & Sautereau	Eocene (Ypresian))		France	A member of the family Harpidae.
Currylimacina asperita^[73]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Cypraecassis minirufa^[62]	Sp. nov	Valid	Landau et al.	Miocene		Indonesia	A species of Cypraecassis.
Cyrillia michalidesi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Cyrillia.
Daphnella groeneveldi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Daphnella.
Donaldina sakmaraensis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Ebala ornatissima^[56]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Ebala.
Emarginula rara^[74]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Emarginula.
Enneopupa priabonica^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Eoconus cambieni^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France
Eoconus vanhoutenae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France
Eofavartia dharmai^[77]	Sp. nov	Valid	Merle & Landau	Middle Miocene		Indonesia	A member of the family Muricidae.
Eohispania^[67]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species E. dominicii.
Eotrichophorus^[63]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cyclophoridae. Genus includes new species E. kachin.
Erato crassus^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato fragilis^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato pinguis^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato princeps^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato prokimakowiczi^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato quasiplanulosa^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Erato tenuipustulata^[78]	Sp. nov	Valid	Fehse	Miocene		France	A species of Erato.
Eulimella redoniana^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Eulimella.
Eulimella semilaeve^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Eulimella.
Eusassia^[71]	Gen. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Cymatiidae. The type species is "Triton (Lampusia)" chalmasi Vasseur (1882); genus also includes "Eutritonium (Sassia)" dubusi Cossmann & Pissarro (1905).
Euthema annae^[79]	Sp. nov	In press	Balashov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Euthema.
Euthema truncatellina^[80]	Sp. nov	Valid	Balashov, Perkovsky & Vasilenko	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Euthema.
Euthriofusus postumus^[60]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain
Freboldia carinii^[81]	Sp. nov	Valid	Pieroni & Nützel	Middle Triassic	Esino Limestone	Italy
Fusulculus hanseni^[82]	Nom. nov	Valid	Pacaud	Early Paleocene		Denmark	A species of Fusulculus; a replacement name for Epalxis ? rosenkrantzi Hansen (2019).
Gosseletina microstriata^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand	A member of the family Gosseletinidae. Originally described as a species of Gosseletina; Mazaev (2021) transferred it to the genus Nemaspira.^[83]
Haedropleura brebioni^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Haedropleura.
Haedropleura gallica^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Haedropleura.
Haedropleura ligeriana^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Haedropleura.
Headonia^[67]	Gen. et comb. nov	Valid	Harzhauser et al.	Late Eocene	Headon Hill Formation	France United Kingdom	The type species is "Planorbis" euomphalus Sowerby (1816)
Heliconoides hodgkinsoni^[73]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States
Helvetataphrus^[74]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	France Switzerland	A member of the family Ataphridae. The type species is "Turbo" subrugosus Buvignier (1852).
Heterogen praelongispira^[84]	Sp. nov	Valid	Matsuoka & Hirano	Pleistocene	Katata Formation	Japan	A species of Heterogen.
Heterosubulites longusapertura^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Incisilabium trochiformis^[85]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Eocene (Priabonian)		France	A member of the family Modulidae.
Ividella tuberculata^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A member of the family Pyramidellidae. Originally described as a species of Ividella, but subsequently transferred to the genus Mulderia.^[86]
Katosira ? bicarinata^[87]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of the family Zygopleuridae.
Klavlia minuta^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Laminifera nannodes^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Laxecostula^[67]	Gen. et comb. nov	Valid	Harzhauser et al.	Eocene (Lutetian)	Bouxwiller Formation	France	Genus includes "Helix" laxecostulata Sandberger (1871).
Leufroyia annegienae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Leufroyia hesseli^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Leufroyia ligeriana^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Leufroyia renauleauensis^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Leufroyia riccardoi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Leufroyia seani^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Leufroyia.
Limacina parvabrazensis^[73]	Sp. nov	Valid	Garvie & Janssen in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina pseudopygmaea^[73]	Sp. nov	Valid	Garvie & Janssen in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina texanopsis^[73]	Sp. nov	Valid	Garvie in Garvie, Goedert & Janssen			United States	A species of Limacina.
Limacina yadongensis^[88]	Sp. nov	Valid	Li et al.	Early Eocene	Zhepure Formation	China	A species of Limacina.
Loriolotrema^[89]	Gen. et sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic		Switzerland	A member of the family Brachytrematidae. Genus includes new species L. liriola.
Lozouetina naonedosca^[90]	Sp. nov	Valid	Pacaud & Gómez	Eocene (Bartonian)		France Spain	A member of the family Ovulidae belonging to the subfamily Pediculariinae.
Macromphalina alinemulderae^[60]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Macromphalina.
Macropupina^[63]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pupinidae. Genus includes new species M. electricus.
Mangelia burgersae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Mangelia.
Margarites hayashii^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A species of Margarites.
Megastomia pseudopolysarcula^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Megastomia.
Menestho bertieae^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Menestho.
Microsassia^[71]	Gen. et sp. nov	Valid	Craig & Tracey in Craig et al.	Eocene (Lutetian)		France	A member of the family Cymatiidae. The type species is M. inconspicua Craig & Tracey.
Microtriton^[71]	Gen. et 2 sp. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Cymatiidae. The type species is M. biplicoides Craig & Tracey; genus also includes new species M. dollfusi Craig & Tracey, as well as "Lampusia (Simpulum)" bourdoti Cossmann (1897).
Mitromorpha mulderi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Mitromorpha.
Modulus benoisti^[85]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Miocene (Serravallian)		France	A species of Modulus.
Moelleria jansseni^[91]	Sp. nov	Valid	Hoeksema, Rijken & Simons	Pleistocene		France	A species of Moelleria.
Monoplex dissimilis^[62]	Sp. nov	Valid	Landau et al.	Miocene		Indonesia	A species of Monoplex.
Monoplex gradatus^[71]	Sp. nov	Valid	Craig, Tracey & Gain in Craig et al.	Eocene		France	A species of Monoplex.
Nassitriton^[71]	Gen. et sp. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Cymatiidae. The type species is N. conicus Craig, Tracey & Le Renard; genus might also include "Lampusia (Sassia)" lennieri Cossmann & Pissarro (1901).
Naticopsis sokensis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Nemdaella usensis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Neniopsilla^[67]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species N. amoena.
Neritopsis brasili^[92]	Sp. nov	Valid	Symonds, Gain & Le Renard	Eocene		France	A species of Neritopsis.
Neritopsis toddi^[92]	Sp. nov	Valid	Symonds, Gain & Le Renard	Eocene		France	A species of Neritopsis.
Niveria ledoni^[93]	Sp. nov	Valid	Fehse & Vicián	Late Miocene		France	A species of Niveria.
Niveria miocarinata^[93]	Sp. nov	Valid	Fehse & Vicián	Late Miocene		France	A species of Niveria.
Ocinebrina bertai^[94]	Sp. nov	Valid	Kovács	Middle Miocene		Hungary	A species of Ocinebrina.
Odostomia fortistriata^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Odostomia.
Odostomia miopupa^[64]	Nom. nov	Valid	Landau et al.			France	A species of Odostomia; a replacement name for Odontostomia rissoides var. pupa Gougerot (1969).
Odostomia robustissima^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Odostomia.
Ovicarychium^[55]	Gen. et 2 sp. et comb. nov	Valid	Kadolsky	Eocene		France Germany	A member of the family Ellobiidae belonging to the subfamily Carychiinae. The type species is O. ronlederi; genus also includes new species O.hennigeri, as well as O. bigeminatum (Deshayes, 1863), O. carbonarium (Wenz, 1932), O. interferens (Deshayes, 1863), O. polysarcum (Cossmann, 1914), O. quadridens(Andreae, 1884), O. remiense (Boissy, 1848) and O. sparnacense (Deshayes, 1863).
Oxfordina^[74]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Eucyclidae. The type species is "Trochus" crassicosta Buvignier (1852).
Palaeostoa costellata^[55]	Sp. nov	Valid	Kadolsky	Early Eocene		Germany	A member of the family Palaeostoidae.
Paleodiplommatina^[63]	Gen. et sp. nov	Disputed	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diplommatinidae. Genus includes new species P. spelomphalos. Considered to be a junior synonym of the genus Euthema by Balashov (2020).^[79]
Paracraticula? catalaunica^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Parasassia^[71]	Gen. et comb. et 2 sp. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Cymatiidae. The type species is "Triton" multigraniferum Deshayes (1835); genus also includes "Triton" bicinctum Deshayes (1835), "Triton" dumortieri Baudon (1853), "Lampusia (Simpulum)" polysarca Cossmann & Pissarro (1901), "Lampusia (Simpulum)" pustulifera Cossmann & Pissarro (1901) and "Murex" reticulosus Lamarck (1803), as well as new species P. globosa Craig & Tracey and P. polysarcoides Craig & Tracey.
Parthenina brebioni^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina chauvereauensis^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina clementiensis^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina lamellata^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina ligeriana^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina milleti^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina pouweri^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina redoniana^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina sceauxensis^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina tenuicostata^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Parthenina wesselinghi^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Parthenina.
Patella staceata^[74]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A species of Patella.
Perissocyclos^[63]	Gen. et sp. nov	Valid	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cyclophoridae. Genus includes new species P. kyrtostoma.
Peruvispira brasilensis^[95]	Sp. nov	In press	Simões et al.	Late Paleozoic	Taciba Formation	Brazil
Planitriton^[71]	Gen. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene	Calcaire Grossier d'Etrechy	France	A member of the family Cymatiidae. The type species is "Triton" planicostatum Deshayes (1835).
Platyzona gradata^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Pleurotomaria pogibshiensis^[96]	Sp. nov	Valid	Ferrari et al.	Early Jurassic (Hettangian)	Pogibshi Formation	United States ( Alaska)
Pleurotomoides barnardi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France
Pleurotomoides vanderdoncki^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France
Praealaerato venusta^[78]	Sp. nov	Valid	Fehse	Miocene		France	A member of the family Eratoidae.
Profundimitra lacuiensis^[97]	Nom. nov	Valid	Nielsen & Ampuero	Early Miocene		Chile	A species of Profundimitra; a replacement name for Mitra martini Philippi (1887).
Prososthenia krijgsmani^[66]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Prososthenia milosevici^[98]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Prososthenia? naissensis^[98]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Prososthenia rundici^[98]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Hydrobiidae belonging to the subfamily Pyrgulinae.
Protocharonia^[71]	Gen. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Charoniidae. The type species is "Triton" formosum Deshayes (1865); genus also includes "Eutritonium (Sassia) gouetense" var. semitextum Cossmann (1919).
Protoplex^[71]	Gen. et sp. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France United Kingdom	A member of the family Cymatiidae. The type species is P. gervillei Craig & Tracey; genus also includes "Murex" nodularius Lamarck (1803), "Sassia" websteri Wrigley (1932) and "Sassia" biserialis Wrigley (1932).
Provanna azurini^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A species of Provanna.
Pseudangaria^[74]	Gen. et comb. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Angariidae. The type species is "Delphinula" kobyi De Loriol in De Loriol & Koby (1895).
Pseudoneptunea constantinensis^[99]	Sp. nov	Valid	Gain & Belliard	Eocene		France
Pseudoneptunea prima^[99]	Sp. nov	Valid	Gain & Belliard	Paleocene (Thanetian)		France
Pseudoneptunea trinodularia^[99]	Sp. nov	Valid	Gain & Belliard	Eocene		France
Pseudoscilla breitenbergeri^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Pseudoscilla.
Pseudotectonica^[67]	Gen. et sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain	Genus includes new species P. pulchra.
Pyrgulina cancellatissima^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Pyrgulina.
Pyrgulina presselierensis^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Pyrgulina.
Raphitoma breitenbergeri^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Raphitoma.
Raphitoma dellabellaorum^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Raphitoma.
Raphitoma soniusae^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Raphitoma.
Raphitoma vogeli^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Raphitoma.
Rimula lobilloensis^[60]	Sp. nov	Valid	Landau & Mulder	Pliocene		Spain	A species of Rimula.
Rimulopsis torulosus^[74]	Sp. nov	Valid	Gründel, Hostettler & Gfeller	Late Jurassic (Oxfordian)	St-Ursanne-Formation	Switzerland	A member of the family Fissurellidae.
Rotatrigonostoma^[100]	Gen. et sp. et comb. nov	Valid	Landau, Breitenberger & Harzhauser	Miocene		Austria Romania	A member of the family Cancellariidae. Genus includes new species R. reinholdkunzi, as well as "Pseudomalaxis" boettgeri Cossmann (1916).
Sassiella^[71]	Gen. et 11 sp. et comb. nov	Valid	Craig & Tracey in Craig et al.	Eocene		France	A member of the family Cymatiidae. The type species is S. lacertina Craig & Tracey; genus also includes new species S. angulata Craig & Tracey, S. bacata Craig & Tracey, S. caputdraconis Craig & Tracey, S. crassilabrum Craig & Tracey, S. decussata Craig & Tracey, S. multivittata Craig & Tracey, S. puncticarinata Craig & Tracey, S. radulfivillensis Craig & Tracey, S. scalaris Craig & Tracey and S. vincta Craig & Tracey, as well as "Murex" colubrinus Lamarck (1803) and "Lampusia (Simpulum) polyzonalis" var. parameces Cossmann & Pissarro (1901).
Schilderina charlesi^[101]	Sp. nov	Valid	Dolin & Aguerre	Miocene		France	A member of the family Cypraeidae.
Schilderina vaesseni^[101]	Sp. nov	Valid	Dolin & Aguerre	Miocene		France	A member of the family Cypraeidae.
Scissurella nesbittae^[102]	Sp. nov	Valid	Geiger & Goedert	Eocene or Oligocene (latest Priabonian or earliest Rupelian)	Gries Ranch Formation	United States ( Washington)	A species of Scissurella.
Scutus mirus^[103]	Sp. nov	Valid	Gard	Late Oligocene to early Miocene		New Zealand	A species of Scutus.
Seguenzia statiana^[104]	Sp. nov	Valid	Sosso, Bertolaso & Dell'Angelo	Miocene (Tortonian)	Marne di S. Agata Fossili Formation	Italy	A species of Seguenzia.
Selvovum^[68]	Gen. et comb. nov	Valid	Juárez-Ruiz, Matamales-Andreu & Kadolsky	Late Eocene		Spain	A member of the family Vidaliellidae; a new genus for "Ampullaria" selvensis.
Semicypraea italorossii^[105]	Sp. nov	Valid	Checchi, Zamberlan & Alberti	Eocene (Ypresian-Lutetian)		Italy	A member of the family Cypraeidae.
Semiricinula preturbinoides^[106]	Sp. nov	Valid	Landau et al.	Miocene (Langhian)		Indonesia	A species of Semiricinula.
Skenea wesselinghi^[107]	Sp. nov	Valid	Hoeksema & Simons	Pliocene		Belgium Netherlands	A species of Skenea.
Skeneopsoides^[91]	Gen. et sp. nov	Valid	Hoeksema, Rijken & Simons	Pliocene		Netherlands	A member of the family Skeneidae. Genus includes new species S. dejongi.
Spiralinella pagoda^[64]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Spiralinella.
Spinatus^[108]	Gen. et 2 sp. et comb. nov	Valid	Dekkers et al.	Miocene		Indonesia	A member of the family Strombidae. The type species is S. echinatus; genus also includes new species S. wonosariensis, as well as "Strombus" tjilonganensis Martin (1899).
Steenbergia wohlmuthorum^[109]	Sp. nov	Valid	Walther & Groh	Late Pleistocene		Madeira	A member of the family Geomitridae.
Stegocoelia samaraensis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Stephaniphera^[110]	Gen. et sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of Cerithioidea belonging to the family Hemisinidae. The type species is S. coronata.
Strebloceras kobayashii^[111]	Sp. nov	Valid	Haga	Middle Pleistocene	Toyohashi Formation	Japan	A member of the family Caecidae.
Streptacis? khaokhadensis^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Streptacis volgenesis^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Striactaenonina elegans^[87]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of Architectibranchia belonging to the family Tubiferidae.
Subtemenia^[110]	Gen. et comb. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Iran Oman	A member of the family Pseudomelaniidae. The type species is "Paryphostoma" morgani Douvillé (1904).
Siphocypraea daughenbaughi^[112]	Sp. nov		Berschauer & Waller		Tamiami Formation	United States ( Florida)	A member of the family Cypraeidae.
Subpterynotus feliciae^[113]	Sp. nov		Petuch & Berschauer			United States ( Florida)	A species of Subpterynotus.
Subpterynotus gabbi^[113]	Sp. nov		Petuch & Berschauer			United States ( Florida)	A species of Subpterynotus.
Teretia horroi^[59]	Sp. nov	Valid	Landau, Van Dingenen & Ceulemans	Miocene (Tortonian)		France	A species of Teretia.
Trachydomia suwanneeae^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Trachyspira eleganta^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Triptychia catalanica^[67]	Sp. nov	Valid	Harzhauser et al.	Late Eocene		Spain
Triptychia tomerdingensis^[114]	Sp. nov	Valid	Kadolsky	Miocene (Aquitanian)		Germany	A member of the family Filholiidae.
Trochomodulus stampinicus^[85]	Sp. nov	Valid	Lozouet, Cahuzac & Charles	Oligocene (Rupelian)		France	A member of the family Modulidae.
Truncatellina dilatatus^[115]	Sp. nov	Valid	Yu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Originally described as a species of Truncatellina; Balashov (2021) transferred it to the genus Euthema.^[116]
Trypanocochlea lopburiensis^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Turbo histrioides^[117]	Sp. nov	Valid	Kase et al.	Miocene		Japan	A species of Turbo.
Turbo izuensis^[117]	Sp. nov	Valid	Kase et al.	Miocene		Japan	A species of Turbo.
Turricarychium^[55]	Gen. et sp. nov	Valid	Kadolsky	Early Eocene		Germany	A member of the family Ellobiidae belonging to the subfamily Carychiinae. The type species is T. muelleri.
Turritelloidea stepheni^[87]	Sp. nov	Valid	Ferrari, Little & Atkinson	Early Jurassic (Toarcian)	Blea Wyke Sandstone Formation	United Kingdom	A member of the family Gordenellidae.
Umbilia darryli^[118]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Umbilia.
Umbilia hallani^[118]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Umbilia.
Vermetujanulus^[109]	Gen. et sp. nov	Valid	Walther & Groh	Late Pleistocene		Madeira	A member of the family Gastrodontidae. Genus includes new species V. boessnecki.
Vetulonia philippinensis^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A species of Vetulonia.
Vexillum pseudoschafferi^[119]	Sp. nov	Valid	Biskupič	Miocene (Serravallian)	Studienka Formation	Slovakia	A species of Vexillum.
Vexillum svagrovskyi^[119]	Sp. nov	Valid	Biskupič	Miocene (Serravallian)	Studienka Formation	Slovakia	A species of Vexillum.
Vicetia bizzottoi^[120]	Sp. nov	Valid	Dominici, Fornasiero & Giusberti	Eocene (Priabonian)	Possagno Marl Formation	Italy	A member of the family Cypraeidae belonging to the subfamily Gisortiinae.
Worthenia humiligrada^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Worthenia morovi^[61]	Sp. nov	Valid	Mazaev	Permian (Sakmarian)		Russia
Xenostoma^[63]	Gen. et sp. nov	Disputed	Bullis et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species X. lophopleura. Originally described as a member of the family Pupinidae; Balashov (2021) considered Xenostoma to be a junior synonym of the diplommatinid genus Euthema, and transferred X. lophopleura to the latter genus.^[116]
Yunnania inflata^[58]	Sp. nov	Valid	Ketwetsuriya et al.	Permian (Roadian)	Khao Khad Formation	Thailand
Zuella^[55]	Gen. et sp. nov	Valid	Kadolsky	Early Eocene		Germany	A member of the family Ellobiidae belonging to the subfamily Carychiinae. The type species is Z. venusta.

Research

A study on the evolutionary history of pteropods, based on molecular data and revised fossil evidence, is published by Peijnenburg et al. (2020).^[121]

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Adelfia^[122]	Gen. et comb. et sp. nov	Valid	Alvarez & Río	Eocene	La Meseta Formation Loreto Formation Submeseta Formation	Antarctica Chile	A member of the family Veneridae. The type species is "Eumarcia" austrolissa Stilwell & Zinsmeister (1992); genus also includes "Venus" arenosa Ortmann (1899), as well as new species A. omega.
Aguileria romualdoensis^[123]	Sp. nov	In press	Rodrigues et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil	A member of the family Bakevelliidae.
Amudariella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Uzbekistan	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" amudariensis Arkhanguelsky (1916); genus also includes "Korobkovitrigonia" subamudariensis Saveliev (1958).
Araripenaia^[125]	Gen. et sp. nov	Valid	Da Silva et al.	Early Cretaceous	Crato Formation	Brazil	An unionid bivalve with possible hyriid affinities. Genus includes new species A. elliptica.
Araripevellia^[123]	Gen. et sp. nov	In press	Rodrigues et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil	A member of the family Bakevelliidae. Genus includes new species A. musculosa.
Arca budoensis^[126]	Sp. nov	Valid	Amano in Amano & Kurita	Early Miocene	Asahi Formation	Japan	A species of Arca.
Archivesica pastori^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A member of the family Vesicomyidae.
Astarte danningeri^[127]	Sp. nov	Valid	Mandic et al.	Early Miocene		Austria	A species of Astarte.
Avichlamys hoferhauseri^[128]	Sp. nov	Valid	Szente	Early Triassic	Werfen Formation	Austria
Bobkovaella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Russia	A member of Trigoniida belonging to the group Myophorelloidea. Genus includes "Trigonia" sydorenkoeremeievae Scheglova-Borodina (1960) and "Korobkovitrigonia" tobolica Pojarkova (1990).
Bogdanovaella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Turkmenistan Uzbekistan	A member of Trigoniida belonging to the group Myophorelloidea. Genus includes "Korobkovitrigonia" conspecta Vinokurova (1970), "Trigonia" darwaseana Romanovsky (1890), "Trigonia" ferganensis Arkhanguelsky (1916), "Korobkovitrigonia" kugitangensis Burkova (1963), "Trigonia" rhombifera Romanovsky (1890) and "Korobkovitrigonia" tagamensis Belyakova (1970).
Cochisella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		United States ( Arizona Texas)	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" guildi Stoyanow (1949); genus also includes "Trigonia" dumblei Stoyanow (1949), "Trigonia" gordoni Whitney (1952), "Trigonia" mearnsi Stoyanow (1949), "Trigonia" resoluta Stoyanow (1949), "Trigonia" saavedra Stoyanow (1949) and "Trigonia" taffi Cragin (1893).
Channelaxinus antipoloensis^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Thyasiridae.
Conchocele majimai^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A member of the family Thyasiridae.
Conchocele visayaensis^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Thyasiridae.
Cratonaia^[129]	Gen. et sp. nov	Valid	Da Silva et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A bivalve belonging to the order Unionida and to the group Silesunionoidea. Genus includes new species C. novaolindensis.
Cristaflabellum^[130]	Gen. et comb. nov	Valid	Hautmann et al.	Triassic	San Cassiano Formation	Bosnia and Herzegovina Italy	A member of the group Aviculopectinoidea. The type species is "Pecten" volaris Bittner (1902); genus also includes "Pecten" interstriatus Münster (1841).
Diplodonta oesia^[131]	Nom. nov	Valid	Pacaud	Eocene (Lutetian)		France	A replacement name for Diplodonta profunda Deshayes (1857)
Dulcina liogliogensis^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A member of the family Lucinidae.
Dulcina magoi^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene	Bata Formation	Philippines	A member of the family Lucinidae.
Elliptiolucina fernandoi^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Late Pliocene to early Pleistocene	Bata Formation	Philippines	A member of the family Lucinidae.
Eothyasira^[132]	Gen. et comb. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany	Genus includes "Venus" antiqua Münster (1841).
Geloina amithoscutana^[110]	Sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of the family Cyrenidae.
Gibbolucina aremorica^[131]	Nom. nov	Valid	Pacaud	Eocene		France	A member of the family Lucinidae; a replacement name for Lucina profunda Dufour (1881)
Globodiscus^[130]	Gen. et 2 sp. nov	Valid	Hautmann et al.	Triassic		Indonesia Romania	A member of the group Aviculopectinoidea. Genus includes new species G. kiliani and G. vinzenti.
Gyrostrea bogdanovae^[133]	Sp. nov	Valid	Kosenko & Metelkin	Early Cretaceous (Hauterivian–Barremian)	Kugusem Formation	Kazakhstan	An oyster.
Illyricocongeria forcakovici^[66]	Sp. nov	Valid	Mandic, Harzhauser & Neubauer	Middle Miocene		Bosnia and Herzegovina
Lima allerdingensis^[127]	Sp. nov	Valid	Mandic et al.	Early Miocene		Austria	A species of Lima.
Limopsis quenstedti^[132]	Nom. nov	Valid	Karapunar et al.				A replacement name for Nucula aurita Quenstedt (1856).
Lucinoma canudai^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Lucinidae.
Lucinoma kosatorea^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Lucinidae.
Lucinoma tinagoensis^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Lucinidae.
Lucinoma velosoi^[72]	Sp. nov	Valid	Kiel, Aguilar & Kase	Early Pleistocene	Bata Formation	Philippines	A member of the family Lucinidae.
Lutetigonia^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		France Tunisia	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" palmata Leymerie (1842); genus also includes "Trigonia" cherahilensis Mongin (1951).
Lyropecten terrysmithae^[134]	Sp. nov	Valid	Powell, Millard & Garcia	Miocene	Monterey Formation Santa Margarita Sandstone	United States ( California)	A scallop.
Macheromenilia^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		France	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" constantii d'Orbigny (1844).
Malletia tsubetsuensis^[135]	Sp. nov	Valid	Matsubara in Matsubara et al.	Miocene (Aquitanian)	Tsubetsu Formation	Japan	A member of the family Malletiidae
Marciachlys^[122]	Gen. et comb. nov	Valid	Alvarez & Río	Eocene	Submeseta Formation	Antarctica	A member of the family Veneridae. The type species is "Eurhomalea" inflata Zinsmeister (1984).
Mezounio^[136]	Gen. et sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia	An unionid bivalve. Genus includes new species M. zverkovi.
Monginellopsis^[125]	Gen. et sp. nov	Valid	Da Silva et al.	Early Cretaceous	Crato Formation	Brazil	A bivalve with possible trigonioidid affinities. Genus includes new species M. bellaradiata.
Monopleura alesta^[137]	Sp. nov	Valid	Masse & Fenerci-Masse	Early Cretaceous (Barremian)		France	A rudist bivalve.
Monopleura jurensis^[137]	Sp. nov	Valid	Masse & Fenerci-Masse	Early Cretaceous (Hauterivian)		France	A rudist bivalve.
Muscatella^[110]	Gen. et sp. nov	Valid	Schneider, Kollmann & Pickford	Late Cretaceous	Al-Khodh Formation	Oman	A member of the family Cyrenidae. The type species is M. biszczukae.
Myonera inouei^[138]	Sp. nov	Valid	Amano in Amano & Kurita	Paleocene (Selandian)	Tomikawa Formation	Japan	A member of the family Cuspidariidae.
Myonia aparicioi^[139]	Sp. nov	Valid	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Nanogyra brevisulcata^[140]	Sp. nov	Valid	Toscano & Lazo	Early Cretaceous	Neuquén Basin	Argentina	A bivalve belonging to the family Gryphaeidae.
Neoquadratotrigonia^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Belgium France United Kingdom	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" quadrata Agassiz (1840); genus also includes "Trigonia daedalea" bracquegniensis Gillet (1920, raised to the rank of the species N. bracquegniensis), "Trigonia daedalea" var. confusa Lycett (1875, raised to the rank of the species N. confusa), "Trigonia" daedalea Parkinson (1811), "Trigonia" rudis Parkinson (1811) and "Trigonia" spectabilis Sowerby (1826).
Nicaniella schoberti^[132]	Sp. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Oesterheldia^[141]	Gen. et sp. nov	Valid	Pérez & Giachetti	Late Oligocene – early Miocene		Argentina	A member of the family Carditidae. The type species is "Venericardia" cannada Ihering (1907); genus also includes "Cyclocardia" dalek Pérez & del Río (2017).
Palaeomutela australis^[142]	Sp. nov	Valid	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the group Actinodontida and the family Palaeomutelidae. Announced in 2019; the final version of the article naming it was published in 2020.
Paracuspidaria^[132]	Gen. et comb. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany	Genus includes "Cypricardia" goldfussiana Kuhn (1935) and "Cypricardia" walteri Kuhn (1935).
Permophorus? yrigoyeni^[139]	Sp. nov	Valid	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Pholadomya miyanoharaensis^[143]	Nom. nov	Valid	Matsubara	Cretaceous		Japan	A replacement name for Pholadomya japonica Amano (1956).
Pratulum kurrawchipellu^[144]	Sp. nov	Valid	Rojas & Nielsen	Early Miocene	Ipún Beds	Chile	A species of Pratulum.
Pratulum terpoorteni^[144]	Sp. nov	Valid	Rojas & Nielsen	Early Miocene	Navidad Formation	Chile	A species of Pratulum.
Promytilus rochacamposi^[139]	Sp. nov	Valid	Díaz Saravia & González	Carboniferous (Pennsylvanian)	Santa Elena Formation	Argentina
Protocardia broilii^[145]	Sp. nov	Valid	Yanin	Early Cretaceous (Berriasian)		Crimean Peninsula
Relogiincola^[142]	Gen. et sp. nov	Valid	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the group Edmondioidea and the family Pachydomidae. Genus includes new species R. delicata. Announced in 2019; the final version of the article naming it was published in 2020.
Retroceramus alliensis^[136]	Sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia
Retroceramus (?) inopinatus^[136]	Sp. nov	Valid	Zakharov in Zakharov et al.	Middle Jurassic (Bajocian)		Russia
Rollieria franconica^[132]	Sp. nov	Valid	Karapunar et al.	Early Jurassic (Pliensbachian)		Germany
Rollieria goldfussi^[132]	Nom. nov	Valid	Karapunar et al.				A replacement name for Nucula subovalis Goldfuss (1837).
Romanovskajaella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous			A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Korobkovitrigonia" mediasiatica Belyakova (1970); genus also includes "Korobkovitrigonia" akkaptschigensis Belyakova (1970), "Korobkovitrigonia" bobkovae Belyakova (1970), "Korobkovitrigonia" gissarensis Belyakova (1970) and "Korobkovitrigonia" rugosa Belyakova (1970).
Romanovskiella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Uzbekistan	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Trigonia" romanovskii Arkhanguelsky (1916); genus also includes "Trigonia" aralensis Arkhangelsky (1912).
Saveligonia^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Russia Turkmenistan	A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Litschkovitrigonia" media Saveliev (1958); genus also includes "Litschkovitrigonia inguschensis" alta Prozorovskii (1961, raised to the rank of the species S. alta), "Trigonia" inguschensis Renngarten (1926), "Trigonia" subdaedalea Renngarten (1926) and "Litschkovitrigonia" tenuituberculata Saveliev (1958).
Shajia^[146]	Gen. et sp. nov	Valid	Rao et al.	Early Cretaceous (Aptian or Albian)	Langshan Formation	China India	A rudist bivalve. Genus includes new species S. tibetica.
Terraia decarinata^[142]	Sp. nov	Valid	Guerrini et al.	Permian	Rio do Rasto Formation	Brazil	A bivalve belonging to the family Astartidae. Announced in 2019; the final version of the article naming it was published in 2020.
Trigonipraxis madjerensis^[98]	Sp. nov	Valid	Neubauer et al.	Miocene		Serbia	A member of the family Dreissenidae belonging to the subfamily Congeriinae.
Vinokurovaella^[124]	Gen. et comb. nov	Valid	Cooper	Cretaceous			A member of Trigoniida belonging to the group Myophorelloidea. The type species is "Korobkovitrigonia" grumosa Vinokurova (1970); genus also includes "Quadratotrigonia" turkmenica Vinokurova (1970).

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acanthochitona interglacialis^[147]	Sp. nov	Valid	Dell'Angelo et al.	Late Pleistocene		Saudi Arabia	A chiton, a species of Acanthochitona.
Apoptopegma joyceae^[148]	Sp. nov	Valid	Smith, Brock & Paterson	Cambrian (Guzhangian)	Shannon Formation	Australia	A rostroconch.
Catalanispira^[149]	Gen. et sp. et comb. nov	Valid	Ebbestad, Kolata & Isakar	Ordovician (Darriwilian and Sandbian)	Kõrgekallas Platteville	Estonia United States ( Illinois)	A member of the family Onychochilidae. The type species is C. plattevillensis; genus also includes "Clisospira" reinwaldti Öpik (1930).
Craspedochiton fontlevensis^[150]	Sp. nov	Valid	Dell'Angelo et al.	Miocene (Burdigalian)		France	A chiton, a species of Craspedochiton.
Craspedochiton lozoueti^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Craspedochiton.
Crassaplax^[151]	Gen. et sp. nov	Valid	Högström, Ebbestad & Suzuki	Ordovician (Katian)	Boda Limestone	Sweden	A member of Paleoloricata. The type species is C. collicola.
Lepidochitona chalossensis^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lepidochitona charlesi^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lepidochitona larratensis^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lepidochitona tarbelliana^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lepidochitona tessellata^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lepidochitona vancuycki^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Lepidochitona.
Lucilina aqabaensis^[147]	Sp. nov	Valid	Dell'Angelo et al.	Late Pleistocene		Saudi Arabia	A chiton, a species of Lucilina.
Qaleruaqia^[152]	Gen. et sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A member of the family Mattheviidae. The type species is Q. sodermanorum.
Pseudoacanthochitona marsanensis^[150]	Sp. nov	Valid	Dell'Angelo et al.	Miocene (Burdigalian)		France	A chiton.
Stenosemus radiatus^[150]	Sp. nov	Valid	Dell'Angelo et al.	Miocene (Burdigalian)		France	A chiton.
Tonicella adunca^[150]	Sp. nov	Valid	Dell'Angelo et al.	Miocene (Aquitanian)		France	A chiton, a species of Tonicella.
Tonicella nuda^[150]	Sp. nov	Valid	Dell'Angelo et al.	Oligocene (Rupelian)		France	A chiton, a species of Tonicella.

2020 in paleomalacology

New taxa

General research

Other cephalopods

New taxa

General research

Gastropods

New taxa

Research

Bivalves

New taxa

Other mollusks

New taxa

Research

References

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