2018 in paleontology

From Wikipedia, the free encyclopedia

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2018.

Extinct animals named in 2018

Plants

Fungi

Name Novelty Status Authors Age Type locality Country Notes Images

Chaenotheca succina[2]

Sp. nov

Valid

Rikkinen & Schmidt in Rikkinen et al.

Eocene (Priabonian)

Baltic amber

Russia
( Kaliningrad Oblast)

A fungus, a species of Chaenotheca.

Notothyrites (?) leptostrobi[3]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Early and Middle Jurassic

Prisayanskaya Formation

Russia

A member of the family Microthyriaceae.

Palaeomycus[4]

Gen. et sp. nov

Valid

Poinar

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A fungus described on the basis of pycnidia. Genus includes new species P. epallelus. Announced in 2018; the final version of the article naming it was published in 2020.

Paleoambrosia[5]

Gen. et sp. nov

Valid

Poinar & Vega

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

An ambrosia fungus associated with the beetle Palaeotylus femoralis.
Genus includes new species P. entomophila.

Perexiflasca[6]

Gen. et sp. nov

Valid

Krings, Harper & Taylor

Devonian (Pragian)

Rhynie chert

United Kingdom

A small, chytrid-like organism. Genus includes new species P. tayloriana.

Phyllopsora magna[7]

Sp. nov

Valid

Kaasalainen, Rikkinen & Schmidt in Kaasalainen et al.

Miocene

Dominican amber

Dominican Republic

A lichenized fungus, a species of Phyllopsora.

Retesporangicus[8]

Gen. et sp. nov

Valid

Strullu-Derrien in Strullu-Derrien et al.

Early Devonian

Rhynie chert

United Kingdom

A fungus belonging to the group Blastocladiomycota, of uncertain phylogenetic placement within the latter group. Genus includes new species R. lyonii.

Vizellopsidites[9]

Gen. et sp. nov

Valid

Khan, Bera & Bera

Late Pliocene to early Pleistocene

Kimin Formation

India

A fossil fungus found on the surface of fossilized leaf fragments. Genus includes new species V. siwalika.

Windipila pumila[10]

Sp. nov

Valid

Krings & Harper

Early Devonian

Rhynie chert

United Kingdom

A fungal reproductive unit.

Cnidarians

Research

New taxa

Name Novelty Status Authors Age Type locality Country Notes

Acropora incogtita[14]

Sp. nov

Valid

Berezovsky & Satanovska

Eocene

Ukraine

A stony coral, a species of Acropora.

Actinoseris riyadhensis[15]

Sp. nov

Valid

Gameil, El-Sorogy & Al-Kahtany

Late Cretaceous (Campanian)

Aruma Formation

Saudi Arabia

A solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.

Antheria fedorowskii[16]

Sp. nov

Valid

Wang, Gorgij & Yao

Late Carboniferous

Iran

A rugose coral.

Antheria robusta[16]

Sp. nov

Valid

Wang, Gorgij & Yao

Late Carboniferous

Iran

A rugose coral.

Asteroseris arabica[15]

Sp. nov

Valid

Gameil, El-Sorogy & Al-Kahtany

Late Cretaceous (Campanian)

Aruma Formation

Saudi Arabia

A solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.

Astraraeatrochus[17]

Gen. et sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Haplaraeoidea and the family Astraraeidae. The type species is A. bachi.

Astreoidogyra[18]

Gen. et sp. nov

Valid

Ricci, Lathuilière & Rusciadelli

Late Jurassic

Italy

A member of the family Rhipidogyridae. The type species is A. giadae.

Aulocystis wendti[19]

Sp. nov

Valid

Król, Zapalski & Berkowski

Devonian (Emsian)

Amerboh Group

Morocco

A tabulate coral belonging to the family Aulocystidae.

Bainbridgia bipartita[19]

Sp. nov

Valid

Król, Zapalski & Berkowski

Devonian (Emsian)

Kess-Kess Formation

Morocco

A tabulate coral belonging to the family Pyrgiidae.

Battersbyia coactilis[20]

Sp. nov

Valid

McLean

Devonian

Canada

A rugose coral.

Battersbyia sentosa[20]

Sp. nov

Valid

McLean

Devonian

Canada

A rugose coral.

Cambrorhytium gracilis[21]

Sp. nov

Valid

Chang et al.

Early Cambrian

China

Caryophyllia (Caryophyllia) imamurai[22]

Sp. nov

Valid

Niko

Miocene

Bihoku Group

Japan

A species of Caryophyllia.

Catenipora jingyangensis[23]

Sp. nov

Valid

Liang, Elias & Lee

Ordovician (Katian)

Beiguoshan Formation

China

A tabulate coral.

Catenipora tiewadianensis[23]

Sp. nov

Valid

Liang, Elias & Lee

Ordovician (Katian)

Beiguoshan Formation

China

A tabulate coral.

Catenipora tongchuanensis[23]

Sp. nov

Valid

Liang, Elias & Lee

Ordovician (Sandbian)

Jinghe Formation

China

A tabulate coral.

Clausastrea eliasovae[18]

Sp. nov

Valid

Ricci, Lathuilière & Rusciadelli

Late Jurassic

Italy

A member of the family Montlivaltiidae.

Crinopora ireneae[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Heterocoenioidea and the family Carolastraeidae.

Crinopora thomasi[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Heterocoenioidea and the family Carolastraeidae.

Cunnolites (Plesiocunnolites) riyadhensis[15]

Sp. nov

Valid

Gameil, El-Sorogy & Al-Kahtany

Late Cretaceous (Campanian)

Aruma Formation

Saudi Arabia

A solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.

Deltocyathoides bihokuensis[22]

Sp. nov

Valid

Niko

Miocene

Bihoku Group

Japan

A stony coral.

Fuchungopora huilongensis[24]

Sp. nov

Valid

Liang et al.

Devonian (Famennian)

Etoucun Formation

China

A syringoporoid tabulate coral.

Geroastrea[17]

Gen. et sp. et comb. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria
France
Iran

A stony coral belonging to the superfamily Cyclolitoidea and the family Synastraeidae. The type species is G. alexi; genus also includes G. audiensis (Reig Oriol, 1992), G. haueri (Reuss, 1854) and G. parvistella (Oppenheim, 1930).

Gosaviaraea aimeae[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral.

Kozaniastrea[25]

Gen. et sp. nov

Valid

Löser, Steuber & Löser

Late Cretaceous (Cenomanian)

Greece

A stony coral belonging to the superfamily Felixaraeoidea and the family Lamellofungiidae. The type species is K. pachysepta.

Lithophyllon comptus[26]

Sp. nov

Valid

Berezovsky & Satanovska

Eocene

Ukraine

A stony coral, a species of Lithophyllon.

Lonsdaleia carnica[27]

Sp. nov

Valid

Rodríguez, Schönlaub & Kabon

Carboniferous (Mississippian)

Kirchbach Formation

Austria

A rugose coral belonging to the family Axophyllidae.

Lyrielasma landryense[20]

Sp. nov

Valid

McLean

Devonian

Canada

A rugose coral.

Nefocoenia seewaldi[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Phyllosmilioidea and the family Phyllosmiliidae.

Nefocoenia werneri[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Phyllosmilioidea and the family Phyllosmiliidae.

Neopilophyllia[28]

Gen. et comb. nov

Valid

Wang in Wang et al.

Silurian (Telychian)

Ningqiang Formation

China

A rugose coral belonging to the new family Amplexoididae. The type species is "Ningqiangophyllum" crassothecatum Cao (1975); genus also includes "Ningqiangophyllum" tenuiseptatum irregulare Cao (1975) (raised to the rank of a separate species Neopilophyllia irregularis), "Ningqiangophyllum" ephippium Cao (1975) and "Pilophyllia" alternata Chen in Wang et al. (1986).

Oculina complanatis[29]

Sp. nov

Valid

Berezovsky & Satanovska

Eocene

Ukraine

A stony coral, a species of Oculina.

Opolestraea[30]

Gen. et comb. nov

Valid

Morycowa

Middle Triassic (Anisian)

Karchowice Beds

Poland

A stony coral belonging to the family Eckastraeidae. The type species is "Coelocoenia" exporrecta Weissermel (1925).

Pachyheterocoenia[17]

Gen. et sp. et comb. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria
Spain

A stony coral belonging to the superfamily Heterocoenioidea and the family Heterocoeniidae. The type species is P. leipnerae; genus also includes P. grandis (Reuss, 1854) and P. fuchsi (Felix, 1903).

Pachyphylliopsis[17]

Gen. et sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria
Iran
United Arab Emirates

A stony coral belonging to the superfamily Phyllosmilioidea and the family Phyllosmiliidae. The type species is P. magnum.

Paractinacis[17]

Gen. et sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria
Germany
Spain

A stony coral belonging to the superfamily Cyclolitoidea and the family Negoporitidae. The type species is P. uliae; genus might also include P. ? elegans (Reuss, 1854).

Plesiolites[25]

Gen. et sp. nov

Valid

Löser, Steuber & Löser

Late Cretaceous (Cenomanian)

Greece

A stony coral belonging to the superfamily Misistelloidea. The type species is P. winnii.

Proplesiastraea rivkae[17]

Sp. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Cladocoroidea and the family Columastraeidae.

Psydracophyllum hinnuleum[20]

Sp. nov

Valid

McLean

Devonian

Canada

A rugose coral.

Striatopora marsupia[19]

Sp. nov

Valid

Król, Zapalski & Berkowski

Devonian (Emsian)

Amerboh Group

Morocco

A tabulate coral belonging to the family Pachyporidae.

Styloheterocoenia[25]

Gen. et 2 sp. nov

Valid

Löser, Steuber & Löser

Late Cretaceous (Cenomanian)

Greece

A stony coral belonging to the superfamily Heterocoenioidea and the family Heterocoeniidae. The type species is S. hellenensis; genus also includes S. brunni.

Stylophora kibiensis[31]

Sp. nov

Valid

Niko, Suzuki & Taguchi

Miocene

Katsuta Group

Japan

A species of Stylophora.

Sutherlandia jamalensis[32]

Sp. nov

Valid

Niko et al.

Early Permian

Jamal Formation

Iran

A tabulate coral belonging to the order Favositida and the family Favositidae.

Synhydnophora[17]

Gen. et sp. et comb. nov

Valid

Löser & Heinrich

Late Cretaceous

Austria

A stony coral belonging to the superfamily Cyclolitoidea and the family Synastraeidae. The type species is S. wagreichi; genus also includes and S. multilamellosa (Reuss, 1854).

Wendticyathus[33]

Gen. et sp. nov

Valid

Berkowski

Devonian (Emsian)

Morocco

A rugose coral. Genus includes new species W. nudus.

Xystriphylloides distinctus[34]

Sp. nov

Valid

Yu

Early Devonian

China

A rugose coral.

Xystriphyllum helenense[20]

Sp. nov

Valid

McLean

Devonian

Canada

A rugose coral.

Arthropods

Bryozoans

New taxa

Name Novelty Status Authors Age Type

locality

Country Notes

Acanthodesia variegata[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Membraniporidae.

Calyptotheca sidneyi[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Bitectiporidae.

Characodoma wesselinghi[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Cleidochasmatidae.

Cystomeson[36]

Gen. nov

Valid

Ernst, Krainer and Lucas

Mississippian

Lake Valley Formation

United States

A cystoporate bryozoan of the family Fistuliporidae.

Pleurocodonellina javanensis[35]

Sp. nov

Valid

Di Martino & Taylor

Early Pleistocene

Pucangan Formation

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Smittinidae.

Turbicellepora yasuharai[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Celleporidae.

Brachiopods

Research

  • Studies on the ontogenetic development of early acrotretoid brachiopods based on well preserved specimens of the earliest Cambrian species Eohadrotreta zhenbaensis and Eohadrotreta? zhujiahensis from the Shuijingtuo Formation (China) are published by Zhang et al. (2018).[37][38]
  • A study on the extinction and origination of members of the order Strophomenida during the Late Ordovician mass extinction is published by Sclafani et al. (2018).[39]
  • A study on the body size of several brachiopod assemblages recorded into the extinction interval prior to the Toarcian turnover, collected from representative localities around the Iberian Massif (Spain and Portugal), is published by García Joral, Baeza-Carratalá & Goy (2018).[40]

New taxa

Name Novelty Status Authors Age Type locality Country Notes

Acrotreta calabozoi[41]

Sp. nov

Valid

Lavié

Ordovician (Sandbian)

Las Plantas Formation

Argentina

Adygella socotrana[42]

Sp. nov

Valid

Gaetani in Gaetani et al.

Middle Triassic

Yemen

A member of Terebratulida belonging to the family Dielasmatidae.

Ahtiella famatiniana[43]

Sp. nov

Valid

Benedetto

Ordovician

Argentina

Ahtiella tunaensis[43]

Sp. nov

Valid

Benedetto

Ordovician

Argentina

Ala alatiformis[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Choristitidae.

Alebusirhynchia vorosi[45]

Sp. nov

Valid

Baeza-Carratalá, Dulai & Sandoval

Early Jurassic

Spain

A member of Rhynchonellida.

Alekseevathyris[46]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Givetian)

Coronados Volcanics

United States
( Alaska)

A member of Terebratulida belonging to the family Stringocephalidae. The type species is A. coronadosensis.

Altaethyrella tarimensis[47]

Sp. nov

Valid

Sproat & Zhan

Ordovician (late Katian)

Hadabulaktag Formation

China

Ambocoelia yidadeensis[48]

Sp. nov

Valid

Zhang & Ma

Devonian (Frasnian)

Yidade Formation

China

Arpaspirifer[49][50]

Gen. et comb. nov

Valid

Gretchishnikova in Alekseeva et al.

Devonian (Famennian)

Armenia
Azerbaijan

A member of the family Cyrtosririferidae. The type species is "Spirifer" latus Abrahamian (1974).

Aulacella finitima[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (EifelianGivetian)

Azerbaijan

Betaneospirifer stepanovi[51][52]

Sp. nov

Valid

Poletaev

Carboniferous

Russia
( Bashkortostan)

A member of the family Trigonotretidae.

Biernatium sucoi[53]

Sp. nov

Valid

García-Alcalde

Devonian (Givetian)

Portilla Formation

Spain

A member of Orthida belonging to the family Mystrophoridae.

Broggeria omaguaca[54]

Sp. nov

Valid

Benedetto, Lavie & Muñoz

Ordovician (Tremadocian)

Argentina

Buxtonia sulcata[55]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Buxtoniidae.

Callaiapsida divitiae[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Rhynchonellida belonging to the family Stenoscismatidae.

Calliprotonia kerrae[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Echinoconchidae.

Calliprotonia umbonalis[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Echinoconchidae.

Chelononia minimauris[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Strophomenata belonging to the superfamily Orthotetoidea and the family Schuchertellidae.

Churkinella[46]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Givetian)

Coronados Volcanics

United States
( Alaska)

A member of Terebratulida belonging to the family Stringocephalidae. The type species is C. craigensis.

Cingulodermis pustulatus[56]

Sp. nov

Valid

Mergl

Devonian (Emsian)

Morocco

Commarginalia norrisi[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Proboscidelloidea and the family Paucispinauriidae.

Composita largitas[57]

Sp. nov

Valid

Waterhouse

Carboniferous

Wahoo Formation

Canada
( Yukon)

Coronadothyris[46]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Givetian)

Coronados Volcanics

United States
( Alaska)

A member of Terebratulida belonging to the family Stringocephalidae. The type species is C. mica.

Costisorthis lisae[53]

Sp. nov

Valid

García-Alcalde

Devonian (Givetian)

Candás Formation

Spain

A member of Orthida belonging to the family Dalmanellidae.

Cyrtiorina houi[58]

Sp. nov

Valid

Zong & Ma

Devonian (Famennian)

Hongguleleng Formation

China

A brachiopod belonging to the group Spiriferida.

Cyrtospirifer dansikensis[49][50]

Sp. nov

Valid

Afanasjeva in Alekseeva et al.

Devonian (Famennian)

Azerbaijan

Dalejina aulacelliformis[56]

Sp. nov

Valid

Mergl

Devonian (Emsian)

Morocco

Datnella[59]

Gen. et comb. nov

Valid

Baranov

Early Devonian

Russia

A member of Atrypida. The type species is D. datnensis (Baranov, 1995).

Deltachania elongata[55]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Athyrida belonging to the superfamily Athyroidea.

Desquamatia globosa jozefkae[60]

Subsp. nov

Valid

Baliński in Skompski et al.

Devonian (GivetianFrasnian boundary)

Szydłówek Beds

Poland

A member of Atrypida belonging to the family Atrypidae.

Diazoma ghyumuschlugensis[49][50]

Sp. nov

Valid

Oleneva in Alekseeva et al.

Devonian (Frasnian)

Azerbaijan

Dichospirifer felixi[49][50]

Sp. nov

Valid

Gretchishnikova in Alekseeva et al.

Devonian (Famennian)

Azerbaijan

Dutroproductus[44]

Gen. et sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Retariidae. The type species is D. dutroi.

Echinalosia minuta[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Strophalosioidea and the family Dasyalosiidae.

Echinaria circularis[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Echinoconchidae.

Eopholidostrophia (Megapholidostrophia) gigas[61]

Sp. nov

Valid

Strusz & Percival

Silurian (Wenlock)

Australia

Eressella[62]

Gen. et comb. nov

Valid

Halamski & Baliński

Middle Devonian

Germany
Morocco
Poland

A member of Rhynchonellida belonging to the family Uncinulidae. The type species is "Rhynchonella" coronata Kayser (1871).

Eridmatina[44]

Gen. et comb. nov

Valid

Waterhouse

Carboniferous and Permian

Gaptank Formation

Canada
( Yukon)
United States

A member of Spiriferida belonging to the family Spiriferellidae. The type species is "Eridmatus" marathonensis Cooper & Grant (1976); genus also includes "Eridmatus" petita Waterhouse & Waddington (1982)

Ettrainia[44]

Gen. et sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Choristitoidea and the family Palaeochoristitidae. The type species is E. costellata.

Flexaria echinata[63]

Sp. nov

Valid

Waterhouse

Carboniferous

Hart River Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Buxtoniidae.

Forticosta[44]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Spiriferoidea and the family Neospiriferidae. The type species is F. transversa.

Gemmulicosta undulata[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Buxtoniidae.

Gypidulina grandis[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (EifelianGivetian)

Azerbaijan

Harkeria elongata[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Yakovleviidae.

Harkeria sulcoprofundus[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Yakovleviidae.

Hartea[63]

Gen. et sp. nov

Junior homonym

Waterhouse

Carboniferous

Hart River Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Ambocoeliidae. The type species is H. venustus. The generic name is preoccupied by Hartea Wright (1865).

Heella[44]

Gen. et comb. nov

Valid

Waterhouse

Permian

China

A member of Spiriferida belonging to the family Ambocoeliidae. The type species is "Attenuatella" mengi He, Shi, Feng & Peng (2007)

Heteralosia scotti[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Strophalosioidea and the family Strophalosiidae.

Hustedia quadrifidus[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Rhynchonellata belonging to the group Retziida and the family Retziidae.

Hustedia trifida[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Rhynchonellata belonging to the group Retziida and the family Retziidae.

Isorthis (Arcualla) delegatensis[61]

Sp. nov

Valid

Strusz & Percival

Silurian (Wenlock)

Australia

Jagtithyris[64]

Gen. et comb. nov

Valid

Simon & Mottequin

Late Cretaceous (Maastrichtian)

Netherlands

A relative of Leptothyrellopsis, assigned to the new family Jagtithyrididae. Genus includes "Terebratella (Morrisia?)" suessi Bosquet (1859).

Junglelomia simplex[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Choristitoidea and the family Palaeochoristitidae.

Juxathyris subcircularis[65]

Sp. nov

Valid

Wu et al.

Permian (Changhsingian)

Changxing Formation

China

A member of Athyridida.

Komiella bitteri[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of the family Rugosochonetidae.

Krotovia norfordi[44]

Sp. nov

Valid

Waterhouse

Carboniferous and Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Echinoconchidae.

Kukulkanus[66]

Gen. et sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (ArtinskianKungurian)

Paso Hondo Formation

Mexico

A brachiopod belonging to the group Productida and the family Productidae. The type species is K. spinosus.

"Kutorginella" primigenius[63]

Sp. nov

Valid

Waterhouse

Carboniferous

Hart River Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Retariidae.

Latisulcus[44]

Gen. et comb. nov

Valid

Waterhouse

Permian

Bone Spring Formation

United States
( Texas)

A member of Rhynchonellata belonging to the group Retziida and the family Retziidae. The type species is "Hustedia" hessensis King (1931)

Leiochonetes onimarensis[67]

Sp. nov

Valid

Tazawa

Carboniferous (Mississippian)

Hikoroichi Formation

Japan

A member of the family Rugosochonetidae belonging to the subfamily Svalbardiinae.

Leptaena (Leptaena) australis[61]

Sp. nov

Valid

Strusz & Percival

Silurian (Wenlock)

Australia

Leurosina katasumiensis[68]

Sp. nov

Valid

Afanasjeva, Jun-Ichi & Yukio

Permian (Kungurian)

Nabeyama Formation

Japan

A member of Chonetida belonging to the family Rugosochonetidae.

Levipustula canadensis[55]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Strophalosiidina belonging to the superfamily Scacchinelloidea and the family Levipustulidae.

Martinezchaconia[69]

Gen. et sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (Bashkirian-Moscovian)

Ixtaltepec Formation

Mexico

A member of Productida belonging to the family Linoproductidae. The type species is M. luisae.

Meristorygma donakovae[51][52]

Sp. nov

Valid

Poletaev

Carboniferous

Kizil Formation

Russia

A member of the family Brachythyrididae.

Mirandifera[44]

Gen. et comb. nov

Valid

Waterhouse

Permian

Cathedral Mountain Formation

Canada
( Yukon)
United States
( Texas)

A member of Spiriferida belonging to the family Martiniidae. The type species is "Martinia" miranda Cooper & Grant (1976); genus also includes "Martinia" wolfcampensis King (1931)

Misunithyris[70]

Gen. et sp. nov

Valid

Baeza-Carratalá, Pérez-Valera & Pérez-Valera

Middle Triassic (Ladinian)

Siles Formation

Spain

A brachiopod belonging to the group Terebratellidina and to the superfamily Zeillerioidea. The type species is M. goyi.

Morinorhynchus tucksoni[61]

Sp. nov

Valid

Strusz & Percival

Silurian (Wenlock)

Australia

Muirwoodiciana[55]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Productida belonging to the family Yakovleviidae. The type species is M. inexpectans.

Musalitinispira[59]

Gen. et sp. nov

Valid

Baranov

Early Devonian

Russia

A member of Atrypida. The type species is M. dogdensis.

Mysteronia[55]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Rhynchonellida belonging to the superfamily Rhynchoporoidea and the family Rhynchoporidae. The type species is M. mysticus.

Nahoniella decorus[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferinida belonging to the group Syringothyridina and the family Licharewiidae.

Nassichukia[44]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Buxtoniidae. The type species is N. nodosa.

Nazeriproductus lazarevi[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Productoidea and the family Retariidae.

Neochonetes (Huangichonetes) matsukawensis[71]

Sp. nov

Valid

Tazawa & Araki

Permian (Wordian)

Kamiyasse Formation

Japan

A member of the family Rugosochonetidae.

Newberria alaskensis[46]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Givetian)

Coronados Volcanics

United States
( Alaska)

A member of Terebratulida belonging to the family Stringocephalidae.

Nucleospira quidongensis[61]

Sp. nov

Valid

Strusz & Percival

Silurian (Wenlock)

Australia

Ogilviecoelia initiatus[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Ambocoeliidae.

Ogilviecoelia shii[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Ambocoeliidae.

Opsiconidion bouceki[72]

Sp. nov

Valid

Mergl, Frýda & Kubajko

Silurian (Ludfordian)

Kopanina Formation

Czech Republic

A member of Acrotretoidea belonging to the family Biernatidae.

Opsiconidion parephemerus[72]

Sp. nov

Valid

Mergl, Frýda & Kubajko

Silurian (Ludfordian)

Kopanina Formation

Czech Republic

A member of Acrotretoidea belonging to the family Biernatidae.

Orthotetes dorsosulcata[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Strophomenata belonging to the superfamily Orthotetoidea and the family Orthotetidae.

Papulifera[55]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Martiniidae. The type species is P. plana.

Paucispinifera abramovi[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Paucispiniferidae.

Paucispinifera carboniferica[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Paucispiniferidae.

Paucispinifera sulcata[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Paucispiniferidae.

Pinguispirifer kesskess[56]

Sp. nov

Valid

Mergl

Devonian (Emsian)

Morocco

Piridiorhynchus jafariani[73]

Sp. nov

Valid

Baranov et al.

Devonian (Famennian)

Khoshyeilagh Formation

Iran

A member of Rhynchonellida belonging to the family Trigonirhynchiidae.

Plicatospiriferella undulata[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the family Spiriferellidae.

Poletaevia[44]

Gen. et comb. nov

Valid

Waterhouse

Carboniferous

Hare Fiord Formation

Canada
( Nunavut)

A member of Productida belonging to the superfamily Paucispiniferoidea and the family Anidanthidae. The type species is "Liraria" paucispina Carter & Poletaev (1998)

Pripyatispirifer caucasius[49][50]

Sp. nov

Valid

Afanasjeva in Alekseeva et al.

Devonian (Frasnian)

Azerbaijan

Protoanidanthus monstratus[57]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Paucispiniferoidea and the family Anidanthidae.

Protoanidanthus nichollsi[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Paucispiniferoidea and the family Anidanthidae.

Pumilusia[44]

Gen. et comb. nov

Valid

Waterhouse

Carboniferous

La Prasada Formation

United States
( New Mexico)

A member of Productida belonging to the superfamily Linoproductoidea and the family Ovatiidae. The type species is "Linoproductus" pumilus Sutherland & Harlow (1973)

Punctospirifer iwatensis[67]

Sp. nov

Valid

Tazawa

Carboniferous (Mississippian)

Hikoroichi Formation

Japan

A member of Spiriferinida belonging to the family Punctospiriferidae.

Resserella dagnensis[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (EmsianEifelian)

Azerbaijan

Reticulariopsis rotunda[49][50]

Sp. nov

Valid

Oleneva in Alekseeva et al.

Devonian (Givetian)

Azerbaijan

Rhipidomella arpensis[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (Givetian)

Azerbaijan

Rhipidomella borealis[57]

Sp. nov

Valid

Waterhouse

Carboniferous

Wahoo Formation

Canada
( Yukon)

A member of Orthida belonging to the family Rhipidomellidae.

Rhynchopora grigorievae[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Rhynchonellida belonging to the superfamily Rhynchoporoidea and the family Rhynchoporidae.

Rhynchoporusia[63]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Hart River Formation

Canada
( Yukon)

A member of Rhynchonellida belonging to the superfamily Rhynchoporoidea and the family Rhynchoporidae. The type species is R. multiplicata.

Rorespirifer prodigium[57]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Ingelarelloidea and the family Rorespiriferidae.

Rugaria arcula[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of the family Rugosochonetidae.

Rugivestigia[44]

Gen. et sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Paucispiniferidae. The type species is R. commarginalis.

Rugosochonetes multistriatus[49][50]

Sp. nov

Valid

Afanasjeva in Alekseeva et al.

Carboniferous (Tournaisian)

Azerbaijan

Saltospirifer gibberosus[55]

Sp. nov

Valid

Waterhouse

Carboniferous

Blackie Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Spiriferoidea and the family Spiriferidae.

Sangredonia alaminata[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Horridoniidae.

Sarytchevinella praecursor[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Linoproductoidea and the family Striatiferidae.

Schizambon langei[74]

Sp. nov

Valid

Freeman, Miller & Dattilo

Cambrian–Ordovician boundary

United States
( Texas)

A linguliform brachiopod.

Schizophoria lata[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (EmsianEifelian)

Azerbaijan

Schizophoria schnuri altera[49][50]

Subsp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (Givetian)

Azerbaijan

Septatrypa tumulorum[75]

Sp. nov

Valid

Baliński & Halamski

Devonian (Emsian)

Morocco

Septospirifer hughi[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Spiriferoidea and the family Neospiriferidae.

Sieberella parva[49][50]

Sp. nov

Valid

Alekseeva & Gretchishnikova in Alekseeva et al.

Devonian (EmsianEifelian)

Azerbaijan

Sphenospira dansikensis[49][50]

Sp. nov

Valid

Gretchishnikova in Alekseeva et al.

Devonian (Famennian)

Azerbaijan

Spinatrypina (Spinatrypina) krivensis[59]

Sp. nov

Valid

Baranov

Early Devonian

Russia

A member of Atrypida.

Spinocyrtia irinae[49][50]

Sp. nov

Valid

Afanasjeva in Alekseeva et al.

Devonian (Eifelian and Givetian)

Azerbaijan

Spirelytha biakovi[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Elitoidea and the family Toryniferidae.

Spiriferinaella simplicata[44]

Sp. nov

Valid

Waterhouse

Carboniferous and Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Paeckelmanelloidea and the family Pterospiriferidae.

Stataria[44]

Gen. et comb. nov

Valid

Waterhouse

Permian

Cathedral Mountain Formation

United States
( Texas)

A member of Rhynchonellata belonging to the group Retziida and the family Retziidae. The type species is "Hustedia" stataria Cooper & Grant (1976)

Stenorhynchia ulrici[75]

Sp. nov

Valid

Halamski & Baliński

Devonian (Emsian)

Morocco

Tegulispirifer? placitus[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferida belonging to the superfamily Spiriferoidea and the family Spiriferidae.

Tethysiella impudens[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Paucispiniferidae.

Thomasaria caucasica[49][50]

Sp. nov

Valid

Oleneva in Alekseeva et al.

Devonian (Eifelian)

Azerbaijan

Trigonatrypa drotae[56]

Sp. nov

Valid

Mergl

Devonian (Emsian)

Morocco

Tuberculatella bunnakia[44]

Nom. nov

Valid

Waterhouse

Carboniferous

Thailand

A member of Productida belonging to the family Avoniidae; a replacement name for Tuberculatella tuberculata Waterhouse (1982).

Tubersulculus ovalis[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the family Echinoconchidae.

Tumarinia solominae[44]

Sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Spiriferinida belonging to the group Syringothyridina and the family Licharewiidae.

Undispirifer dansikensis[49][50]

Sp. nov

Valid

Oleneva in Alekseeva et al.

Devonian (Eifelian)

Azerbaijan

Unispirifer arpensis[49][50]

Sp. nov

Valid

Afanasjeva in Alekseeva et al.

Carboniferous (Tournaisian)

Azerbaijan

Villaconcha planiconcha[44]

Sp. nov

Valid

Waterhouse

Permian

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Echinoconchoidea and the family Waagenoconchidae.

Yanzaria[44]

Gen. et comb. et sp. nov

Valid

Waterhouse

Carboniferous and Permian

Fenestella Shales

Canada
( Yukon)
India

A member of Rhynchonellida belonging to the family Tetracameridae. The type species is "Camarophoria" dowhatensis Diener (1915); genus also includes new species Y. solitarius.

Yukonalosia[44]

Gen. et sp. nov

Valid

Waterhouse

Carboniferous

Jungle Creek Formation

Canada
( Yukon)

A member of Productida belonging to the superfamily Strophalosioidea and the family Dasyalosiidae. The type species is Y. arctica.

Zaigunrostrum nakhichevanense[76]

Sp. nov

Valid

Pakhnevich

Devonian (Famennian)

Azerbaijan

A brachiopod belonging to the group Rhynchonellida and the family Trigonirhynchiidae.

Zezinia[76]

Gen. et sp. nov

Valid

Pakhnevich

Devonian (Frasnian)

Azerbaijan

A brachiopod belonging to the group Rhynchonellida and the family Uncinulidae. The type species is Z. multicostata.

Molluscs

Echinoderms

Conodonts

Research

  • A study testing the proposed models of growth of conodont elements is published by Shirley et al. (2018).[77]
  • A study on the histological sections of Ordovician and Permian conodont dental elements from the Bell Canyon Formation (Texas, United States), Harding Sandstone (Colorado, United States), Ali Bashi Formation (Iran) and Canadian Arctic, examining those fossils for the presence and distribution of soft tissue biomarkers, is published by Terrill, Henderson & Anderson (2018).[78]
  • A study evaluating the δ18O variation within a species-rich conodont assemblage from the Ordovician (Floian) Factory Cove Member of the Shallow Bay Formation, Cow Head Group (western Newfoundland, Canada), as well as assessing the implications of these data for determining the paleothermometry of ancient oceans and conodont ecologic models, is published by Wheeley et al. (2018).[79][80][81]
  • A study on the body size and diversity of Carnian conodonts from South China and their implications for inferring the biotic and environmental changes during the Carnian Pluvial Event is published by Zhang et al. (2018).[82]
  • A study assessing the similarity of late Paleozoic to Triassic conodont faunas known from the Cache Creek terrane (Canada) is published by Golding (2018).[83]
  • Reconstruction of the multi-element apparatus of the Middle Triassic conodont from British Columbia (Canada) belonging to the Neogondolella regalis group within the genus Neogondolella is presented by Golding (2018).[84]
  • Reconstruction of the number and arrangement of elements in the apparatus of Hindeodus parvus published by Zhang et al. (2017)[85] is criticized by Agematsu, Golding & Orchard (2018);[86] Purnell et al. (2018) defend their original conclusions.[87]
  • A cluster of icriodontid conodonts belonging to the species Caudicriodus woschmidti, providing new information on the apparatus structure of icriodontid conodonts, is described from the Lower Devonian sediments in southern Burgenland (Austria) by Suttner, Kido & Briguglio (2018).[88]
  • A study on the species belonging to the genus Neognathodus, evaluating whether previously defined morphotype groups are reliably distinct from one another, is published by Zimmerman, Johnson & Polly (2018).[89]

New taxa

Name Novelty Status Authors Age Type locality Country Notes

Ancyrogondolella diakowi[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Ancyrogondolella equalis[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Ancyrogondolella inequalis[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Ancyrogondolella? praespiculata[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Ancyrogondolella transformis[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Baltoniodus cooperi[91]

Sp. nov

Valid

Carlorosi, Sarmiento & Heredia

Ordovician (Dapingian)

Santa Gertrudis Formation

Argentina

Declinognathodus intermedius[92]

Sp. nov

Valid

Hu, Qi & Nemyrovska

Carboniferous

China

Declinognathodus tuberculosus[92]

Sp. nov

Valid

Hu, Qi & Nemyrovska

Carboniferous

China

Gedikella[93]

Gen. et sp. nov

Valid

Kılıç, Plasencia & Önder

Middle Triassic (Anisian)

Turkey

A member of the family Gondolellidae. The type species is G. quadrata.

Gnathodus mirousei[94]

Sp. nov

Valid

Sanz-López & Blanco-Ferrera

Carboniferous (Mississippian)

Alba Formation
Aspe-Brousset Formation
Black Rock Limestone

Belgium
China
Ireland
Italy
Spain
United Kingdom United States
( Illinois)

Idiognathodus abdivitus[95]

Sp. nov

Valid

Hogancamp & Barrick

Carboniferous

Atrasado Formation
Eudora Shale

United States
( New Mexico)

Originally described as a species of Idiognathodus, but subsequently transferred to the genus Heckelina.[96]

Idiognathodus centralis[95]

Sp. nov

Valid

Hogancamp & Barrick

Carboniferous

Atrasado Formation
Eudora Shale

United States
( New Mexico)

Idiognathodus sweeti[95]

Sp. nov

Valid

Hogancamp & Barrick

Carboniferous

Atrasado Formation
Eudora Shale

United States
( New Mexico)

Idiognathoides chaagulootus[97]

Sp. nov

Valid

Frederick & Barrick

Carboniferous (early Pennsylvanian)

Ladrones Limestone

United States
( Alaska)

Kamuellerella rectangularis[93]

Sp. nov

Valid

Kılıç, Plasencia & Önder

Middle Triassic (Anisian)

Turkey

A member of the family Gondolellidae.

Ketinella goermueshi[93]

Sp. nov

Valid

Kılıç, Plasencia & Önder

Middle Triassic (Anisian)

Turkey

A member of the family Gondolellidae.

Magnigondolella[98]

Gen. et 5 sp. et comb. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

Favret Formation
Toad Formation

Canada
( British Columbia)
China
United States
( Nevada)

A member of the family Gondolellidae. The type species is M. salomae;
genus also includes new species M. alexanderi, M. cyri, M. julii and M. nebuchadnezzari,
as well as "Neogondolella" regale Mosher (1970) and "Neogondolella" dilacerata Golding & Orchard (2016).

Mesogondolella hendersoni[99]

Sp. nov

Valid

Yuan, Zhang & Shen

Permian (Changhsingian)

Selong Group

China

Mockina? spinosa[90]

Sp. nov

Valid

Orchard

Late Triassic (Norian)

Pardonet Formation

Canada
( British Columbia)

A member of the family Gondolellidae.

Neopolygnathus fibula[100]

Sp. nov

Valid

Hartenfels & Becker

Devonian (Famennian)

Morocco

Neospathodus arcus[101]

Sp. nov

Valid

Maekawa in Maekawa, Komatsu & Koike

Early Triassic

Taho Formation

Japan

Novispathodus shirokawai[101]

Sp. nov

Valid

Maekawa in Maekawa, Komatsu & Koike

Early Triassic

Taho Formation

Japan

Novispathodus tahoensis[101]

Sp. nov

Valid

Maekawa in Maekawa, Komatsu & Koike

Early Triassic

Taho Formation

Japan

'Ozarkodina'? chenae[102]

Sp. nov

Valid

Lu et al.

Devonian (Emsian)

Ertang Formation

China

'Ozarkodina'? wuxuanensis[102]

Sp. nov

Valid

Lu et al.

Devonian (Emsian)

Ertang Formation

China

Polygnathus linguiformis saharicus[103]

Subsp. nov

Valid

Narkiewicz & Königshof

Devonian (late Eifelian–middle Givetian)

Ispena Formation
Si Phai Formation

Morocco
Spain
Tajikistan
Turkey
Vietnam

Polygnathus linguiformis vietnamicus[103]

Subsp. nov

Valid

Narkiewicz & Königshof

Devonian (Givetian)

Plum Brook Shale
Si Phai Formation

Germany
Morocco
United States
( Ohio)
Vietnam

Polygnathus praeinversus[102]

Sp. nov

Valid

Lu et al.

Devonian (Emsian)

Ertang Formation

China

Polygnathus rhenanus siphai[103]

Subsp. nov

Valid

Narkiewicz & Königshof

Devonian (Givetian)

Candás Formation
Si Phai Formation

China
Morocco
Spain
Vietnam

Polygnathus xylus bacbo[103]

Subsp. nov

Valid

Narkiewicz & Königshof

Devonian (Givetian)

Si Phai Formation

Vietnam

Pseudognathodus posadachaconae[104]

Sp. nov

Valid

Sanz-López, Blanco-Ferrera & Miller

Carboniferous (Mississippian)

Prestatyn Limestone

United Kingdom

A member of the family Gnathodontidae.

Pseudopolygnathus primus tafilensis[100]

Subsp. nov

Valid

Hartenfels & Becker

Devonian (Famennian)

Morocco

Pustulognathus[105]

Gen. et 2 sp. nov

Valid

Golding & Orchard in Golding

Permian (Guadalupian to Lopingian)

Copley Limestone
Horsefeed Formation

Canada
( British Columbia)
China?

A member of the family Sweetognathidae. The type species is P. monticola; genus also includes P. vigilans.

Quadralella (Quadralella) postica[106]

Sp. nov

Valid

Zhang et al.

Late Triassic (Carnian)

China

Quadralella robusta[106]

Sp. nov

Valid

Zhang et al.

Late Triassic (Carnian)

China

Quadralella wignalli[106]

Sp. nov

Valid

Zhang et al.

Late Triassic (Carnian)

China

Quadralella yongningensis[106]

Sp. nov

Valid

Zhang et al.

Late Triassic (Carnian)

China

Scandodus choii[107]

Sp. nov

Valid

Lee

Ordovician (Darriwilian)

South Korea

Sweetognathus duplex[108]

Sp. nov

Valid

Read & Nestell

Permian (Sakmarian)

Riepe Spring Limestone

United States
( Nevada)

Sweetognathus wardlawi[108]

Sp. nov

Valid

Read & Nestell

Permian (Sakmarian)

Riepe Spring Limestone

United States
( Nevada)

"Tortodus" sparlingi[109]

Sp. nov

Valid

Aboussalam & Becker in Brett et al.

Devonian (Givetian)

Poland
Spain
United States
( Kentucky
Ohio)

Walliserognathus[110]

Gen. et comb. nov

Valid

Corradini & Corriga

Silurian (Ludlow)

Henryhouse Formation
Roberts Mountains Formation

Austria
China
Hungary
Italy
Spain
Sweden
United States
( Nevada
Oklahoma)

A member of the family Spathognathodontidae; a new genus for Spathognathodus inclinatus posthamatus Walliser (1964), raised to the rank of the species Walliserognathus posthamatus.

Fish

Amphibians

Reptiles

Synapsids

Non-mammalian synapsids

Research

  • A description of the postcranial material referable to the caseid species Ennatosaurus tecton is published by Romano, Brocklehurst & Fröbisch (2018).[111]
  • A study on the anatomy and phylogenetic relationships of Milosaurus mccordi is published by Brocklehurst & Fröbisch (2018).[112]
  • A skull of a juvenile specimen of Anteosaurus magnificus is described from the Permian Abrahamskraal Formation (South Africa) by Kruger, Rubidge & Abdala (2018).[113]
  • A study on the evolution of the trigeminal nerve innervation in anomodonts is published by Benoit et al. (2018).[114]
  • A study on the stable oxygen and carbon isotope compositions of dentine apatite in the teeth of twenty-eight specimens of Diictodon feliceps, and on their implications for inferring the potential role of climate in driving the late Capitanian mass extinction of terrestrial tetrapods, is published by Rey et al. (2018).[115]
  • Description of the anatomy of six new skulls of the dicynodont Abajudon kaayai from the Permian (Guadalupian) lower Madumabisa Mudstone Formation (Zambia) and a study on the phylogenetic relationships of the species is published by Olroyd, Sidor & Angielczyk (2018).[116]
  • A study on the anatomy of the bony labyrinth of the specimens of the dicynodont genus Endothiodon collected from the Permian K5 Formation (Mozambique), comparing it with the closely related genus Niassodon, is published by Araújo et al. (2018).[117]
  • A study on the taphonomic history of a monotypic bonebed composed by several individuals attributable to the dicynodont Dinodontosaurus collected in a classic Middle Triassic locality in Brazil, and on its implications for inferring possible gregarious behaviour in Dinodontosaurus, is published online by Ugalde et al. (2018).[118]
  • Redescription of the dicynodont genus Sangusaurus and a study on its feeding system and phylogenetic relationships is published by Angielczyk, Hancox & Nabavizadeh (2018).[119]
  • Partial hindlimb of a dicynodont nearing the size of Stahleckeria potens is described from the Triassic Lifua Member of the Manda Beds (Tanzania) by Kammerer, Angielczyk & Nesbitt (2018), representing the largest dicynodont postcranial element from the Manda Beds reported so far.[120]
  • Description of plant remains and palynomorphs preserved in the coprolites produced by large dicynodonts from the Triassic Chañares Formation (Argentina), and a study on their implications for inferring the diet of dicynodonts, is published by Perez Loinaze et al. (2018).[121]
  • Tetrapod tracks, probably produced by dicynodonts, are described from the Upper Triassic Vera Formation of the Los Menucos Group (Argentina) by Citton et al. (2018).[122]
  • A study on the age of putative Rhaetian dicynodont from Lipie Śląskie (Poland) is published online by Racki & Lucas (2018), who consider it more likely that this dicynodont was of Norian age.[123]
  • A study on the anatomy of the skull of Cynariops robustus is published by Bendel et al. (2018).[124]
  • A study on rates of enamel development in a range of non-mammalian cynodont species, inferred from incremental markings, is published by O'Meara, Dirks & Martinelli (2018).[125]
  • Description of the morphology of the skull of Cynosaurus suppostus and a study on the phylogenetic relationships of the species is published by van den Brandt & Abdala (2018).[126]
  • Fossils of Cynognathus crateronotus are described for the first time from the Triassic Ntawere Formation (Zambia) and Manda Beds (Tanzania) by Wynd et al. (2018).[127]
  • A study on the postcranial anatomy of a specimen of Diademodon tetragonus recovered from the Upper Omingonde Formation (Namibia) is published by Gaetano, Mocke & Abdala (2018).[128]
  • Partial skull and postcranial skeleton of a member of the species Cricodon metabolus is described from the Triassic Ntawere Formation (Zambia) by Sidor & Hopson (2018), who also study the phylogenetic relationships of members of the family Trirachodontidae.[129]
  • A study on the musculature, posture and range of motion of the forelimb of Massetognathus pascuali is published by Lai, Biewener & Pierce (2018).[130]
  • New specimen of Trucidocynodon riograndensis, almost 20% larger than the holotype specimen, is described from the Carnian of Candelária Sequence (southern Brazil) by Stefanello et al. (2018).[131]
  • Right dentary with teeth of Prozostrodon brasiliensis is described from the Late Triassic of Brazil by Pacheco et al. (2018), representing the second known specimen of this species.[132]
  • Description of the anatomy of the postcranial skeleton of Prozostrodon brasiliensis is published by Guignard, Martinelli & Soares (2018).[133]
  • A study on the limb bone histology and life histories of Prozostrodon brasiliensis, Irajatherium hernandezi, Brasilodon quadrangularis and Brasilitherium riograndensis is published by Botha-Brink, Bento Soares & Martinelli (2018).[134]
  • A study on the origin and relationships of ictidosaurian cynodonts, i.e. tritheledontids and therioherpetids, is published by Bonaparte & Crompton (2018).[135]
  • A large (comprising at least 38 individuals) clutch of well-preserved perinates of Kayentatherium wellesi, found with a presumed maternal skeleton, is described from the Lower Jurassic sediments of the Kayenta Formation (found on lands of the Navajo Nation) by Hoffman & Rowe (2018);[136] in light of this finding, a new interpretation of earlier records of associations between adult and juvenile cynodonts is proposed by Benoit (2019).[137]
  • Cynodont teeth (representing a brasilodontid and a Riograndia-like form) found in the Triassic locality in Brazil which also yielded the fossils of Sacisaurus agudoensis are described by Marsola et al. (2018).[138]
  • A study on the evolution of the mammalian jaw is published by Lautenschlager et al. (2018), who find no evidence for a concurrent reduction in jaw-joint stress and increase in bite force in key non-mammaliaform taxa in the cynodont–mammaliaform transition.[139]
  • Tetrapod burrows, likely produced by small eucynodonts, are described from the Triassic Chañares Formation (Argentina) by Fiorelli et al. (2018).[140]
  • A study on the morphological diversity of vertebral regions in non-mammalian synapsids, and on its implication for elucidating the evolution of anatomically distinct regions of the mammalian spines, is published by Jones et al. (2018).[141]
  • A study on teeth ontogeny in wide range of extinct synapsid lineages is published by LeBlanc et al. (2018), who interpret their findings as indicating that the ligamentous tooth attachment system is not unique to crown mammals within Synapsida.[142]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Ascendonanus[143]

Gen. et sp. nov

Valid

Spindler et al.

Permian (Sakmarian-Artinskian transition)

Chemnitz petrified forest
(Leukersdorf Formation)

Germany

A member of the family Varanopidae. Genus includes new species A. nestleri.

Gordodon[144]

Gen. et sp. nov

Valid

Lucas, Rinehart & Celeskey

Early Permian (early Wolfcampian)

Bursum Formation

United States
( New Mexico)

A member of the family Edaphosauridae. The type species is G. kraineri.

Gorynychus[145]

Gen. et sp. nov

Valid

Kammerer & Masyutin

Permian

Kotelnich red beds

Russia
( Kirov Oblast)

A therocephalian. The type species is G. masyutinae.

Leucocephalus[146]

Gen. et sp. nov

Valid

Day et al.

Permian (early Wuchiapingian)

Tropidostoma Assemblage Zone of the Main Karoo Basin

South Africa

A biarmosuchian belonging to the family Burnetiidae. The type species is L. wewersi.

Lisowicia[147]

Gen. et sp. nov

Sulej & Niedźwiedzki

Late Triassic (late Norian-earliest Rhaetian)

Poland

A gigantic dicynodont reaching an estimated body mass of 9 tons. The type species is L. bojani. Announced in 2018; the final version of the article naming it was published in 2019.

Microvaranops[143]

Gen. et sp. nov

Valid

Spindler et al.

Permian (Guadalupian)

Abrahamskraal Formation

South Africa

A member of the family Varanopidae. Genus includes new species M. parentis.

Nochnitsa[148]

Gen. et sp. nov

Valid

Kammerer & Masyutin

Permian

Kotelnich red beds

Russia
( Kirov Oblast)

A gorgonopsian. The type species is N. geminidens.

Pentasaurus[149]

Gen. et sp. nov

Valid

Kammerer

Late Triassic

Elliot Formation

South Africa

A dicynodont belonging to the family Stahleckeriidae. The type species is P. goggai.

Polonodon[150]

Gen. et sp. nov

Valid

Sulej et al.

Late Triassic (Carnian)

Poland

A non-mammaliaform eucynodont. Genus includes new species P. woznikiensis. Announced in 2018; the final version of the article naming it was published in 2020.

Siriusgnathus[151]

Gen. et sp. nov

Valid

Pavanatto et al.

Late Triassic (Carnian or Norian[152])

Santa Maria Supersequence

Brazil

A traversodontid cynodont. Genus includes new species S. niemeyerorum.

Mammals

Other animals

Research

  • A review and synthesis of studies on the timing and environmental context of landmark events in early animal evolution is published by Sperling & Stockey (2018).[153]
  • A study on the phylogenetic relationships of the rangeomorphs, dickinsoniomorphs and erniettomorphs as indicated by what is known of the ontogeny of the rangeomorph Charnia masoni, dickinsoniomorph Dickinsonia costata and erniettomorph Pteridinium simplex is published by Dunn, Liu & Donoghue (2018), who consider at least the rangeomorphs and dickinsoniomorphs to be metazoans.[154]
  • A study on the phylogenetic relationships of the rangeomorphs is published by Dececchi et al. (2018).[155]
  • A study on the size distribution and morphological features of a population of juvenile specimens of Dickinsonia costata from the Crisp Gorge fossil locality in the Flinders Ranges (Australia) is published by Reid, García-Bellido & Gehling (2018).[156]
  • A study on the phylogenetic relationships of Dickinsonia based on data from lipid biomarkers extracted from organically preserved Ediacaran macrofossils is published by Bobrovskiy et al. (2018), who interpret their findings as indicating that Dickinsonia was an animal.[157]
  • A study on the anatomy and phylogenetic relationships of Stromatoveris, based on data from new specimens from the Chengjiang Konservat-Lagerstätte (China), is published by Hoyal Cuthill & Han (2018), who interpret Stromatoveris as a member of early animal group Petalonamae that also included Arborea, Pambikalbae, rangeomorphs, dickinsoniomorphs and erniettomorphs.[158]
  • The first reliable occurrence of abundant penetrative trace fossils, providing trace fossil evidence for Precambrian bilaterians with complex behavioural patterns, is reported from the latest Ediacaran of western Mongolia by Oji et al. (2018).[159]
  • Trace fossils produced by Ediacaran animals which burrowed within sediment are described from the shallow-marine deposits of the Urusis Formation (Nama Group, Namibia) by Buatois et al. (2018), who name a new ichnotaxon Parapsammichnites pretzeliformis.[160]
  • New trace fossils from the Ediacaran Shibantan Member of the upper Dengying Formation (China), including burrows and possible trackways which were probably made by millimeter-sized animals with bilateral appendages, are described by Chen et al. (2018).[161]
  • An aggregation of members of the genus Parvancorina, providing evidence of two size-clusters and bimodal orientation in this taxon, is described from the Ediacara Conservation Park (Australia) by Coutts et al. (2018).[162]
  • New, three-dimensional specimens of Charniodiscus arboreus (Arborea arborea), allowing for a detailed reinterpretation of its functional morphology and taxonomy, are described from the Ediacara Member, Rawnsley Quartzite of South Australia by Laflamme, Gehling & Droser (2018).[163]
  • 3D reconstructions of Cloudina aggregates are presented by Mehra & Maloof (2018).[164]
  • A study on Namacalathus and Cloudina skeletons from the Ediacaran Omkyk Member of the Nama Group (Namibia) is published by Pruss et al. (2018), who interpret their findings as indicating that both organisms originally produced aragonitic skeletons, which later underwent diagenetic conversion to calcite.[165]
  • A study on the substrate growth dynamics, mode of biomineralization and possible affinities of Namapoikia rietoogensis is published by Wood & Penny (2018).[166]
  • A review of evidence for existence of swimming animals during the Neoproterozoic is published by Gold (2018).[167]
  • A study on the age of the Cambrian Chengjiang biota (China) is published by Yang et al. (2018).[168]
  • Description of coprolites from the Cambrian (Drumian) Rockslide Formation (Mackenzie Mountains, Canada) produced by an unknown predator, and a study on their implications for reconstructing the Cambrian food web, is published by Kimmig & Pratt (2018).[169]
  • A study on the nature and biological affinity of the Cambrian taxon Archaeooides is published by Yin et al. (2018), who interpret the fossils of Archaeooides as embryonic remains of animals.[170]
  • Zumberge et al. (2018) report a new fossil sterane biomarker, possessing a rare hydrocarbon skeleton that is uniquely found within extant demosponge taxa, from late NeoproterozoicCambrian sedimentary rocks and oils, and interpret this finding as indicating that demosponges, and hence multicellular animals, were prominent in some late Neoproterozoic marine environments at least extending back to the Cryogenian period.[171]
  • Diverse, abundant sponge fossils from the Ordovician–Silurian boundary interval are reported from seven localities in South China by Botting et al. (2018), who produce a model for the distribution and preservation of the sponge fauna.[172]
  • A study on the phylogenetic relationships of extant and fossil demosponges is published by Schuster et al. (2018).[173]
  • An assemblage of animal fossils, including the oldest known pterobranchs, preserved in the form of small carbonaceous fossils is described from the Cambrian Buen Formation (Greenland) by Slater et al. (2018).[174]
  • Description of new morphological features of the Cambrian mobergellan Discinella micans is published by Skovsted & Topper (2018).[175]
  • A study on the interrelationships between the eldonioid Pararotadiscus guizhouensis and associated fossil taxa from the Kaili Biota is published by Zhao et al. (2018).[176]
  • A study on the slab with a dense aggregation of members of the species Banffia constricta recovered from the Cambrian Burgess Shale (Canada) and its implications for life habits of the animal is published by Chambers & Brandt (2018).[177]
  • A study on the morphology and phylogenetic affinities of Yuyuanozoon magnificissimi, based on new specimens, is published by Li et al. (2018).[178]
  • A study on the fossil record of early Paleozoic graptoloids and on the factors influencing rates of diversification within this group is published by Foote et al. (2018).[179]
  • A study on the impact of the long-period astronomical cycles (Milankovitch "grand cycles") associated with Earth's orbital eccentricity and obliquity on the variance in species turnover probability (extinction probability plus speciation probability) in Early Paleozoic graptoloids is published by Crampton et al. (2018).[180]
  • A redescription of the species Malongitubus kuangshanensis from the Cambrian Chengjiang Lagerstätte (China) is published by Hu et al. (2018), who interpret this taxon as a pterobranch.[181]
  • A study on the morphology of the palaeoscolecid worm Palaeoscolex from the Lower Ordovician Fezouata Lagerstätte (Morocco), using computed microtomography and providing new information on the internal anatomy of this animal, is published by Kouraiss et al. (2018).[182]
  • The first occurrence of the tommotiid species Paterimitra pyramidalis from the Xinji Formation (China) is reported by Pan et al. (2018).[183]
  • A study on the temporal distribution of lophotrochozoan skeletal species from the upper Ediacaran to the basal Miaolingian of the Siberian Platform, and on its implications for understanding the evolutionary dynamics of the Cambrian explosion, is published by Zhuravlev & Wood (2018).[184]
  • Eggs of ascaridoid nematodes found in crocodyliform coprolites are described from the Upper Cretaceous Bauru Group (Brazil) by Cardia et al. (2018).[185]
  • A study reinterpreting the putative Cambrian lobopodian Mureropodia apae as a partial isolated appendage of a member of the genus Caryosyntrips, published by Pates & Daley (2017)[186] is criticized by Gámez Vintaned & Zhuravlev (2018);[187] Pates, Daley & Ortega-Hernández (2018) defend their original conclusions.[188]
  • A study on the early evolution of stem and crown-arthropods as indicated by Ediacaran and Cambrian body and trace fossils is published by Daley et al. (2018).[189]
  • A study on the evolution of ecdysozoan vision, focusing on the evolution of arthropod multi-opsin vision, as indicated by molecular data and data from fossil record, is published by Fleming et al. (2018).[190]
  • A juvenile specimen of Lyrarapax unguispinus, providing new information on the frontal appendages and feeding mode in this taxon, is described from the Cambrian Chiungchussu Formation (China) by Liu et al. (2018).[191]
  • A study evaluating likely swimming efficiency and maneuverability of Anomalocaris canadensis is published by Sheppard, Rival & Caron (2018).[192]
  • Cambrian animal Pahvantia hastata from the Wheeler Shale (Utah, United States), originally classified as a possible arthropod,[193] is reinterpreted as a suspension-feeding radiodont by Lerosey-Aubril & Pates (2018).[194]
  • The presence of metameric midgut diverticulae is reported for the first time in the stem-arthropod Fuxianhuia protensa by Ortega-Hernández et al. (2018), who interpret their finding as indicative of a predatory or scavenging ecology of fuxianhuiids.[195]
  • Liu et al. (2018) reinterpret putative remains of the nervous and cardiovascular systems in numerous articulated individuals of Fuxianhuia protensa as more likely to be microbial biofilms that developed following decomposition of the intestine, muscle and other connective tissues.[196]
  • A study on the post-embryonic development of Fuxianhuia protensa is published by Fu et al. (2018).[197]
  • Redescription of the fuxianhuiid Liangwangshania biloba is published by Chen et al. (2018).[198]
  • New specimens of the stem-arthropod species Kerygmachela kierkegaardi, providing new information on the anatomy of this species and on the ancestral condition of the panarthropod brain, are described from the Cambrian Stage 3 of the Buen Formation (Sirius Passet, Greenland) by Park et al. (2018).[199]
  • Fossils of spindle- or conotubular-shaped animals of uncertain phylogenetic placement are described from the Ordovician Martinsburg Formation (Pennsylvania, United States) by Meyer et al. (2018).[200]
  • Evidence of macrofauna living at depths of up to 8 metres below the seabed is reported from the Permian Fort Brown Formation (Karoo Basin, South Africa) by Cobain et al. (2018).[201]
  • A study on the morphology of the hyolithid Paramicrocornus zhenbaensis from the lower Cambrian Shuijingtuo Formation (China) is published by Zhang, Skovsted & Zhang (2018), who report that this species lacked helens, and also report the oldest known hyolith muscle scars preserved on the opercula of this species.[202]
  • A study on the feeding strategies and locomotion of Cambrian hyolithids, based on specimens preserved in coprolites from the Chengjiang biota and associated with a Tuzoia carcass from the Balang Fauna (China), is published by Sun et al. (2018).[203]
  • Digestive tract of a specimen of the hyolith species Circotheca johnstrupi from the Cambrian Læså Formation (Bornholm, Denmark) is described by Berg-Madsen, Valent & Ebbestad (2018).[204]
  • The oldest stromatoporoidbryozoan reefs reported so far are described from the middle Ordovician Duwibong Formation (South Korea) by Hong et al. (2018).[205]
  • Small bioconstructions formed solely by microconchid tube worms, representing the stratigraphically oldest exclusively metazoan bioconstructions from the earliest Triassic (mid-Induan) strata in East Greenland, are reported by Zatoń et al. (2018).[206]
  • The oldest known evidence of trematode parasitism of bivalves in the form of igloo-shaped traces found on shells of the freshwater bivalve Sphaerium is reported from the Upper Cretaceous Judith River Formation (Montana, United States) by Rogers et al. (2018).[207]
  • A study on the predatory drill holes in Late Cretaceous and Paleogene molluscan and serpulid worm prey from Seymour Island (Antarctica) and their implications for inferring the effects of the Cretaceous–Paleogene extinction event on predator-prey dynamics at this site is published by Harper, Crame & Sogot (2018).[208]
  • A study on burrows from Lower–Middle Triassic successions in South China assigned to the ichnotaxon Rhizocorallium, and on their implications for inferring the course of biotic recovery following the Permian–Triassic extinction event, is published by Feng et al. (2018).[209]
  • A study evaluating how different species of fossil and extant free-living cupuladriid bryozoans responded to the environmental changes in the Southwest Caribbean over the last 6 million years is published by O'Dea et al. (2018).[210]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Acanthodesia variegata[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Membraniporidae.

Acoscinopleura albaruthenica[211]

Sp. nov

Valid

Koromyslova, Martha & Pakhnevich

Late Cretaceous (late Campanian)

Belarus

A bryozoan belonging to the group Flustrina and the family Coscinopleuridae.

Acoscinopleura crassa[211]

Sp. nov

Valid

Koromyslova, Martha & Pakhnevich

Late Cretaceous (Maastrichtian)

Germany

A bryozoan belonging to the group Flustrina and the family Coscinopleuridae.

Acoscinopleura dualis[211]

Sp. nov

Valid

Koromyslova, Martha & Pakhnevich

Late Cretaceous (Maastrichtian)

Germany

A bryozoan belonging to the group Flustrina and the family Coscinopleuridae.

Acoscinopleura occulta[211]

Sp. nov

Valid

Koromyslova, Martha & Pakhnevich

Late Cretaceous (Maastrichtian)

Germany

A bryozoan belonging to the group Flustrina and the family Coscinopleuridae.

'Aechmella' viskovae[212]

Sp. nov

Valid

Koromyslova, Baraboshkin & Martha

Late Cretaceous

Kazakhstan

A bryozoan.

Aechmellina[213]

Gen. et comb. nov

Valid

Taylor, Martha & Gordon

Cretaceous (Cenomanian) to Paleocene (Danian).

Denmark
France
Germany
United Kingdom
United States

A bryozoan belonging to the group Flustrina and the family Onychocellidae. The type species is "Aechmella" falcifera Voigt (1949); genus also includes "Homalostega" anglica Brydone (1909), "Aechmella" bassleri Voigt (1924), "Homalostega" biconvexa Brydone (1909), "Cellepora" hippocrepis Goldfuss (1826), "Aechmella" indefessa Taylor & McKinney (2006), "Aechmella" latistoma Berthelsen (1962), "Aechmella" linearis Voigt (1924), "Aechmella" parvilabris Voigt (1924), "Aechmella" pindborgi Berthelsen (1962), "Semieschara" proteus Brydone (1912), "Monoporella" seriata Levinsen (1925), "Aechmella" stenostoma Voigt (1930), "Reptescharinella" transversa d'Orbigny (1852) and "Aechmella" ventricosa Voigt (1924).

Alacaris[214]

Gen. et sp. nov

Valid

Yang et al.

Cambrian Stage 3

Hongjingshao Formation

China

A stem-arthropod related to Chengjiangocaris. The type species is A. mirabilis.

Allonnia nuda[215]

Sp. nov

Valid

Cong et al.

Cambrian Stage 3

Chengjiang Lagerstätte

China

A chancelloriid.

Allonnia tenuis[216]

Sp. nov

Valid

Zhao, Li & Selden

Early Cambrian

China

A chancelloriid.

Arnaopora[217]

Gen. et sp. nov

Valid

Suárez Andrés & Wyse Jackson

Devonian

Moniello Formation

Spain

A bryozoan belonging to the group Fenestrata. Genus includes new species A. sotoi.

Aspidostoma armatum[218]

Sp. nov

Valid

Pérez, López-Gappa & Griffin

Early Miocene

Monte León Formation

Argentina

A cheilostome bryozoan belonging to the family Aspidostomatidae.

Aspidostoma roveretoi[218]

Sp. nov

Valid

Pérez, López-Gappa & Griffin

Late Miocene

Puerto Madryn Formation

Argentina

A cheilostome bryozoan belonging to the family Aspidostomatidae.

Aspidostoma tehuelche[218]

Sp. nov

Valid

Pérez, López-Gappa & Griffin

Early to middle Miocene

Chenque Formation

Argentina

A cheilostome bryozoan belonging to the family Aspidostomatidae.

Austroscolex sinensis[219]

Sp. nov

Valid

Liu et al.

Cambrian (Paibian)

China

A palaeoscolecid.

Axilosoecia[220]

Gen. et 2 sp. nov

Valid

Taylor & Brezina

Paleocene (Danian) to early Miocene

Roca Formation

Argentina
New Zealand

A bryozoan belonging to the group Tubuliporina and the family Oncousoeciidae. The type species is A. giselae; genus also includes A. mediorubiensis.

Burocratina[221]

Gen. et sp. nov

Wachtler & Ghidoni

Early-Middle Triassic

Italy

A polychaete. The type species is B. kraxentrougeri.

Catenagraptus[222]

Gen. et sp. nov

Valid

VandenBerg

Ordovician (late Floian)

Australia

A graptolite belonging to the group Sinograptina and the family Sigmagraptidae. The type species is C. communalis.

Characodoma wesselinghi[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Cleidochasmatidae.

Cheethamia aktolagayensis[212]

Sp. nov

Valid

Koromyslova, Baraboshkin & Martha

Late Cretaceous

Kazakhstan

A bryozoan.

Codositubulus[223]

Gen. et sp. nov

Valid

Gámez Vintaned et al.

Cambrian

Spain

A tubicolous animal of uncertain phylogenetic placement. The type species is C. grioensis.

Colospongia lenis[224]

Sp. nov

Valid

Malysheva

Late Permian

Russia
( Primorsky Krai)

A sponge.

Cornulites gondwanensis[225]

Sp. nov

Valid

Gutiérrez-Marco & Vinn

Ordovician (Hirnantian)

Morocco

A member of Cornulitida.

Cupitheca convexa[226]

Sp. nov

Valid

Sun et al.

Cambrian

Manto Formation

China

A member of Hyolitha.

Cystomeson[227]

Gen. et sp. nov

Valid

Ernst, Krainer & Lucas

Carboniferous (Mississippian)

Lake Valley Formation

United States
( New Mexico)

A bryozoan belonging to the group Cystoporata. Genus includes new species C. sierraensis.

Decoritheca? hageni[228]

Sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A member of Hyolitha.

Demirastrites campograptoides[229]

Sp. nov

Valid

Štorch & Melchin

Silurian (Aeronian)

Czech Republic

A graptolite belonging to the family Monograptidae.

Dictyocyathus aranosensis[230]

Sp. nov

Valid

Perejón et al.

Early Cambrian

Namibia

A member of Archaeocyatha.

Didymograptellus kremastus[231]

Sp. nov

Valid

Vandenberg

Ordovician (Floian)

Australia
New Zealand
Norway
United States

A graptolite belonging to the group Dichograptina and the family Pterograptidae.

Erismacoscinus ganigobisensis[230]

Sp. nov

Valid

Perejón et al.

Early Cambrian

Namibia

A member of Archaeocyatha.

'Escharoides' charbonnieri[232]

Sp. nov

Valid

Di Martino, Martha & Taylor

Late Cretaceous (Maastrichtian)

Madagascar

A bryozoan.

Fehiborypora[232]

Gen. et comb. nov

Valid

Di Martino, Martha & Taylor

Late Cretaceous (Maastrichtian)

Madagascar

A bryozoan; a new genus for "Cribilina" labiatula Canu (1922).

Gibbavasis[233]

Gen. et sp. nov

Vaziri, Majidifard & Laflamme

Ediacaran

Kushk Series

Iran

A vase-shaped organism of uncertain phylogenetic placement, possibly a poriferan-grade animal. The type species is G. kushkii.

Homoctenus katzerii[234]

Sp. nov

Valid

Comniskey & Ghilardi

Devonian (late Pragian or late Emsian)

Ponta Grossa Formation

Brazil

A member of Tentaculitoidea belonging to the order Homoctenida and the family Homoctenidae.

Kalaallitia[228]

Gen. et sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A member of Hyolitha. Genus includes new species K. myliuserichseni.

Kamilocella[213]

Gen. et comb. nov

Valid

Taylor, Martha & Gordon

Late Cretaceous (Cenomanian) to Campanian).

Czech Republic
France
Germany

A bryozoan belonging to the group Flustrina and the family Onychocellidae. The type species is "Eschara" latilabris Reuss (1872); genus also includes "Eschara" acis d'Orbigny (1851), "Onychocella" barbata Martha, Niebuhr & Scholz (2017), "Eschara" cenomana d'Orbigny (1851) and "Eschara" labiata Počta (1892).

Kenocharixa[235]

Gen. et sp. et comb. nov

Valid

Dick, Sakamoto & Komatsu

Cretaceous to Eocene

Japan
New Zealand

A cheilostome bryozoan. Genus includes new species K. kashimaensis, as well as "Charixa goshouraensis Dick, Komatsu, Takashima & Ostrovsky (2013) and "Conopeum" stamenocelloides Gordon & Taylor (2015).

Khmeria minima[236]

Sp. nov

Valid

Wendt

Late Triassic (Carnian)

Italy

An ascidian belonging to the new order Khmeriamorpha.

Khmeria stolonifera[236]

Sp. nov

Valid

Wendt

Late Permian, possibly also Carboniferous

Cambodia
Thailand
Vietnam

An ascidian belonging to the new order Khmeriamorpha.

Kimberella persii[233]

Sp. nov

Vaziri, Majidifard & Laflamme

Ediacaran

Kushk Series

Iran

A stem-mollusc bilaterian.

Kootenayscolex[237]

Gen. et sp. nov

Valid

Nanglu & Caron

Cambrian

Burgess Shale

Canada
( British Columbia)

A polychaete. Genus includes new species K. barbarensis.

Laminacaris[238]

Gen. et sp. nov

Valid

Guo et al.

Cambrian Stage 3

China
United States?[239]

A member of Radiodonta. Genus includes new species L. chimera.

Lenisambulatrix[240]

Gen. et sp. nov

Valid

Ou & Mayer

Cambrian Stage 3

Heilinpu Formation

China

A lobopodian. The type species is L. humboldti.

Lunulites marambionis[241]

Sp. nov

Valid

Hara et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A bryozoan belonging to the group Cheilostomata and the family Lunulitidae.

Marginaria prolixa[235]

Sp. nov

Valid

Dick, Sakamoto & Komatsu

Late Cretaceous (Campanian)

Himenoura Group

Japan

A cheilostome bryozoan.

Matteolaspongia[242]

Gen. et sp. nov

Valid

Botting, Zhang & Muir

Ordovician (Hirnantian)

Wenchang Formation

China

A sponge, possibly a stem-rossellid. The type species is M. hemiglobosa.

Melychocella biperforata[218]

Sp. nov

Valid

Pérez, López-Gappa & Griffin

Early Miocene

Chenque Formation
Monte León Formation

Argentina

A cheilostome bryozoan belonging to the family Aspidostomatidae.

Micrascidites gothicus[243]

Sp. nov

Valid

Sagular, Yümün & Meriç

Quaternary

Turkey

A didemnid ascidian.

Micropora nordenskjoeldi[241]

Sp. nov

Valid

Hara et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A bryozoan belonging to the group Cheilostomata and the family Microporidae.

Minitaspongia[244]

Gen. et sp. nov

Valid

Carrera et al.

Carboniferous (Tournaisian)

Agua de Lucho Formation

Argentina

A hexactinellid sponge belonging to the family Dictyospongiidae. The type species is M. parvis.

Monniotia minutula[243]

Sp. nov

Valid

Sagular, Yümün & Meriç

Quaternary

Turkey

A didemnid ascidian.

Nasaaraqia[228]

Gen. et sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A member of Hyolitha. Genus includes new species N. hyptiotheciformis.

Neotrematopora lyaoilensis[245]

Sp. nov

Valid

Tolokonnikova & Ponomarenko

Devonian (Frasnian)

Lyaiol Formation

Russia

A bryozoan.

Nevadotheca boerglumensis[228]

Sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A member of Hyolitha.

Nidelric gaoloufangensis[216]

Sp. nov

Valid

Zhao, Li & Selden

Early Cambrian

China

An animal with single-element spines.

Nogrobs moroccensis[246]

Sp. nov

Valid

Schlögl et al.

Middle Jurassic (Bajocian)

Agoudim Formation

Morocco

A serpulid polychaete.

Onuphionella corusca[247]

Sp. nov

Valid

Muir et al.

Ordovician (Sandbian)

First Bani Group

Morocco

Agglutinated tubes produced by unknown animal. Published online in 2018; the final version of the article naming it was published in 2022.

Otionellina antarctica[241]

Sp. nov

Valid

Hara et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A bryozoan belonging to the group Cheilostomata and the family Otionellidae.

Otionellina eocenica[241]

Sp. nov

Valid

Hara et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A bryozoan belonging to the group Cheilostomata and the family Otionellidae.

Pedunculotheca[248]

Gen. et sp. nov

Valid

Sun, Zhao & Zhu in Sun et al.

Cambrian Stage 3

Yu'anshan Formation

China

A member of Hyolitha belonging to the group Orthothecida. Genus includes new species P. diania.

'Plagioecia' antanihodiensis[232]

Sp. nov

Valid

Di Martino, Martha & Taylor

Late Cretaceous (Maastrichtian)

Madagascar

A bryozoan.

Platychelyna secunda[249]

Sp. nov

Valid

López-Gappa, Pérez & Griffin

Early Miocene

Monte León Formation

Argentina

A bryozoan.

Pleurocodonellina javanensis[35]

Sp. nov

Valid

Di Martino & Taylor

Early Pleistocene

Pucangan Formation

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Smittinidae.

Protohertzina compressa[250]

Sp. nov

Valid

Slater, Harvey & Butterfield

Cambrian (Terreneuvian)

Lontova Formation
Voosi Formation

Estonia

A member of the total group of Chaetognatha.

Qinscolex[251]

Gen. et sp. nov

Valid

Liu et al.

Cambrian (Fortunian)

China

A cycloneuralian tentatively assigned to total-group Scalidophora. Genus includes new species Q. spinosus.

Ramskoeldia[252]

Gen. et 2 sp. nov

Valid

Cong et al.

Cambrian

Maotianshan Shales

China

A member of Radiodonta related to Amplectobelua. Genus includes new species R. platyacantha and R. consimilis.

Reptomultisparsa stratosa[253]

Sp. nov

Valid

Viskova & Pakhnevich

Middle Jurassic (Callovian)

Russia

A bryozoan.

Rhagasostoma aralense[254]

Sp. nov

Valid

Koromyslova et al.

Late Cretaceous (Campanian)

Uzbekistan

A bryozoan belonging to the group Flustrina and the family Onychocellidae.

Rhagasostoma brydonei[254]

Sp. nov

Valid

Koromyslova et al.

Late Cretaceous (Turonian and Coniacian)

United Kingdom

A bryozoan belonging to the group Flustrina and the family Onychocellidae.

Rhagasostoma operculatum[254]

Sp. nov

Valid

Koromyslova et al.

Late Cretaceous (Campanian)

Turkmenistan

A bryozoan belonging to the group Flustrina and the family Onychocellidae.

Schistodictyon webbyi[255]

Sp. nov

Valid

Zhen

Late Silurian

Australia

A sponge belonging to the class Stromatoporoidea, order Clathrodictyida and the family Anostylostromatidae.

Seqineqia[256]

Gen. et sp. nov

Valid

Peel

Cambrian (Guzhangian)

Holm Dal Formation

Greenland

A sponge. The type species is S. bottingi.

"Serpula" calannai[257]

Sp. nov

Valid

Sanfilippo et al.

Permian

Italy.

A serpulid polychaete.

"Serpula" prisca[257]

Sp. nov

Valid

Sanfilippo et al.

Permian

Italy.

A serpulid polychaete.

Shaanxiscolex[258]

Gen. et sp. nov

Valid

Yang et al.

Cambrian Stage 4

China

A palaeoscolecid. The type species is S. xixiangensis.

Shanscolex[251]

Gen. et sp. nov

Valid

Liu et al.

Cambrian (Fortunian)

China

A cycloneuralian tentatively assigned to total-group Scalidophora. Genus includes new species S. decorus.

Sisamatispongia[256]

Gen. et sp. nov

Valid

Peel

Cambrian (Guzhangian)

Holm Dal Formation

Greenland

A sponge. The type species is S. erecta.

Sonarina[259]

Gen. et sp. nov

Valid

Taylor & Di Martino

Late Cretaceous (late Campanian or early Maastrichtian)

Kallankurichchi Formation

India

A cheilostome bryozoan belonging to the family Onychocellidae. Genus includes new species S. tamilensis.

Stanleycaris[188]

Gen. et sp. nov

Valid

Pates, Daley & Ortega-Hernández

Cambrian

Stephen Formation
Wheeler Formation

Canada
( British Columbia)
United States
( Utah)

A member of Radiodonta belonging to the group Hurdiidae. The type species is S. hirpex. The original description of the taxon appeared in an online supplement to the article published by Caron et al. (2010),[260] making in invalid until it was validated by Pates, Daley & Ortega-Hernández (2018).[187][188]

Styliolina langenii[234]

Sp. nov

Valid

Comniskey & Ghilardi

Devonian (middle to late Emsian)

Ponta Grossa Formation

Brazil

A member of Tentaculitoidea belonging to the order Dacryoconarida and the family Styliolinidae.

Sullulika[228]

Gen. et sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A selkirkiid stem-priapulid. Genus includes new species S. broenlundi.

Tallitaniqa[256]

Gen. et sp. nov

Valid

Peel

Cambrian (Guzhangian)

Holm Dal Formation

Greenland

A sponge. The type species is T. petalliformis.

Tarimspira artemi[261]

Sp. nov

Valid

Peel

Cambrian Stage 4

Henson Gletscher Formation

Greenland

An animal of uncertain phylogenetic placement described on the basis of fossil sclerites, possibly representing a stage in paraconodont evolution prior to the development of a basal cavity.

Tentaculites kozlowskii[234]

Sp. nov

Valid

Comniskey & Ghilardi

Devonian (late Pragian or late Emsian)

Ponta Grossa Formation

Brazil

A member of Tentaculitoidea belonging to the order Tentaculitida and the family Tentaculitidae.

Tentaculites paranaensis[234]

Sp. nov

Valid

Comniskey & Ghilardi

Devonian (late Pragian or late Emsian)

Ponta Grossa Formation

Brazil

A member of Tentaculitoidea belonging to the order Tentaculitida and the family Tentaculitidae.

Thanahita[262]

Gen. et sp. nov

Siveter et al.

Silurian (Wenlock)

Herefordshire Lagerstätte

United Kingdom.

A relative of Hallucigenia. The type species is T. distos.

Trapezovitus malinkyi[228]

Sp. nov

Valid

Peel & Willman

Cambrian Series 2

Buen Formation

Greenland

A member of Hyolitha.

Turbicellepora yasuharai[35]

Sp. nov

Valid

Di Martino & Taylor

Holocene

Indonesia

A bryozoan belonging to the group Cheilostomata and the family Celleporidae.

Uniconus ciguelii[234]

Sp. nov

Valid

Comniskey & Ghilardi

Devonian (late Pragian or late Emsian)

Ponta Grossa Formation

Brazil

A member of Tentaculitoidea belonging to the order Tentaculitida and the family Uniconidae.

Zardinisoma[236]

Gen. et 5 sp. nov

Valid

Wendt

Permian (Wordian) to Triassic (Carnian)

San Cassiano Formation

Italy
Japan

An ascidian belonging to the new order Khmeriamorpha. The type species is Z. cassianum; genus also includes Z. japonicum, Z. pauciplacophorum, Z. pyriforme and Z. polyplacophorum.

Zhijinites tumourifomis[263]

Sp. nov

Valid

Pan, Feng & Chang

Cambrian (Terreneuvian)

Yanjiahe Formation

China

A small shelly fossil.

Foraminifera

Research

  • A study on the effects of differential ocean acidification at the Cretaceous-Paleocene transition on the planktonic foraminiferal assemblages from the Farafra Oasis (Egypt) is published by Orabi et al. (2018).[264]
  • A wide variety of morphological abnormalities in planktic foraminiferal tests from the earliest Danian, mainly from Tunisian sections, is described by Arenillas, Arz & Gilabert (2018).[265]
  • A study on the impact of the climatic and environmental perturbation on the morphology of foraminifera living during the Paleocene–Eocene Thermal Maximum is published by Schmidt et al. (2018).[266]
  • Taxonomic compilation and partial revision of early Eocene deep-sea benthic Foraminifera is presented by Arreguín-Rodríguez et al. (2018).[267]
  • A study on the responses of two species of foraminifera (extant Truncorotalia crassaformis and extinct Globoconella puncticulata) to climate change during the late Pliocene to earliest Pleistocene intensification of Northern Hemisphere glaciation (3.6–2.4 million years ago) is published by Brombacher et al. (2018).[268]

New taxa

Name Novelty Status Authors Age Type locality Country Notes

Alabamina heyae[269]

Sp. nov

Valid

Fox et al.

Oligocene

Germany

A member of Rotaliida belonging to the family Alabaminidae.

Alicantina[270]

Gen. et comb. nov

Valid

Soldan, Petrizzo & Silva

Eocene

Dunghan Formation
Langley Formation
Lizard Springs Formation
Navet Formation
Richmond Formation
Shaheed Ghat Formation
Thebes Formation
Universidad Formation

Cuba
Egypt
Italy
Jamaica
Pakistan
Spain
Syria
Trinidad and Tobago
Tunisia
Atlantic Ocean
Indian Ocean
(Kerguelen Plateau)
Pacific Ocean
(Caroline Abyssal Plain
Shatsky Rise)

A member of the family Globigerinidae. The type species is "Globigerina" lozanoi Colom (1954); genus also includes "Globigerina" prolata Bolli (1957).

Ammobaculites deflectus[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

Ammobaculites knorringensis[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

Ammobaculoides dhrumaensis[272]

Sp. nov

Valid

Kaminski, Malik & Setoyama

Middle Jurassic (Bajocian)

Dhruma Formation

Saudi Arabia

A member of Lituolida belonging to the family Spiroplectamminidae.

Asterigerinella jonesi[273]

Sp. nov

Valid

Rögl & Briguglio

Miocene (Burdigalian)

Quilon Formation

India

Brizalina keralensis[273]

Sp. nov

Valid

Rögl & Briguglio

Miocene (Burdigalian)

Quilon Formation

India

Chiloguembelina adriatica[274]

Sp. nov

Valid

Premec Fucek, Hernitz Kucenjak & Huber

Eocene and Oligocene

Cipero Formation

Cuba
Syria
Trinidad and Tobago
Adriatic Sea
Gulf of Mexico
Pacific Ocean
(Ontong Java Plateau)

A member of Guembelitrioidea belonging to the family Chiloguembelinidae.

Chiloguembelina andreae[274]

Sp. nov

Valid

Premec Fucek, Hernitz Kucenjak & Huber

Late Eocene and early Oligocene

France
Syria
United States
( New Jersey)

A member of Guembelitrioidea belonging to the family Chiloguembelinidae.

Ciperoella[275]

Gen. et comb. nov

Valid

Olsson & Hemleben in Olsson et al.

Late Eocene to early Miocene

Cipero Formation
Tingnaro Formation

Australia
Austria
Belgium
Colombia
Cuba
France
Italy
Malta
Romania
Spain
Tanzania
Trinidad and Tobago
United States
( Mississippi)
Venezuela
Atlantic Ocean
Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae. The type species is "Globigerina" ciperoensis Bolli (1954); genus also includes "Globigerina" anguliofficinalis Blow (1969), "Globigerina ciperoensis" angulisuturalis Bolli (1957) (raised to the rank of the species Ciperoella angulisuturalis) and "Globigerina" fariasi Bermúdez (1961).

Colominella piriniae[276]

Sp. nov

Valid

Mancin & Kaminski

Pliocene

Italy

A member of Textulariida.

Cyclammina saidovae[277]

Nom. nov

Valid

Hanagata

Neogene

Japan

A species of Cyclammina; a replacement name for Cyclammina pseudopusilla Hanagata (2003).

Dentoglobigerina eotripartita[278]

Sp. nov

Valid

Pearson, Wade & Olsson in Wade et al.

Eocene and Oligocene

Navet Formation

Indonesia
Tanzania
Trinidad and Tobago
United States
( Mississippi)
Adriatic Sea

A member of Globigerinoidea belonging to the family Globigerinidae.

Douglassites[279]

Gen. et sp. nov

Valid

Read & Nestell

Carboniferous (late Pennsylvanian)

Riepe Spring Limestone

United States
( Nevada)

A member of Fusulinida belonging to the family Schubertellidae. Genus includes new species D. sprucensis.

Elazigina siderea[280]

Sp. nov

Valid

Consorti & Rashidi

Late Cretaceous (Maastrichtian)

Tarbur Formation

Iran
Oman
Turkey

A member of the group Rotaliida belonging to the family Rotaliidae.

Globigerina archaeobulloides[281]

Sp. nov

Valid

Hemleben & Olsson in Spezzaferri et al.

Oligocene

Shubuta Formation

United States
( Alabama)

A species of Globigerina.

Globigerinella roeglina[281]

Sp. nov

Valid

Spezzaferri & Coxall in Spezzaferri et al.

Oligocene, possibly Miocene

Romania
Gulf of Mexico
Indian Ocean

A member of Globigerinoidea belonging to the family Globigerinidae.

Globigerinoides joli[282]

Sp. nov

Valid

Spezzaferri in Spezzaferri, Olsson & Hemleben

Miocene

Caribbean Sea
Gulf of Mexico
South Atlantic Ocean
Indian Ocean
(Kerguelen Plateau)

A species of Globigerinoides.

Globigerinoides neoparawoodi[282]

Sp. nov

Valid

Spezzaferri in Spezzaferri, Olsson & Hemleben

Miocene

North-western Pacific Ocean

A species of Globigerinoides.

Globoconella pseudospinosa[283]

Sp. nov

Valid

Crundwell

Early Pliocene

Southwest Pacific Ocean

Globorotaloides atlanticus[284]

Sp. nov

Valid

Spezzaferri & Coxall

Oligocene and Miocene

Atlantic Ocean
Indian Ocean
Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae.

Globoturborotalita eolabiacrassata[285]

Sp. nov

Valid

Spezzaferri & Coxall in Spezzaferri et al.

Eocene to Miocene

Belgium
France
Romania
Tanzania
United States
( New Jersey)
Atlantic Ocean
Indian Ocean
(Kerguelen Plateau)
Pacific Ocean
(Nazca Plate)

A member of Globigerinoidea belonging to the family Globigerinidae.

Globoturborotalita paracancellata[285]

Sp. nov

Valid

Olsson & Hemleben in Spezzaferri et al.

Eocene and Oligocene

Western Atlantic Ocean
Gulf Coast of the United States

A member of Globigerinoidea belonging to the family Globigerinidae.

Globoturborotalita pseudopraebulloides[285]

Sp. nov

Valid

Olsson & Hemleben in Spezzaferri et al.

Oligocene and Miocene

Australia
Austria
Tanzania
Trinidad and Tobago
Gulf of Mexico
South Atlantic Ocean
Western equatorial Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae.

Haplophragmoides perlobatus[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

Hemisphaerammina apta[286]

Sp. nov

Valid

McNeil & Neville

Early Eocene

Beaufort Sea

A member of the order Astrorhizida and the suborder Hemisphaeramminineae.

Ichnusella senerae[287]

Sp. nov

Valid

Rigaud, Schlagintweit & Bucur

Early Cretaceous (Barremian–early Aptian)

Austria
France
Italy
Romania
Turkey
Croatia?
Serbia?
Ukraine?

A member of the group Spirillinida belonging to the family Spirillinidae.

Labrospira lenticulata[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

Lenticulina stewarti[269]

Sp. nov

Valid

Fox et al.

Oligocene (Rupelian)

Germany

A member of the group Nodosarioidea belonging to the family Vaginulinidae.

Moulladella[288]

Gen. et sp. nov

Valid

Bucur & Schlagintweit

Early Cretaceous (Valanginian-Barremian)

Austria
Bulgaria
France
Romania
Serbia
Spain

A member of Loftusiida belonging to the family Pfenderinidae. The type species is "Meyendorffina (Paracoskinolina)" jourdanensis Foury & Moullade (1966).

Neodubrovnikella[289]

Gen. et sp. nov

Valid

Schlagintweit & Rashidi

Late Cretaceous (Maastrichtian)

Tarbur Formation

Iran

Genus includes new species N. maastrichtiana.

Neonavarella[290]

Gen. et sp. nov

Valid

Giusberti, Kaminski & Mancin

Paleocene (Thanetian)

Scaglia Rossa Formation

Italy

A member of Lituolida belonging to the family Ammobaculinidae. The type species is N. sudalpina.

Neotrocholina theodori[287]

Sp. nov

Valid

Rigaud, Schlagintweit & Bucur

Early Cretaceous (Barremian–early Aptian)

Austria
France
Iran
Poland
Romania
Turkey

A member of the group Spirillinida belonging to the family Spirillinidae.

Nonion cepa[269]

Sp. nov

Valid

Fox et al.

Late Oligocene to early Miocene

Central North Sea basin
Netherlands

A member of the group Rotaliida belonging to the family Nonionidae.

Nummulites fayumensis[291]

Sp. nov

Valid

Al Menoufy & Boukhary

Eocene (Lutetian)

Egypt

A nummulite.

Nummulites tenuissimus[291]

Sp. nov

Valid

Al Menoufy & Boukhary

Eocene (Lutetian)

Egypt

A nummulite.

Omphalocyclus macroporus ellipsoides[292]

Subsp. nov

Valid

Al Nuaimy

Late Cretaceous (Maastrichtian)

Aqra Formation

Iraq

Omphalocyclus macroporus maukabensis[292]

Subsp. nov

Valid

Al Nuaimy

Late Cretaceous (Maastrichtian)

Aqra Formation

Iraq

Palaeoelphidium[293]

Gen. et comb. nov

Valid

Consorti, Schlagintweit & Rashidi

Late Cretaceous (Maastrichtian)

Iran
Iraq
Qatar

A member of the family Elphidiellidae; a new genus for "Elphidiella" multiscissurata Smout (1955).

Paralachlanella[273]

Gen. et sp. nov

Valid

Rögl & Briguglio

Miocene (Burdigalian)

Quilon Formation

India

Genus includes new species P. pilleri.

Pseudomassilina quilonensis[273]

Sp. nov

Valid

Rögl & Briguglio

Miocene (Burdigalian)

Quilon Formation

India

Pseudopeneroplis[294]

Gen. et sp. nov

Valid

Consorti in Consorti et al.

Late Cretaceous (late Cenomanian)

Peru

A member of the superfamily Soritoidea and the family Praerhapydioninidae. Genus includes new species P. oyonensis.

Quiltyella[281]

Gen. et comb. nov

Valid

Coxall & Spezzaferri in Spezzaferri et al.

Oligocene and Miocene

Austria
Romania
East Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae. The type species is "Clavigerinella" nazcaensis Quilty (1976); genus also includes "Hastigerinella" clavacella Rögl (1969).

Ranikothalia daviesi[295]

Sp. nov

Valid

Sirel & Deveciler

Early Eocene

Turkey

A member of the group Rotaliida belonging to the family Nummulitidae.

Reophax pyriloculus[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

Schubertella luisorum[296]

Sp. nov

Valid

Villa in Villa, Merino-Tomé & Martín Llaneza

Carboniferous (Moscovian)

La Nueva Limestone
Meruxalín Limestone
Sutu Limestone

Spain

A member of Fusulinida.

Streptochilus tasmanensis[297]

Sp. nov

Valid

Smart & Thomas

Oligocene

South Tasman Rise

A member of Bolivinoidea belonging to the family Bolivinidae.

Subbotina projecta[298]

Sp. nov

Valid

Olsson, Pearson & Wade in Wade et al.

Late Eocene and Oligocene

Yazoo Formation

Tanzania
United States
( Alabama
Mississippi)
Atlantic Ocean
Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae.

Textularia pernana[271]

Sp. nov

Valid

Hjalmarsdottir, Nakrem & Nagy

Late Jurassic - Early Cretaceous

Agardhfjellet Formation

Norway

A species of Textularia.

Trilobatus altospiralis[282]

Sp. nov

Valid

Spezzaferri in Spezzaferri, Olsson & Hemleben

Miocene

South Pacific Ocean

A member of Globigerinoidea belonging to the family Globigerinidae.

Trochammina jakovlevae[299]

Sp. nov

Valid

Glinskikh & Nikitenko

Middle Jurassic (late Bajocian-early Bathonian)

Churkino Formation

Russia

A member of the family Trochamminidae.

Uvigerina kingi[269]

Sp. nov

Valid

Fox et al.

Middle Miocene

Netherlands
Southern and central North Sea

A member of the group Rotaliida belonging to the family Uvigerinidae.

Other organisms

Research

  • A study on putative stromatolites described from the 3,700-Myr-old rocks from the Isua supracrustal belt (Greenland) by Nutman et al. (2016)[300] is published by Allwood et al. (2018), who interpret these putative stromatolites as more likely to be structures of non-biological origin.[301]
  • Carbon isotope analyses of 11 microbial fossils from the ~3,465-million-year-old Apex chert (Australia) are published by Schopf et al. (2018), who interpret two of the five studied species as primitive photosynthesizers, one as an Archaeal methane producer, and two as methane consumers.[302]
  • The oldest well-preserved microbial mats fabrics are described from the ≈3,472-million-year-old Middle Marker horizon, Barberton Greenstone Belt (South Africa) by Hickman-Lewis et al. (2018).[303]
  • Direct fossil evidence for life on land 3,220 million years ago in the form of terrestrial microbial mats is reported from the Moodies Group (South Africa) by Homann et al. (2018).[304]
  • Microfossils representing 18 morphotypes are reported from the c. 2.4 billion years old Turee Creek Group (Western Australia) by Barlow & Van Kranendonk (2018).[305]
  • Ten representative types of exceptionally well-preserved mat-related structures, interpreted as likely to be of biological origin and including putative microbial mats and discoidal microbial colonies, are reported from the 2.1-billion-year-old Francevillian series in Gabon by Aubineau et al. (2018).[306]
  • A study on the chemical, isotopic and molecular structural characteristics of the putative multicellular eukaryote fossils from carbonaceous compressions in the 1.63 billion years old Tuanshanzi Formation (China) is published by Qu et al. (2018).[307]
  • Intact porphyrins, the molecular fossils of chlorophylls, are described from 1,100-million-year-old marine black shales of the Taoudeni Basin (Mauritania) by Gueneli et al. (2018), who also study the nitrogen isotopic values of the fossil pigments, and interpret their findings as indicating that the oceans of that time were dominated by cyanobacteria, while larger planktonic algae were scarce.[308]
  • A study on the evolutionary history of bacteria is published by Louca et al. (2018), who interpret their findings as indicating that most bacterial lineages ever to have inhabited Earth are extinct.[309]
  • Bobrovskiy et al. (2018) report molecular fossils from organically preserved specimens of Beltanelliformis, and interpret the fossils as representing large spherical colonies of cyanobacteria.[310]
  • Discoid imprints sampled from the Precambrian terranes of central Dobruja (Romania) are described and assigned to the species Beltanelliformis brunsae by Saint Martin & Saint Martin (2018).[311]
  • A study on the age of the fossil red alga Bangiomorpha pubescens is published by Gibson et al. (2018).[312]
  • A reassessment of the anatomy and taxonomy of Orbisiana, based on a restudy of the rediscovered original type material of O. simplex, is published by Kolesnikov et al. (2018).[313]
  • A study on the positions of fossil specimens in the assemblages of Ediacaran fossils from Mistaken Point (Canada), as well as on their implications for inferring the interactions and associations between the Ediacaran organisms, is published by Mitchell & Butterfield (2018).[314]
  • A study on the height of Ediacaran organisms from Mistaken Point, evaluating the link between the increase of height and resource competition or greater offspring dispersal, is published by Mitchell & Kenchington (2018).[315]
  • Evidence of a radiation of the Ediacaran biota that witnessed the emergence and widespread implementation of novel, animal-style ecologies is presented by Tarhan et al. (2018), who argue that this transition was linked to the expansion of Ediacaran taxa into dynamic, shallow marine environments characterized by episodic disturbance and complex and diverse organically bound substrates, and propose that younger, second-wave Ediacaran communities resulting from said radiation were part of an ecological and evolutionary continuum with Phanerozoic ecosystems.[316]
  • A study on the size range, ontogeny and palaeoenvironment of Rugoconites is published by Hall, Droser & Gehling (2018).[317]
  • Elliptical body fossils are described from the Ediacaran–Fortunian deposits of central Brittany (France) by Néraudeau et al. (2018), representing the first body fossils described from these deposits.[318]
  • A study on the sandstone- and limestone-hosted occurrences of Palaeopascichnus linearis (including material from a new locality in Arctic Siberia), indicative of a greater range of taxonomic and taphonomic variation, is published by Kolesnikov et al. (2018).[319]
  • A study on the organic-walled microfossils from the Cambrian strata in the stratotype section of the Precambrian–Cambrian boundary in the Burin Peninsula (Canada) is published by Palacios et al. (2018).[320]
  • Fossils interpreted as threads of filamentous cyanobacteria are described from the Cambrian (Guzhangian) Alum Shale Formation (Sweden) by Castellani et al. (2018).[321]
  • Enigmatic Devonian taxon Protonympha is interpreted as a possible post-Ediacaran vendobiont by Retallack (2018).[322]
  • Description of fossils of nonmarine diatoms belonging to the genus Actinocyclus from the Lower to Middle Miocene lacustrine deposits in Japan and a study on the possible causal links between the evolution of nonmarine planktonic diatoms and the climatic and environmental changes that occurred during the Miocene is published by Hayashi et al. (2018).[323]
  • A study on the cell-size frequency distributions across calcareous nanoplankton communities through the Paleocene–Eocene Thermal Maximum, on their population biomass and on the impact of climate change on their cellular characteristics is published by Gibbs et al. (2018).[324]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Alievium mangalensiense[325]

Sp. nov

Valid

Bragina & Bragin

Late Cretaceous

Cyprus

A radiolarian belonging to the family Pseudoaulophacidae.

Angochitina plicata[326]

Sp. nov

Valid

Noetinger, di Pasquo & Starck

Devonian

Argentina

A chitinozoan.

Anhuithrix[327]

Gen. et comb. nov

Pang et al.

Tonian

Liulaobei Formation

China

A member of Cyanobacteria; a new genus for "Omalophyma" magna Steiner (1994).

Attenborites[328]

Gen. et sp. nov

Valid

Droser et al.

Ediacaran

Rawnsley Quartzite

Australia

An organism of uncertain phylogenetic placement, described on the basis of a well-defined irregular oval to circular fossil. Genus includes new species A. janeae. Announced in 2018; the final version of the article naming it was published in 2020.

Cyclotella cassandrae[329]

Sp. nov

Valid

Paillès et al.

Pleistocene

Guatemala

A diatom.

Cyclotella petenensis[329]

Sp. nov

Valid

Paillès et al.

Pleistocene

Guatemala

A diatom.

Doulia[330]

Gen. et sp. nov

Valid

Lian et al.

Cambrian Stage 3

Hongjingshao Formation

China

A possible planktonic alga of uncertain phylogenetic placement. Genus includes new species D. rara.

Eolaminaria simigladiola[330]

Sp. nov

Valid

Lian et al.

Cambrian Stage 3

Hongjingshao Formation

China

A macroalga of uncertain phylogenetic placement.

Epistacheoides bozorgniai[331]

Sp. nov

Valid

Falahatgar, Vachard & Sarfi

Carboniferous (Viséan)

Iran

An alga of uncertain phylogenetic placement.

Girvanella lianiformis[332]

Sp. nov

Valid

Peel

Cambrian (Drumian)

Ekspedition Bræ Formation

Greenland

A member of Cyanobacteria belonging to the family Cyanophyceae.

Girvanella pituutaq[332]

Sp. nov

Valid

Peel

Cambrian (Drumian)

Ekspedition Bræ Formation

Greenland

A member of Cyanobacteria belonging to the family Cyanophyceae.

Gorgonisphaeridium impexus[326]

Sp. nov

Valid

Noetinger, di Pasquo & Starck

Devonian

Argentina

An acritarch.

Hylaecullulus[333]

Gen. et sp. nov

Valid

Kenchington, Dunn & Wilby

Ediacaran

United Kingdom

A rangeomorph. The type species is H. fordi.

Leiosphaeridia gorda[334]

Sp. nov

Valid

Loron & Moczydłowska

Tonian

Visingsö Group
Wynniatt Formation

Canada
Sweden

A unicellular microorganism of algal affinities.

Lontohystrichosphaera[250]

Gen. et sp. nov

Valid

Slater, Harvey & Butterfield

Cambrian (Terreneuvian)

Lontova Formation

Estonia

A large ornamented acritarch of unresolved biological affinity, probably an ontogenetically and metabolically active eukaryotic organism rather than a dormant protistan cyst. Genus includes new species L. grandis.

Mallomonas aperturae[335]

Sp. nov

Valid

Siver

Middle Eocene

Giraffe Pipe locality

Canada

A synurid, a species of Mallomonas.

Mallomonas bakeri[336]

Sp. nov

Valid

Siver

Middle Eocene

Giraffe Pipe locality

Canada

A synurid, a species of Mallomonas.

Mallomonas skogstadii[336]

Sp. nov

Valid

Siver

Middle Eocene

Giraffe Pipe locality

Canada

A synurid, a species of Mallomonas.

Mispertonia[337]

Gen. et sp. nov

Valid

McLean et al.

Carboniferous (Mississippian) to Late Permian or Early Triassic

India
United Kingdom

An organic-walled microfossil of uncertain phylogenetic placement. Genus includes new species M. desiccata.

Obamus[338]

Gen. et sp. nov

Valid

Dzaugis et al.

Ediacaran

Rawnsley Quartzite

Australia

A torus-shaped organism, similar in gross morphology to some poriferans and benthic cnidarians. Genus includes new species O. coronatus. Announced in 2018; the final version of the article naming it was published in 2020.

Orpikania[332]

Gen. et sp. nov

Valid

Peel

Cambrian (Drumian)

Ekspedition Bræ Formation

Greenland

A member of the family Epiphytaceae (a group of organisms of uncertain phylogenetic placement). Genus includes new species O. freucheni.

Pakupaku[339]

Gen. et sp. nov

Valid

Riedman, Porter & Calver

Tonian

Black River Dolomite

Australia

A vase-shaped microfossil. Genus includes new species P. kabin.

Pierceites deccanensis[340]

Sp. nov

Valid

Prasad et al.

Late Cretaceous (Maastrichtian)

India

A dinoflagellate belonging to the family Peridiniaceae.

Pseudoalievium[325]

Gen. et 2 sp. nov

Valid

Bragina & Bragin

Late Cretaceous

Cyprus

A radiolarian belonging to the family Pseudoaulophacidae. Genus includes new species P. parekklisiense and P. inflatum.

Pseudoaulophacus decoratus[325]

Sp. nov

Valid

Bragina & Bragin

Late Cretaceous

Cyprus

A radiolarian belonging to the family Pseudoaulophacidae.

Retiranus[250]

Gen. et sp. nov

Valid

Slater, Harvey & Butterfield

Cambrian (Terreneuvian)

Lontova Formation
Voosi Formation

Estonia
Lithuania

A sheet-like or funnel-shaped organism of unresolved biological affinity. Genus includes new species R. balticus.

Rugophyca[330]

Gen. et sp. nov

Valid

Lian et al.

Cambrian Stage 3

Hongjingshao Formation

China

A macroalga of uncertain phylogenetic placement. Genus includes new species R. longa.

Saarinomorpha[341]

Gen. et sp. nov

Valid

Kolosov & Sofroneeva

Vendian

Russia

A tubiform organic-walled segmented microfossil, resembling Saarina juliae but smaller by one–two orders of magnitude. Genus includes new species S. infundibularis.

Singulariphyca[330]

Gen. et sp. nov

Valid

Lian et al.

Cambrian Stage 3

Hongjingshao Formation

China

A macroalga of uncertain phylogenetic placement. Genus includes new species S. ramosa.

Stellarossica[342]

Gen. et comb. nov

Valid

Vorob'eva & Sergeev

Precambrian

Ura Formation

Russia

A large acanthomorph acritarch. Genus includes new species S. ampla.

Synsphaeridium parahioense[343]

Sp. nov

Valid

Yin et al.

Cambrian Series 3

India

An acritarch.

Tristratothallus[344]

Gen. et sp. nov

Valid

Edwards et al.

Silurian (Ludfordian)

Downton Castle Sandstone Formation

United Kingdom

A nematophyte belonging to the family Nematothallaceae. Genus includes new species T. ludfordensis.

Vendotaenia pavimentpes[345]

Sp. nov

Valid

Yang & Qin in Yang et al.

Ediacaran

Dengying Formation

China

An alga.

Vendotaenia sixiense[345]

Sp. nov

Valid

Yang & Qin in Yang et al.

Ediacaran

Dengying Formation

China

An alga.

History of life in general

Other research

References

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