2021 in arthropod paleontology

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2021 in arthropod paleontology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2021.

New taxa

Name Novelty Status Authors Age Type locality Location Notes Images

Allochthonius balticus[1]

Sp. nov

In press

Schwarze et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A pseudoscorpion belonging to the family Pseudotyrannochthoniidae.

Alterphyxioschemoides[2]

Gen. et sp. nov

Valid

Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

Possibly a member of the family Dipluridae. The type species is A. spicula.

Alticorona[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of the family Tetrablemmidae. The type species is A. plenfemur.

Autotomiana brevisetosa[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Pholcochyroceridae.

Barbutia theroni[3]

Sp. nov

Valid

Khaustov et al.

Late Eocene

Rovno amber

Ukraine

A mite belonging to the group Raphignathoidea and the family Barbutiidae.

Boavista[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae. The type species is B. crassifemora.

Burmadictyna crassembolus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the family Salticoididae.

Burmadictyna fissura[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the family Salticoididae.

Burmadictyna similis[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the family Salticoididae.

Burmaspiralis[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Araneoidea and the family Zarqaraneidae. The type species is B. trispinae.

Burmesiana[2]

Nom. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Nemesiidae; a replacement name for Burmesia Wunderlich (2020).

Burmorchestina bo[4]

Sp. nov

Valid

Jiang & Li in Xin et al.

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Oonopidae.

Burmorchestina convexa[4]

Sp. nov

Valid

Jiang & Li in Xin et al.

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Oonopidae.

Burmorsolus longitibia[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Burmorsolidae.

Centrochthonius bitterfeldicus[1]

Sp. nov

In press

Schwarze et al.

Eocene

Bitterfeld amber

Germany

A pseudoscorpion belonging to the family Pseudotyrannochthoniidae.

Chaerilobuthus meggeri[5]

Sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Burmese amber

Myanmar

A scorpion belonging to family Chaerilobuthidae.

Cornutheridion[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Theridiidae. The type species is C. concavum.

Crassicephalus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Crassicephalidae. The type species is C. parvibulbus.

Crassitibia sicilicula[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Araneoidea and the family Zarqaraneidae.

Cretaceousopisthacanthus[6]

Gen. et sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Burmese amber

Myanmar

A scorpion belonging to the family Protoischnuridae. The type species is C. smeelei.

Cretapalpus[7]

Gen. et sp. nov

Valid

Downen & Selden

Early Cretaceous

Crato Formation

Brazil

A spider belonging to the family Palpimanidae. Genus includes new species C. vittari.

Dolichocybe elongata[8]

Sp. nov

Valid

Khaustov et al.

Late Eocene

Rovno amber

Ukraine

A mite belonging to the group Heterostigmata and the family Dolichocybidae.

Dubiodeinopsis[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Dubiodeinopsidae. The type species is D. spinifemora.

Dubiouloborix[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Dubiouloboridae. The type species is D. incompletus.

Dubiouloborus[2]

Gen. et 2 sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Dubiouloboridae. The type species is D. praeta; genus also includes D. procerembolus.

Electroblemma spermaferens[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of the family Tetrablemmidae.

Eoagelenomorphus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider of uncertain phylogenetic placement, possibly a member of an early branch of the RTA clade. The type species is E. cretaceus.

Furcembolus chuandiani[9]

Sp. nov

Valid

Jiang & Li in Xin et al.

Late Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Pacullidae.

Furcembolus fengzheni[9]

Sp. nov

Valid

Jiang & Li in Xin et al.

Late Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Pacullidae.

Furcembolus martensi[9]

Sp. nov

Valid

Jiang & Li in Xin et al.

Late Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Pacullidae.

Hiatomegops[10]

Gen. et sp. nov

Valid

Guo, Selden & Ren

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Lagonomegopidae. Genus includes new species H. spinalis.

Hoplocheylus neosimilis[8][3]

Sp. nov

Valid

Khaustov et al.

Late Eocene

Rovno amber

Ukraine

A mite belonging to the group Heterostigmata and the family Tarsocheylidae.

Hungarosilia[11]

Gen. et sp. nov

In press

Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

Hungary

A spider belonging to the family Hersiliidae. Genus includes new species H. verdesi.

Kachinarachne[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Pholcochyroceridae. The type species is K. oblonga.

Levantoglyphus[12]

Gen. et sp. nov

Valid

Klimov et al.

Early Cretaceous (Barremian)

Lebanese amber

Lebanon

A mite belonging to the group Astigmata. The type species is L. sidorchukae.

Liacarus (Procorynetes) shtanchaevae[13]

Sp. nov

In press

Arillo & Subías in Arillo, Subías & Álvarez-Parra

Early Cretaceous (Albian)

Escucha Formation

Spain

A mite belonging to the family Liacaridae.

Longissipalpus cochlea[2]

Sp. nov

Junior homonym

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Pholcochyroceridae. The specific name is preoccupied Longissipalpus cochlea Wunderlich (2017). Wunderlich (2022) coined a replacement name Longissipalpus aliter.[14]

Longissipalpus impudicus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Pholcochyroceridae.

Megasetae[2]

Gen. et sp. nov

Valid

Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the new family Megasetidae. The type species is M. colphepeiroides.

Metaphalangium martensi[15]

Sp. nov

Valid

Mitov, Perkovsky & Dunlop

Eocene

Rovno amber

Ukraine

A harvestman, a species of Metaphalangium.

Micropalpimanus gibber[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Palpimanoidea and the family Micropalpimanidae.

Microtheridion[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Theridiidae. The type species is M. longissispinae.

Microuloborus oblongus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Neoveterator[16]

Nom. nov

Ceccolini & Cianferoni

Miocene

Mexican amber

Dominican Republic
Mexico

A spider belonging to the family Trochanteriidae; a replacement name for Veterator Petrunkevitch (1963).

Orchestina globus[4]

Sp. nov

Valid

Jiang & Li in Xin et al.

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Oonopidae, a species of Orchestina.

Palaeodiplothrombidium[17]

Gen. et sp. nov

In press

Rivas & Vega

Miocene

Mexican amber

Mexico

A mite belonging to the family Johnstonianidae. The type species is P. microscutum.

Palaeoleptoneta fissura[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Leptonetidae.

Palaeothele onoi[18]

Sp. nov

Valid

Selden

Carboniferous (Moscovian)

Mazon Creek fossil beds

United States
( Illinois)

A spider belonging to the group Mesothelae and to the new family Palaeothelidae.

Palaeozearchaea[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Mecysmaucheniidae. The type species is P. depressa.

Paradactylidium sineunguis[8]

Sp. nov

Valid

Khaustov et al.

Late Eocene

Rovno amber

Ukraine

A mite belonging to the group Heterostigmata and the family Acarophenacidae.

Parahistricostoma[15]

Gen. et comb. nov

Valid

Mitov, Perkovsky & Dunlop

Eocene

Baltic amber
Bitterfeld amber
Rovno amber

Germany
Ukraine

A harvestman belonging to the family Nemastomatidae; a new genus for "Nemastoma" tuberculatum Koch & Berendt (1854).

Paramiagrammopes appendix[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes curvatus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes furca[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes granulatus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes inaequalis[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes inclinatus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes multifemurspinae[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes paracurvatus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes pilosus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes pollex[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes semiapertus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes simplex[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes sulcus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes texter[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Paramiagrammopes unibrevispina[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Planarchaea incompleta[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Palpimanoidea and the family Planarchaeidae.

Platythelae[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Palpimanoidea and the family Planarchaeidae. The type species is P. longicorpus.

Praetervetiator[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Palpimanoidea and the family Vetiatoridae. The type species is P. circulus. The paper naming it also uses the spelling of the genus name Praetervetianus; both names refer to the same genus.[14]

Prionochthonius[19]

Gen. et sp. nov

Valid

Wriedt et al.

Cretaceous

Burmese amber

Myanmar

A pseudoscorpion belonging to the family Chthoniidae. Genus includes new species P. burmiticus.

Priscaleclercera christae[20]

Sp. nov

Valid

Magalhaes et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A spider belonging to the family Psilodercidae.

Proadactylidium fossibilis[8]

Sp. nov

Valid

Khaustov et al.

Late Eocene

Rovno amber

Ukraine

A mite belonging to the group Heterostigmata and the family Acarophenacidae.

Proalbiorix[21]

Gen. et 2 sp. nov

In press

Geißler et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A pseudoscorpion belonging to the family Ideoroncidae. Genus includes new species P. gracilis and P. compactus.

Proaraneoides lanceatum[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Leptonetoidea and the family Protoaraneoididae.

Procerclypeus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of the family Tetrablemmidae. The type species is P. deformans.

Procervetiator[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Palpimanoidea and the family Vetiatoridae. The type species is P. fruticosus.

Propterkachin bispinatus[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae.

Propterpsiloderces similis[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of Araneomorphae belonging to the family Eopsilodercidae.

Protolycosa suazoi[18]

Sp. nov

Valid

Selden

Carboniferous (Kasimovian)

Atrasado Formation

United States
( New Mexico)

A spider belonging to the group Mesothelae and the family Arthrolycosidae.

Pseudokachin[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae. The type species is P. tuberculatus.

Scopomegops[10]

Gen. et sp. nov

Valid

Guo, Selden & Ren

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Lagonomegopidae. Genus includes new species S. fax.

Scutuloborella[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. admirabilis.

Scutuloboroides[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. pumilio.

Scutuloborus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Deinopoidea and the new family Scutuloboridae. The type species is S. spiralembolus.

Spiniarchaea[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Archaeidae. The type species is S. aberrans.

Spinicymbium unispina[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the group Araneoidea and the family Zarqaraneidae.

Spiniuloborus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Uloboridae. The type species is S. crux.

Spinoburmesebuthus knodelorum[22]

Sp. nov

Valid

Lourenço

Cretaceous

Burmese amber

Myanmar

A scorpion belonging to the group Buthoidea and the family Palaeoburmesebuthidae.

?Telemofila ovalis[2]

Sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A spider belonging to the family Telemidae, possibly a species of Telemofila.

Tenuicephalus[2]

Gen. et sp. nov

Valid

Wunderlich in Wunderlich & Müller

Cretaceous

Burmese amber

Myanmar

A member of the family Tetrablemmidae. The type species is T. penicillus.

Research

  • Revision of the fossil record of whip spiders is published by Haug & Haug (2021).[23]
  • Guo, Selden & Ren (2021) describe an adult lagonomegopid female, part of an egg sac and lagonomegopid spiderlings preserved in four pieces of Burmese amber, and interpret this finding as evidence of maternal care in fossil spiders.[24]
  • An exuvium of a member of the genus Myrmecarchaea is described from the Eocene Cambay amber (India) by Wood, Singh & Grimaldi (2021), representing the first member of the family Archaeidae from Cambay amber reported to date.[25]
  • An indeterminate deutonymph belonging to the family Sejidae is described by Joharchi, Vorontsov and Walter (2021) from Burmese amber, which represents the oldest record of the parasitiform mite clade Mesostigmata.[26]

Crustaceans

New taxa

Malacostracans

Name Novelty Status Authors Age Type locality Country Notes Images

Agolambrus rathbunae[27]

Sp. nov

Valid

Schweitzer, Hyžný & Feldmann

Venezuela

A crab belonging to the family Parthenopidae.

Aptanacalliax[28]

Gen. et sp. nov

In press

Ferratges, Hyžný & Zamora

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of Axiidea belonging to the family Anacalliacidae. Genus includes new species A. enigma.

Aptaxiopsis[28]

Gen. et sp. nov

In press

Ferratges, Hyžný & Zamora

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of Axiidea. Genus includes new species A. longimanus.

Autrigoniscus[29]

Gen. et sp. nov

Valid

Sánchez-García et al.

Early Cretaceous (Albian)

Spanish amber

Spain

A member of Isopoda belonging to the family Trichoniscidae. The type species is A. resinicola.

Bajoprosopon[30]

Gen. et sp. nov

Valid

Van Bakel et al.

Middle Jurassic (Bajocian)

France

A crab belonging to the superfamily Homolodromioidea and the family Prosopidae. The type species is B. piardi.

Bavaricaris[31]

Gen. et sp. nov

Valid

Winkler

Late Jurassic (Tithonian)

Altmühltal Formation

Germany

A member of Caridea, possibly belonging to the family Palaemonidae. The type species is B. haereri.

Berglundus[32]

Gen. et sp. nov

In press

Nyborg et al.

Early to middle Miocene

Astoria Formation

United States
( Washington)

A crab belonging to the group Cyclodorippoida and the family Cyclodorippidae. The type species is B. bretoni.

Bericirinia[33]

Gen. et sp. nov

In press

De Angeli & Garassino

Eocene (Priabonian)

Italy

A crab belonging to the family Epialtidae. The type species is B. bretoni.

Blaculla anjobea[31]

Sp. nov

Valid

Winkler

Late Jurassic (Tithonian)

Altmühltal Formation

Germany

A member of Caridea.

Bucculentum arnosavelkouli[34]

Sp. nov

Valid

Wallaard et al.

Late Jurassic (Tithonian)

Mörnsheim Formation

Germany

A crab belonging to the group Homolodromioidea and the family Bucculentidae.

Bucculentum horstkuscheli[35]

Sp. nov

Valid

Krzemińska et al.

Late Jurassic (Kimmeridgian)

Germany

A crab belonging to the group Homolodromioidea and the family Bucculentidae.

Bucculentum plettenbergense[35]

Sp. nov

Valid

Krzemińska et al.

Late Jurassic (Oxfordian)

Germany

A crab belonging to the group Homolodromioidea and the family Bucculentidae.

Carpilius cantellii[36]

Sp. nov

Valid

De Angeli & Alberti

Eocene (Bartonian-Priabonian)

Italy

A crab, a species of Carpilius.

Cirolana aptiana[37]

Sp. nov

In press

Bruce et al.

Early Cretaceous (Aptian)

Sierra Madre Formation

Mexico

A member of Isopoda, a species of Cirolana.

Cirolana bretoni[37]

Sp. nov

In press

Bruce et al.

Early Cretaceous (Aptian)

Sierra Madre Formation

Mexico

A member of Isopoda, a species of Cirolana.

Cirolana longirostra[37]

Sp. nov

In press

Bruce et al.

Early Cretaceous (Aptian)

Sierra Madre Formation

Mexico

A member of Isopoda, a species of Cirolana.

Clampethildella[38]

Gen. et sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

Italy

A crab belonging to the family Mathildellidae. The type species is C. spinosa.

Collodes cumarebensis[27]

Sp. nov

Valid

Schweitzer, Hyžný & Feldmann

Venezuela

A crab belonging to the family Inachoididae.

Cretacocalcinus[28]

Gen. et sp. nov

In press

Ferratges, Hyžný & Zamora

Early Cretaceous (Aptian)

Forcall Formation

Spain

A hermit crab. Genus includes new species C. josaensis.

Cretapsara[39]

Gen. et sp. nov

Valid

Luque in Luque et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A crab belonging to the group Eubrachyura and to the new family Cretapsaridae. The type species is C. athanata.

Cristinagalathea[38]

Gen. et sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

Italy

A member of the family Munidopsidae. The type species is C. striata.

Crosniera forcallensis[28]

Sp. nov

In press

Ferratges, Hyžný & Zamora

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of Axiidea belonging to the family Callianideidae.

Cryptolacruma[40]

Gen. et sp. nov

Valid

Schädel et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Isopoda belonging to the group Epicaridea. The type species is C. nidis.

Daciapagurus suebiorum[41]

Sp. nov

Valid

Fraaije, Van Bakel & Jagt

Late Jurassic (Kimmeridgian)

Germany

A hermit crab belonging to the family Schobertellidae.

Electrolana[42]

Gen. et sp. nov

Disputed

Schädel, Hyžný & Haug

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Isopoda belonging to the group Cymothoida. The type species is E. madelineae. Bruce & Rodcharoen (2023) considered Electrolana to be a junior synonym of the genus Cirolana, though the authors maintained E. madelineae as a distinct species within the latter genus.[43]

Emplastron[44]

Gen. et comb. nov

Valid

O'Flynn, Audo & Kawai

Paleocene (Thanetian)

France

A member of the family Astacidae; a new genus for "Astacus" edwardsi Van Straelen (1928).

Eocalcinus gerardbretoni[45]

Sp. nov

Valid

Ferratges, Artal & Zamora

Eocene (Priabonian)

Arguis Formation

Spain

A hermit crab belonging to the family Calcinidae.

Eoinachoides bretoni[46]

Sp. nov

In press

Garassino, Feldmann & Schweitzer

Eocene (Bartonian)

Castle Hayne Limestone

United States
( North Carolina)

A crab belonging to the family Epialtidae.

Eoligiiscus[29]

Gen. et sp. nov

Valid

Sánchez-García et al.

Early Cretaceous (Albian)

Spanish amber

Spain

A member of Isopoda belonging to the family Ligiidae. The type species is E. tarraconensis.

Ethusa orgianensis[47]

Sp. nov

Valid

De Angeli & Garassino

Late Eocene

Italy

A crab, a species of Ethusa.

Gammaroidorum yooling[48]

Sp. nov

Valid

Wei et al.

Late Neogene

Xiaobai Formation

China

A member of Amphipoda.

Glypheopsis tubantiensis[49]

Sp. nov

Valid

Becker, Fraaije & Mulder

Early Cretaceous

Netherlands

A member of the family Glypheidae.

Harthofia heidenreichetfauseri[31]

Sp. nov

Valid

Winkler

Late Jurassic (Tithonian)

Altmühltal Formation

Germany

A member of Caridea.

Heraclitus[29]

Gen. et sp. nov

Valid

Sánchez-García et al.

Early Cretaceous (Albian)

Spanish amber

Spain

A member of Isopoda belonging to the group Crinocheta, possibly a member of the family Detonidae. The type species is H. helenae.

Homarus fami[50]

Sp. nov

Valid

Garassino, Pasini & Nyborg in Garassino et al.

Early Cretaceous (Albian)

Canada
( British Columbia)

A species of Homarus.

Hoploparia bretoni[51]

Sp. nov

In press

Feldmann & Schweitzer

Late Cretaceous (Campanian)

Point Loma Formation

United States
( California)

A lobster.

Jurastenopus[52]

Gen. et sp. nov

Valid

Winkler et al.

Late Jurassic (Tithonian)

Painten Formation

Germany

A member of Stenopodidea. The type species is J. frattigianii.

Linuparus qualitus[53]

Sp. nov

Valid

Feldmann et al.

Late Cretaceous (Turonian)

Kaskapau Formation

Canada
( British Columbia)

A species of Linuparus.

Mecochirus tuberculatus[50]

Sp. nov

Valid

Garassino, Pasini & Nyborg in Garassino et al.

Early Cretaceous (Albian)

Canada
( British Columbia)

A member of the family Mecochiridae.

Metanephrops serendipitus[54]

Sp. nov

Valid

Gašparič et al.

Early Miocene

Slovenia

A species of Metanephrops. Announced in 2021; validated in 2023.[55]

Meticonaxius gracilis[28]

Sp. nov

In press

Ferratges, Hyžný & Zamora

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of Axiidea.

Meyeria libanotica[56]

Sp. nov

Valid

Charbonnier et al.

Early Cretaceous (Barremian)

Lebanon

A member of the family Mecochiridae.

Miocrangon[57]

Gen. et sp. nov

Valid

Garassino, Pasini & Nazarkin

Miocene

Russia
( Sakhalin Oblast)

A member of the family Crangonidae. Genus includes new species M. tartaricus.

Mothocya vallecillae[58]

Sp. nov

In press

Stinnesbeck et al.

Late Cretaceous (Turonian)

Mexico

A member of Isopoda belonging to the family Cymothoidae.

Muellermunida[38]

Gen. et comb. nov

Valid

Beschin, Busulini & Tessier

Eocene

Szépvölgy Formation

Hungary
Italy

A member of the family Munididae. The type species is "Protomunida" pentacantha Müller & Collins (1991); genus also includes "Eumunida" veronensis Beschin, Busulini & Tessier in Beschin et al. (2019).

Nahecaris sabineae[59]

Sp. nov

Valid

Poschmann

Devonian (Emsian)

Germany

A member of Phyllocarida.

Orgianocarcinus[33]

Gen. et sp. nov

In press

De Angeli & Garassino

Eocene (Priabonian)

Italy

A crab belonging to the family Dairidae. The type species is O. bericus.

Palaeastacus haidaensis[50]

Sp. nov

Valid

Garassino, Pasini & Nyborg in Garassino et al.

Early Cretaceous (Albian)

Canada
( British Columbia)

A member of the family Erymidae.

Paradasygyius rodriguezi[60]

Sp. nov

Valid

Lima, Aguilera & Tavares

Late Miocene

Venezuela

A spider crab belonging to the family Inachoididae.

Parapetrochirus[45]

Gen. et sp. nov

Valid

Ferratges, Artal & Zamora

Eocene (Ypresian)

Roda Formation

Spain

A hermit crab belonging to the family Diogenidae. The type species is P. robustus.

Paromola bretoni[61]

Sp. nov

Valid

Ferratges, Domínguez & Ossó

Eocene (Ypresian)

Roda Formation

Spain

A crab belonging to the family Homolidae.

Paronapaguropsis[62]

Gen. et sp. nov

Valid

Beschin et al.

Late Eocene

Italy

A hermit crab. Genus includes new species P. scaligera.

Parvucymoides[63]

Gen. et sp. nov

Valid

Van der Wal et al.

Late Eocene

Czech Republic

An isopod belonging to the family Cymothoidae. The type species is P. dvorakorum.

Petersbuchia[64]

Gen. et sp. nov

Valid

Schweigert

Late Jurassic (Kimmeridgian)

Treuchtlingen Formation

Germany

A crab belonging to the group Homolodromioidea and the family Prosopidae. The type species is P. thauckei. Announced in 2021 in an online-only journal; validated in 2023.[65]

Plakolana chiapaneca[37]

Sp. nov

Valid

Bruce et al.

Early Cretaceous (Aptian)

Sierra Madre Formation

Mexico

A member of Isopoda belonging to the group Cymothoida. Originally described as a species of Plakolana, but subsequently made the type species of the separate genus Pseudoplakolana.[66]

Platuropodus[67]

Gen. et sp. nov

Valid

Pazinato et al.

Permian (Kungurian)

Irati Formation

Brazil

A member of Isopoda. The type species is P. odysseus.

Protaxius paucisaetosus[68]

Sp. nov

In press

Andrada et al.

Early Cretaceous (Hauterivian)

Agrio Formation

Argentina

A member of the family Axiidae.

Pseudonecrocarcinus eichhorni[69]

Sp. nov

In press

Nyborg et al.

Late Cretaceous (Coniacian)

United States
( Montana)

A crab belonging to the group Raninoida and the family Paranecrocarcinidae.

Stenodactylina shotoverigiganti[70]

Sp. nov

Valid

Devillez & Charbonnier

Late Jurassic (Oxfordian)

United Kingdom

A member of Erymoidea.

Tyrannosculda[71]

Gen. et sp. nov

Valid

Haug & Haug

Late Jurassic (Tithonian)

Altmühltal Group (Eichstätt Subformation)

Germany

A mantis shrimp. The type species is T. laurae.

Vicetitrapezia[38]

Gen. et sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

Italy

A crab belonging to the superfamily Trapezioidea; the type genus of the new family Vicetitrapeziidae. The type species is V. exaltissimo.

Willinachoides[60]

Gen. et sp. nov

Valid

Lima, Aguilera & Tavares

Early-middle Miocene

Brazil

A spider crab belonging to the family Inachoididae. Genus includes new species W. santanai.

Xanthodius falconensis[27]

Sp. nov

Valid

Schweitzer, Hyžný & Feldmann

Venezuela

A species of Xanthodius.

Xiaopenaeus[72]

Gen. et sp. nov

Valid

Xing et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Penaeidae. Genus includes new species X. electrinus.

Ostracods

Name Novelty Status Authors Age Type locality Country Notes Images

Aracajuia separatta[73]

Sp. nov

In press

Vázquez García et al.

Cretaceous (Albian–Cenomanian)

Riachuelo Formation

Brazil

Aurikirkbya guadalupensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of Palaeocopida belonging to the family Kirkbyidae.

Aurila bassiounii[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Aurila burolleti[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Aurila castanyi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Aurila dipattii[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Aurila doruki[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Aurila nocellae[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Aurila violai[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Aurila zargounii[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Bairdia elcapitanensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of the family Bairdiidae.

Baschkirina tesakovae[77]

Sp. nov

Valid

Melnikova et al.

Silurian

Syadai Formation

Russia

Bythoceratina antetumida[78]

Nom. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Callistocythere sissinghi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Candona dawenkouensis[79]

Sp. nov

In press

Wang et al.

Middle Eocene to Oligocene

Dawenkou Formation

China

A species of Candona.

Candona phaseolus[80]

Sp. nov

In press

Kshetrimayum et al.

Late Cretaceous (Maastrichtian)

India

A species of Candona.

Capsacythere narjessae[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Carinocythereis carboneli[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cavellina septima[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Australia

Ceratobairdia mescaleroella[74]

Sp. nov

Valid

Forel in Tarnac et al.

Permian (Capitanian)

Bell Canyon Formation

United States
( Texas)

A member of the family Bairdiidae.

Ceratobairdia pratti[74]

Sp. nov

Valid

Forel in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of the family Bairdiidae.

Ceratobairdia sexagintaduella[74]

Sp. nov

Valid

Forel in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of the family Bairdiidae.

Chrysocythere nevianii[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cimbaurila mariellae[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cistacythereis merzeraudi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cistacythereis purii[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cypridea khandae[82]

Sp. nov

In press

Choi et al.

Early Cretaceous (Albian)

Jinju Formation

South Korea

Cytherelloidea alexanderi[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Cytherelloidea ardita[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Cytherelloidea slipperi[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Cytherelloidea temaniae[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Cytheridea jonesi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Cytheropterina crassicostata[83]

Sp. nov

Valid

Wannenmacher et al.

Middle Jurassic (Aalenian)

Achdorf Formation

Germany

A member of the family Cytheruridae.

Cytheropteron laranjeirensis[73]

Sp. nov

In press

Vázquez García et al.

Cretaceous (Albian–Cenomanian)

Riachuelo Formation

Brazil

Denticupachydomella bellcanyonensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Capitanian)

Bell Canyon Formation

United States
( Texas)

A member of Podocopida belonging to the family Microcheilinellidae.

Echinocythereis clavata[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Echinocythereis multituberculata[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Eucytherura eberti[83]

Sp. nov

Valid

Wannenmacher et al.

Middle Jurassic (Aalenian)

Achdorf Formation

Germany

A member of the family Cytheruridae.

Eucytherura fossapunctata[84]

Sp. nov

In press

Maia et al.

Late Pleistocene

Brazil

Eucytherura foveolata[83]

Sp. nov

Valid

Wannenmacher et al.

Middle Jurassic (Aalenian)

Opalinuston Formation

Germany

A member of the family Cytheruridae.

Eucytherura regina[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Eurybolbina uralica[77]

Sp. nov

Valid

Melnikova et al.

Silurian

Syadai Formation

Russia

Evlanella? conversa[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Geisina culbersonensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of Palaeocopida belonging to the family Geisinidae.

Glyptolichvinella acrista[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Australia

Gomphocythere testudo[80]

Sp. nov

In press

Kshetrimayum et al.

Late Cretaceous (Maastrichtian)

India

Healdia cutoffella[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian)

Cutoff Formation

United States
( Texas)

A member of Podocopa belonging to the family Healdiidae.

Healdia mckittrickensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian to Capitanian)

Bell Canyon Formation

United States
( Texas)

A member of Podocopa belonging to the family Healdiidae.

Healdia veeversi[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Healdiacypris drucei[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Healdianella ogmoconchelloides[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Hollinella locula[85]

Sp. nov

In press

Forel in Forel, Poulet-Crovisier & Korat

Late Permian and Early Triassic

Daye Formation
Feixianguan Formation
Kayitou Formation
Longtan Formation
Xuanwei Formation

China

A member of Palaeocopida.

Hollinella williamsranchensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian to Capitanian)

Bell Canyon Formation

United States
( Texas)

A member of Palaeocopida belonging to the family Hollinellidae.

Idiocythere caburnensis[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Isocythereis postelongata[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Jenningsina guilinensis[86]

Sp. nov

In press

Song et al.

Late Devonian

China

Karsteneis oculocosta[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Mauritsina? paradordoniensis[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Mediocytherideis hornei[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Microxestoleberis riachuelensis[73]

Sp. nov

In press

Vázquez García et al.

Cretaceous (Albian–Cenomanian)

Riachuelo Formation

Brazil

Monoceratina minangulata[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Neomonoceratina lajmii[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Neomonoceratina morkhoweni[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Neomonoceratina mostafawii[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Neomonoceratina reussi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Okadaleberis azouzi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Pachycaudites yassinii[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Paracavellina olempskae[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Parahemingwayela fauthi[73]

Sp. nov

In press

Vázquez García et al.

Cretaceous (Albian–Cenomanian)

Riachuelo Formation

Brazil

Paraparchites pecosensis[74]

Sp. nov

Valid

Crasquin in Tarnac et al.

Permian (Roadian to Capitanian)

Cutoff Formation

United States
( Texas)

A member of Podocopa belonging to the family Paraparchitidae.

Patellacythere weaveri[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Peteraurila carbonneli[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Peteraurila moisettei[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Peteraurila nachitei[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Peteraurila ulicznyi[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Pleurocythere khapissovi[87]

Sp. nov

Valid

Glinskikh & Tesakova

Middle Jurassic (Callovian)

Russia

Plumhoffia curvicosta[88]

Sp. nov

Valid

Ohmert & Franz

Middle Jurassic

Ostreenkalk Formation

Germany

A member of the family Cytheruridae.

Polycope lazarusi[89]

Sp. nov

Valid

Forel in Forel & Grădinaru

Triassic

Romania

A member of the family Polycopidae.

Polycope legio[89]

Sp. nov

Valid

Forel in Forel & Grădinaru

Triassic

Romania

A member of the family Polycopidae.

Polycope ludwigi[89]

Sp. nov

Valid

Forel in Forel & Grădinaru

Triassic

Romania

A member of the family Polycopidae.

Pontopolycope jeanlouisi[89]

Sp. nov

Valid

Forel in Forel & Grădinaru

Triassic

Romania

A member of the family Polycopidae.

Pontopolycope papai[89]

Sp. nov

Valid

Forel in Forel & Grădinaru

Triassic

Romania

A member of the family Polycopidae.

?Praeschuleridea plana[88]

Sp. nov

Valid

Ohmert & Franz

Middle Jurassic

Ostreenkalk Formation

Germany

A member of the family Schulerideidae.

Pseudocytherura carolinae[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Pseudomutilus[90]

Gen. et sp. nov

Valid

Sciuto in Sciuto & Benkhedda

Miocene (Tortonian)

Algeria

A member of the family Trachyleberididae. The type species is P. benkheddai.

Pterygocythereis carolinae[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Quasiagrenocythere[90]

Gen. et sp. nov

Valid

Sciuto in Sciuto & Benkhedda

Miocene (Tortonian)

Algeria

A member of the family Trachyleberididae. The type species is Q. tevestaensis.

Quasihermanites? punctata[73]

Sp. nov

In press

Vázquez García et al.

Cretaceous (Albian–Cenomanian)

Riachuelo Formation

Brazil

Rehacythereis stellatus[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Reversocypris australis[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Bonaparte Basin

Australia

Schuleridea langdonensis[78]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

United Kingdom

Syadaikinia[77]

Gen. et sp. nov

Valid

Melnikova et al.

Silurian

Syadai Formation

Russia

The type species is S. tumidosa.

Tenedocythere uffenordei[75]

Sp. nov

Valid

Sciuto & Reitano

Early Pliocene

Italy

Tschigovana thomasi[81]

Sp. nov

Valid

Jones, Kelman & Laurie

Carboniferous (Mississippian)

Australia

Urocythere (Pokornyella) bremani[76]

Sp. nov

Valid

Sciuto, Temani & Ammar

Miocene (Messinian)

Tunisia

Other crustaceans

Name Novelty Status Authors Age Type locality Country Notes Images

Carapacestheria? lulworthensis[91]

Sp. nov

In press

Li, Teng & Wimbledon

Late Jurassic/Early Cretaceous

Purbeck Group

United Kingdom

A clam shrimp.

Chelonibia zanzibarensis[92]

Sp. nov

In press

Collareta & Newman in Collareta et al.

Miocene

Chake-Chake beds

Tanzania

A barnacle, a species of Chelonibia.

Concinnalepas bessinensis[93]

Sp. nov

In press

Gale

Middle Jurassic (Bathonian)

France

A barnacle.

Concinnalepas rugosa[93]

Sp. nov

In press

Gale

Late Jurassic (Tithonian)

Kimmeridge Clay

United Kingdom

A barnacle.

Cornia cheni[94]

Sp. nov

Valid

Liao & Shen

Early Devonian

China

A clam shrimp.

Gibbosaverruca milazzoensis[95]

Sp. nov

Valid

Gale

Plio-Pleistocene

Italy

A barnacle.

Malayacyclus[96]

Gen. et sp. nov

Valid

Tang, Mychko, Feldmann & Schweitzer in Tang et al.

Carboniferous (Viséan)

Malaysia

A member of Cyclida. Genus includes new species M. terengganuensis. Announced in 2021; validated in 2023.[97]

Palaeolimnadiopsis zhangi[94]

Sp. nov

Valid

Liao & Shen

Early Devonian

China

A clam shrimp.

Solidobalanus digeronimoi[95]

Sp. nov

Valid

Gale

Plio-Pleistocene

Italy

A barnacle.

Research

  • A study on the anatomy and phylogenetic relationships of the species "Penaeus" natator from the Santonian of Lebanon is published by Audo, Winkler & Charbonnier (2021), who interpret this species as a relative of Pseudodrobna kenngotti from the Late Jurassic of Germany, and transfer it to the genus Pseudodrobna[98]
  • A study on the anatomy and morphological variation in Beurlenia araripensis, based on data from fossil samples from the Crato Formation (Brazil), is published by Barros et al. (2021).[99]
  • A study on the anatomy of the eyes of Callichimaera perplexa, and on their growth during the ontogeny of this crab, is published by Jenkins, Briggs & Luque (2021).[100]
  • A study on the anatomy and phylogenetic relationships of Oxyuropoda is published by Robin et al. (2021), who interpret this arthropod as the oldest known member of the crown group of Peracarida.[101]
  • Fossil evidence of early colonization of estuarine settings by ostracods is reported from the Silurian Si Ka Formation (Vietnam) by McGairy et al. (2021)[102]
  • A study on evolutionary trends in sexual dimorphism of cytheroid ostracods from the Gulf and Atlantic coastal plain from the Late Cretaceous to the late Eocene is published by Matzke-Karasz & Smith (2021).[103]

Insects

Radiodonts

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Buccaspinea[104]

Gen. et sp. nov

Valid

Pates et al.

Cambrian (Drumian)

Marjum Formation

United States
( Utah)

A hurdiid radiodont.
The type species is B. cooperi.

Houcaris[105]

Gen. et comb. nov

Valid

Wu et al.

Cambrian Series 2

Carrara Formation
Maotianshan Shales
Pioche Formation

China
United States

A tamisiocaridid radiodont.
Type species "Anomalocaris" saron (1995);
Also includes "Anomalocaris" magnabasis (2019).

Houcaris saron

Lenisicaris[106]

Gen. et sp. et comb. nov

In press

Wu et al.

Cambrian

China
United States

An anomalocaridid radiodont.
Type species L. lupata,
L. pennsylvanica moved from "Anomalocaris" pennsylvanica (1929).

Lenisicaris lupata

Paranomalocaris simplex[107]

Sp. nov

Valid

Jiao et al.

Cambrian Stage 4

Wulongqing Formation

China

An anomalocaridid radiodont.

Titanokorys[108]

Gen. et sp. nov

Valid

Caron & Moysiuk

Cambrian (Wuliuan)

Burgess Shale

Canada
( British Columbia)

A hurdiid radiodont.
The type species is T. gainesi.

Research

  • A study on the morphologies of frontal appendages and probable modes of feeding of radiodonts from the Burgess Shale is published by De Vivo, Lautenschlager & Vinther (2021).[109]

Trilobites

New taxa

Name Novelty Status Authors Age Type locality Location Notes Images

Acuticryphops prorotundus[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France

A member of the family Phacopidae.

Acutiphacops[110]

Gen. et 2 sp. et comb. nov

In press

Feist & Klapper

Late Devonian

France
Morocco

A member of the family Phacopidae. The type species is A. medius; genus also includes new species A. tchrafinensis, as well as "Phacops" erfoudensis Richter & Richter (1943).

Ampulliglabella arakii[111]

Sp. nov

Valid

Flick & Shiino

Permian (probably Wordian)

Hoso-o Formation

Japan

Anadesma[112]

Gen. et comb. nov

Valid

Holloway

Silurian (late Aeronian to early Telychian)

Canada

A member of Lichida belonging to the family Lichidae; a new genus for "Amphilichas" shallopensis Twenhofel (1928).

Arctinurus aorter[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of Lichida belonging to the family Lichidae.

Asioptychaspis lata[113]

Sp. nov

In press

Wernette et al.

Cambrian (Furongian)

Myet-Ye Formation

Myanmar

Belenopyge bellerophon[114]

Sp. nov

Valid

Flick

Devonian

Greifenstein Limestone

Germany

Bollandia reviviscens[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Bollandiinae.

Brachymetopus (Acutimetopus) laticeps[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Brachymetopidae belonging to the subfamily Brachymetopinae.

Celadiella[115]

Gen. et sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae. Genus includes new species C. celadensis.

Cheiropyge radula[111]

Sp. nov

Valid

Flick & Shiino

Permian (probably Wordian)

Hoso-o Formation

Japan

Chlupacops angularis[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France

A member of the family Phacopidae.

Chlupacops clapassousensis[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France

A member of the family Phacopidae.

Chlupacops migrans[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

Serre Formation

France

A member of the family Phacopidae.

Chlupacops narbonnensis[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France

A member of the family Phacopidae.

Chlupacops rectannulatus[110]

Sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

Serre Formation

France

A member of the family Phacopidae.

Chotecops morrisoni[116]

Sp. nov

Valid

Flick

Devonian

Günterod Limestone

Germany

Cinerana[115]

Gen. et sp. nov

Valid

Gandl

Carboniferous

Spain

Possibly a member of the family Phillipsiidae. Genus includes new species C. matallanensis.

Cronierella[110]

Gen. et 3 sp. et comb. nov

In press

Feist & Klapper

Late Devonian

Serre Formation

France
Poland

A member of the family Phacopidae. The type species is C. gallica; genus also includes new species C. postera and C. expansa, as well as "Phacops" zofiae Chlupáč (1993).

Ctenopyge magna[117]

Sp. nov

Valid

Nielsen & Andersen

Cambrian (Furongian)

Alum Shale Formation

Denmark

A member of the family Olenidae.

Cydonocephalus tiffanyae[118]

Sp. nov

Valid

Adrain & Pérez-Peris

Ordovician (Darriwilian)

Table Cove Formation

Canada
( Newfoundland and Labrador)

A member of Acanthoparyphinae.

Diacanthaspis tricosa[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of the family Odontopleuridae.

Dicranopeltis eretes[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of Lichida belonging to the family Lichidae.

Dicranopeltis kallaion[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of Lichida belonging to the family Lichidae.

Ditomopyge benkei[111]

Sp. nov

Valid

Flick & Shiino

Permian (probably Wordian)

Hoso-o Formation

Japan

Ditomopyge (Carniphillipsia) cantamudensis[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae.

Ditomopyge (Carniphillipsia) castelleriae[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae.

Dohmiella pachyacanthophilia[119]

Sp. nov

Valid

Van Viersen

Devonian (Emsian)

Spain

Dohmiella pooka[119]

Sp. nov

Valid

Van Viersen

Devonian (Eifelian)

Belgium

Eccaparadoxides epimetheus[120]

Sp. nov

In press

Geyer, Landing & Żylińska

Cambrian (Miaolingian)

Jbel Wawrmast Formation

Morocco

A member of the family Paradoxididae.

Eccaparadoxides? hestia[120]

Sp. nov

In press

Geyer, Landing & Żylińska

Cambrian (Miaolingian)

Jbel Wawrmast Formation

Morocco

A member of the family Paradoxididae.

Eccaparadoxides zelus[120]

Sp. nov

In press

Geyer, Landing & Żylińska

Cambrian (Miaolingian)

Delitzsch Formation

Germany

A member of the family Paradoxididae.

Ehmaniella tupeqarfik[121]

Sp. nov

Valid

Peel

Cambrian (Wuliuan)

Telt Bugt Formation

Greenland

A member of the family Alokistocaridae.

Eisarkaspis jonesi[122]

Sp. nov

Valid

Smith & Laurie

Ordovician (Darriwilian)

Australia

Eldoradia caerulioris[123]

Sp. nov

Valid

Peel

Cambrian (Guzhangian)

Blue Cliffs Formation

Greenland

A member of the family Bolaspididae.

Fieldaspis? iubilaei[121]

Sp. nov

Valid

Peel

Cambrian (Wuliuan)

Telt Bugt Formation

Greenland

A member of the family Zacanthoididae.

Gerastos cornix[119]

Sp. nov

Valid

Van Viersen

Devonian (Eifelian)

Belgium

Ghanaspis[122]

Gen. et sp. nov

Valid

Smith & Laurie

Ordovician (Darriwilian)

Australia

Genus includes new species G. ritchiei.

Girardina[110]

Gen. et 2 sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France
Germany

A member of the family Phacopidae. The type species is G. konradbartzschi; genus also includes new species G. consimilis, and possibly also "Phacops" liopyga Richter (1863).

Gonioteloides moffitti[124]

Sp. nov

Valid

Adrain & Karim

Ordovician (Tremadocian)

United States

Possibly a member of the family Dimeropygidae.

Gonioteloides pankowskii[124]

Sp. nov

Valid

Adrain & Karim

Ordovician (Tremadocian)

United States

Possibly a member of the family Dimeropygidae.

Griffithides inexpectatus[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Griffithidinae.

Harebayaspis[118]

Gen. et sp. nov

Valid

Adrain & Pérez-Peris

Ordovician (Darriwilian)

Table Cove Formation

Canada
( Newfoundland and Labrador)

A member of Pilekiinae. The type species is H. plurima.

Illaenus taoyuanensis[125]

Sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Iridis[122]

Gen. et sp. nov

Valid

Smith & Laurie

Ordovician (Darriwilian)

Australia

Genus includes new species I. schoonorum.

Kaotaia xuanensis[126]

Sp. nov

In press

Hughes et al.

Cambrian (Miaolingian)

Vietnam

Kawina stougei[118]

Sp. nov

Valid

Adrain & Pérez-Peris

Ordovician (Darriwilian)

Table Cove Formation

Canada
( Newfoundland and Labrador)

A member of Acanthoparyphinae.

Koneprusia dongesi[127]

Sp. nov

Valid

Alberti & Müller

Devonian (Emsian)

Rupbach Shales

Germany

A member of the family Odontopleuridae.

Koneprusites amicalis[128]

Sp. nov

Valid

Flick

Devonian

Günterod Limestone

Germany

Koneprusites cervisiaemontis[128]

Sp. nov

Valid

Flick

Devonian

Günterod Limestone

Germany

Leonaspis glaux[129]

Sp. nov

Valid

Flick & Flick

Devonian (Emsian)

Ballersbach Limestone

Germany

A member of the family Odontopleuridae.

Lycophron titan[122]

Sp. nov

Valid

Smith & Laurie

Ordovician (Darriwilian)

Australia

Mainbrookia[118]

Gen. et sp. nov

Valid

Adrain & Pérez-Peris

Ordovician (Darriwilian)

Table Cove Formation

Canada
( Newfoundland and Labrador)

A member of Deiphoninae. The type species is M. becki.

Mioptychopyge chengkouensis[130]

Sp. nov

Valid

Wei et al.

Ordovician

Houping Formation

China

Miraspis capnica[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of the family Odontopleuridae.

Morocops richterorum[116]

Sp. nov

Valid

Flick

Devonian

Greifenstein Limestone

Germany

A member of the family Phacopidae.

Namuropyge alfredi[131]

Sp. nov

Valid

Müller, Hahn & Paul

Carboniferous (Viséan)

Germany

A member of the family Brachymetopidae.

Namuropyge brooksi[131]

Sp. nov

Valid

Müller, Hahn & Paul

Carboniferous (Mississippian)

Cuyahoga Formation

United States
( Ohio)

A member of the family Brachymetopidae.

Namuropyge ulrichi[131]

Sp. nov

Valid

Müller, Hahn & Paul

Carboniferous (Viséan)

Germany

A member of the family Brachymetopidae.

Neogriffithides extremorientalis[111]

Sp. nov

Valid

Flick & Shiino

Permian (probably Wordian)

Hoso-o Formation

Japan

Newfoundlandops[118]

Gen. et sp. nov

Valid

Adrain & Pérez-Peris

Ordovician (Darriwilian)

Table Cove Formation

Canada
( Newfoundland and Labrador)

Possibly a member of Sphaerexochinae. The type species is N. karimae.

Nileus yichongqiaoensis[125]

Sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Norasaphus (Norasaphus) patersoni[122]

Sp. nov

Valid

Smith & Laurie

Ordovician (Darriwilian)

Australia

Nyterops fugax[132] Sp. nov Valid Van Viersen & Koppka Devonian (Givetian) Hanonet Formation Belgium A member of the family Phacopidae.
Nyterops gerolsteinensis[132] Sp. nov Valid Van Viersen & Koppka Devonian (Eifelian?) Germany A member of the family Phacopidae.

Occitanella[110]

Gen. et 2 sp. nov

In press

Feist & Klapper

Devonian (Frasnian)

France
Australia?

A member of the family Phacopidae. The type species is O. hexagonalis; genus also includes new species O. postinflexa, and possibly also "Trimerocephaloides" linguiformis Feist et al. (2009).

Octobronteus? incertus[133]

Sp. nov

Valid

Holloway

Silurian

United States

A member of the family Scutelluidae.

Octobronteus? ormistoni[133]

Sp. nov

Valid

Holloway

Silurian

United States

A member of the family Scutelluidae.

Octobronteus? petasus[133]

Sp. nov

Valid

Holloway

Silurian

United States

A member of the family Scutelluidae.

Panderia striolatus[125]

Sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Paratiresias peculiaris[125]

Sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Pedinopariops apsconditus[132] Sp. nov Valid Van Viersen & Koppka Devonian (Eifelian) Jemelle Formation Belgium A member of the family Phacopidae.

Planiscutellum apollo[133]

Sp. nov

Valid

Holloway

Silurian (Aeronian)

Canada

A member of the family Scutelluidae.

Pseudohelmutia[134]

Gen. et sp. nov

Valid

Basse & Müller

Early Devonian

Germany

A member of the subfamily Proetinae. Genus includes new species P. feisti.

Rupbachella[135]

Gen. et sp. nov

Disputed

Alberti

Devonian (Emsian)

Rupbach Shale

Germany

A member of the family Odontopleuridae. The type species is R. paedomorpha. Van Viersen (2024) considered R. paedomorpha to be a junior synonym of Leonaspis kleini.[136]

Sinagnostus[125]

Gen. et sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Genus includes new species S. mirabilis.

Solenoparia changi[137]

Sp. nov

Valid

Sun, Bentley & Laurie

Cambrian (Guzhangian)

Warburton Basin

Australia

Timsaloproetus alissae[138]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Emsian)

Morocco

A member of the family Proetidae.

Trochurus searcyensis[112]

Sp. nov

Valid

Holloway

Silurian

United States

A member of Lichida belonging to the family Lichidae.

Unguliproetus chamaeleo[128]

Sp. nov

Valid

Flick

Devonian

Ballersbach Limestone

Germany

Unguliproetus fredknoppi[134]

Sp. nov

Valid

Basse & Müller

Early Devonian

Germany

A member of the subfamily Proetinae.

Valberzosa[115]

Gen. et sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Bollandiinae. Genus includes new species V. rara.

Waideggula (Camasobrina) verdenensis[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae.

Waideggula (Waideggula) cantabrica tabardina[115]

Ssp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae.

Waideggula (Waideggula) pedregalensis[115]

Sp. nov

Valid

Gandl

Carboniferous

Spain

A member of the family Phillipsiidae belonging to the subfamily Ditomopyginae.

Weania insulana[111]

Sp. nov

Valid

Flick & Shiino

Permian (probably Wordian)

Hoso-o Formation

Japan

Yanpingia[125]

Gen. et sp. nov

Valid

Wei et al.

Ordovician (Darriwilian)

China

Genus includes new species Y. punctata.

Research

  • A study on middle–late Cambrian trilobite diversity patterns in South China is published by Zhang et al. (2021).[139]
  • Sun, Zeng & Zhao (2021) describe digestive structures of representatives of five trilobite genera from the Cambrian Mantou Formation and Zhangxia Formation (Liaoning, China).[140]
  • Hou, Hughes & Hopkins (2021) report structural details of the upper limb branch of Triarthrus eatoni and Olenoides serratus, and interpret their findings as indicating that the upper limb branch of trilobites served a respiratory function.[141]
  • A study on the morphology of Redlichia rex and Olenoides serratus, aiming to determine whether these trilobites were adapted for durophagy, is published by Bicknell et al. (2021).[142]
  • A study exploring the existence and the nature of growth gradients along the main body axis of Oryctocarella duyunensis is published by Dai et al (2021), who interpret O. duyunensis as the first trilobite with documented determinate growth.[143]
  • Description of all meraspid stages of Oryctocarella duyunensis, based on data from specimens from the Cambrian Balang Formation (Hunan, South China), is published by Dai et al. (2021).[144]
  • A study on the ontogenic moulting sequence of Arthricocephalites xinzhaiheensis, based on data from specimens from the Balang Formation, is published by Wang et al. (2021).[145]
  • A study on post-embryonic axial growth in Estaingia bilobata is published by Holmes, Paterson & García-Bellido (2021).[146]
  • A study on the functional performance of Placoparia cambriensis in water is published by Esteve et al. (2021), who interpret their findings as indicating that this trilobite was not able to swim, but it was capable of hopping locomotion, and it might represent a transitional form between benthic animals without swimming skills and animals able to swim near the benthos.[147]
  • A study on the phylogenetic relationships within Harpetida, and on the impact of the Ordovician–Silurian extinction events on this group, is published by Beech & Lamsdell (2021).[148]
  • The first known Silurian trilobite specimen preserved with soft parts reported to date (a specimen of Dalmanites preserved with appendages and alimentary system) is described from the Herefordshire Lagerstätte (United Kingdom) by Siveter et al. (2021).[149]
  • A study on the long-term evolutionary history of Devonian trilobites in North Africa is published by Bault et al. (2021).[150]
  • A study on the hydrodynamics of queuing behaviour in Trimerocephalus chopini is published by Song et al. (2021).[151]
  • Evidence from trace and body fossils indicative of the presence of trilobites in brackish-water settings is presented by Mángano et al. (2021).[152]
  • A study on the chemical changes in the exoskeleton of trilobites induced by diagenesis, based on data from pygidia of Athabaskia anax from the Miaolingian of San Isidro (Argentina), is published by D'Angelo et al. (2021), is published by D'Angelo et al. (2021), who argue that some morphological characteristics of the trilobite pygidia are in fact results of chemical and structural changes taking place during fossilization, and evaluate possible systematic implications of the chemical data, advising caution when using morphological characteristics of the exoskeletons to establish new taxa.[153]
  • The study on the internal structures of eyes of trilobites belonging to the genera Asaphus and Archegonus published by Scholtz, Staude & Dunlop (2019)[154] is criticized by Schoenemann & Clarkson (2021).[155][156]
  • A study on the biomechanics of the trilobite cephalon is published by Esteve et al. (2021), who interpret their findings as indicating that in the sutured trilobites the cephalon was able to withstand greater stresses than in their non-sutured counterparts, and argue that the ability to withstand greater burrowing loads enabled trilobites to successfully invade bioturbated and more consolidated sediments during the Cambrian substrate revolution.[157]
  • A study on the compound eyes of trilobites belonging to the group Phacopina is published by Schoenemann et al. (2021), who interpret their findings as indicating that these trilobites had hyper-compound eyes hiding an individual compound eye below each of the big lenses, resulting in each of the compound eyes comprising tens or hundreds of small compound eye systems.[158]
  • A study on the systematics of Devonian trochurine trilobites is published by Van Viersen (2021).[159]

Other arthropods

General research

References

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