2021 in paleoichthyology
From Wikipedia, the free encyclopedia
| List of years in paleoichthyology |
|---|
This list of fossil fish research presented in 2021 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2021.
Jawless fishes research
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
In press |
Meng & Gai |
A member of Galeaspida belonging to the family Tridensaspidae. Genus includes new species F. liui. |
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|
Gen. et sp. nov |
Valid |
Liu et al. |
A member of Galeaspida belonging to the family Hanyangaspidae. Genus includes new species H. inexpectatus. |
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|
Gen. et sp. nov |
Valid |
Liu et al. |
A member of Galeaspida belonging to the group Eugaleaspidiformes. The type species is J. retrospina. |
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|
Gen. et sp. nov |
Valid |
Jiang et al. |
Early Devonian |
A member of Galeaspida. Genus includes new species Q. elaia. |
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- A study on the phylogenetic relationships of cyathaspidids is published by Elliott, Lassiter & Blieck (2021).[5]
- Miyashita et al. (2021) report larval and juvenile forms of four stem lampreys from the Paleozoic era (Hardistiella, Mayomyzon, Pipiscius and Priscomyzon), including a hatchling-to-adult growth series of Priscomyzon, and report that the studied larvae display features that are otherwise unique to adult modern lampreys, and lack the defining traits of ammocoetes.[6]
- A study on the anatomy and likely feeding ecology of Mesomyzon mengae, based on data from new, well-preserved specimens, is published by Wu, Chang & Janvier (2021).[7]
- A study on the histology of the dermal skeleton in Procephalaspis oeselensis, Aestiaspis viitaensis, Dartmuthia gemmifera and four species of Tremataspis is published by Bremer et al. (2021), who interpret their findings as indicative of the emergence of the complex pore-canal system in Tremataspis through the modification of the structures already present in other taxa.[8]
- A study aiming to determine whether the earliest vertebrates may have swum under various conditions without a clearly-differentiated tail fin, based on data from an abstracted model of Metaspriggina walcotti, is published by Rival, Yang & Caron (2021).[9]
- A study on the morphological and functional diversity of osteostracan and galeaspid headshields, and on its implications for the knowledge of the ecology of the immediate jawless relatives of jawed vertebrates, is published by Ferrón et al. (2021).[10]
- Redescription of Nochelaspis maeandrine is published by Meng, Zhu & Gai (2021).[11]
- A study on the anatomy of a dorsal head shield of Kalanaspis delectabilis is published by Tinn et al. (2021).[12]
Placoderms
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Valid |
Li, Zhu & Zhu in Li et al. |
A placoderm closely related to the last common ancestor of bony and cartilaginous fishes. The type species is B. micros. |
 | ||||
|
Gen. et sp. nov |
Jobbins et al. |
Devonian (Givetian) |
A member of Arthrodira belonging to the family Plourdosteidae. The type species is L. ziregensis. |
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Placoderm research
- Zhu et al. (2021) use CT scanning to reveal the endocast of Brindabellaspis stensioi, and evaluate the implications of its anatomy for the knowledge of the phylogenetic relationships of early jawed vertebrates.[15]
- Redescription of the anatomy of the headshield of Parayunnanolepis xitunensis is published by Wang & Zhu (2021).[16]
- Description of new fossil material of Palaeacanthaspis vasta from the Devonian (Lochkovian) Chortkiv Formation (Ukraine), and a study on the phylogenetic relationships of this species, is published by Dupret et al. (2021).[17]
Acanthodians
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et comb. nov |
Valid |
Dearden et al. |
A new genus for "Vernicomacanthus" waynensis Miles (1973) |
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|
Sp. nov |
Valid |
Li et al. |
||||||
|
Nostolepis qujingensis[19] |
Sp. nov |
Valid |
Li et al. |
Devonian (Lochkovian) |
Xitun Formation |
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Acanthodian research
- A study on the development of teeth in acanthodians, and on its implications for the knowledge of the evolution of teeth of jawed vertebrates, is published by Rücklin et al. (2021).[20]
- A study on the anatomy of teeth, jaws and associated oral structures of acanthodians, and on their implications for the knowledge of the evolution of dentition of modern cartilaginous fishes, is published by Dearden & Giles (2021).[21]
Cartilaginous fishes
| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Vullo et al. |
Late Cretaceous (Turonian) |
A probable planktivorous shark placed in the new family Aquilolamnidae, of uncertain placement. Possibly a member of Lamniformes. The type species is A. milarcae. |
 | ||||
|
Sp. nov |
Valid |
Canevet & Lebrun |
Miocene |
A species of Carcharhinus. |
||||
|
Sp. nov |
Valid |
Canevet & Lebrun |
Miocene |
A species of Carcharhinus. |
||||
|
Sp. nov |
Valid |
Cicimurri & Ebersole |
A species of Carcharhinus. |
|||||
|
Sp. nov |
Valid |
Feichtinger et al. |
||||||
|
Gen. et sp. nov |
Valid |
Hodnett et al. |
A medium-sized ctenacanthiform shark known from a complete skeleton with soft tissue. The type species is D. hoffmanorum. |
 | ||||
|
Gen. et sp. nov |
Valid |
Stumpf et al. |
Late Jurassic (Tithonian) |
A member of the family Hybodontidae. The type species is D. maiseyi. |
||||
|
Gen. et sp. nov |
Valid |
Li et al. |
A member of Euselachii. Genus includes new species F. orientalis. |
|||||
|
Gen. et sp. nov |
Valid |
Roelofs et al. |
Genus includes new species J. ambiguus. |
|||||
|
Gen. et sp. nov |
Valid |
Li et al. |
Triassic (Ladinian–Carnian) |
A member of Euselachii. Genus includes new species K. nimaiguensis. |
||||
|
Gen. et comb. nov |
Valid |
Ivanov, Duffin & Richter |
A member of the family Jalodontidae. Genus includes "Phoebodus" brodiei Woodward (1893) (interpreted by Ivanov, Duffin & Richter, 2021 as a senior synonym of "Phoebodus" keuperinus Seilacher, 1948). |
|||||
|
Gen. et comb. nov |
Valid |
Greenfield |
late Eocene or Cretaceous (uncertain) |
A possible member of the family Pseudoscapanorhynchidae. The type species is "Archaeolamna" apophysata Li (1997). |
 | |||
|
Gen. et sp. nov |
Valid |
Bronson |
Carboniferous (Mississippian) |
A symmoriiform. The type species is M. ozarkanum. |
||||
|
Gen. et comb. nov |
Valid |
Long et al. |
Devonian (Emsian) |
Cravens Peak Beds |
A member of the family Mcmurdodontidae; a new genus for "Mcmurdodus" whitei Turner & Young (1987). |
|||
|
Gen. et sp. nov |
Valid |
Popov & Shapovalov |
Late Jurassic |
A chimaera belonging to the family Callorhinchidae. Genus includes new species M. robustus. |
||||
|
Gen. et sp. nov |
Valid |
Collareta et al. |
A member of Rajiformes, possibly a skate. The type species is N. wardi. |
|||||
|
Sp. nov |
Valid |
Canevet & Lebrun |
Miocene |
A species of Negaprion. |
||||
|
Sp. nov |
Valid |
Ivanov |
||||||
|
Gen. et sp. nov |
Valid |
Lebedev & Popov in Lebedev et al. |
Dashkovo Formation |
A chimaera. Genus includes new species P. mirabilis. |
||||
|
Sp. nov |
In press |
Jambura, Stumpf & Kriwet |
Late Cretaceous (Cenomanian) |
|||||
|
Sp. nov |
In press |
Amadori et al. |
||||||
|
Gen. et sp. nov |
Valid |
Ivanov in Ivanov et al. |
A member of the family Anachronistidae. Genus includes new species R. lata. |
|||||
|
Gen. et sp. nov |
Valid |
Li et al. |
Triassic (Ladinian–Carnian) |
A member of Elasmobranchii of uncertain phylogenetic placement. Genus includes new species R. xingyiensis. |
||||
|
Sp. nov |
Valid |
Sharma & Singh |
Jaisalmer Formation |
Marine hybodont shark |
||||
|
Strophodus jaisalmerensis[43] |
Sp. nov |
In press |
Kumar et al. |
A hybodont shark. |
||||
|
Sp. nov |
Laurito & Valerio |
Miocene–Pliocene (Messinian–Piacenzian) |
A stingray, a species of Taeniurops. |
|||||
|
Sp. nov |
Valid |
Luccisano et al. |
Early Permian |
Autun Basin |
||||
|
Nom. nov |
Valid |
Duffin |
Late Triassic |
A member of Neoselachii; a replacement name for Vallisia Duffin (1982). |
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Cartilaginous fish research
- Description of new fossil material of Gualepis elegans from the Lower Devonian of Yunnan (China), and a study on the phylogenetic relationships of this fish, is published by Cui et al. (2021).[47]
- Mottequin et al. (2021) reject the interpretation of Spiraxis interstrialis as chondrichthyan egg cases, and evaluate the implications of this reinterpretation for the knowledge of the evolution of oviparity in cartilaginous fishes.[48]
- Description of the first known skull remains of Onchopristis numidus from the Cretaceous Kem Kem Group (Morocco), and a study on the anatomy and phylogenetic relationships of this species, is published by Villalobos-Segura et al. (2021), who name a new family Onchopristidae.[49]
- New, exceptionally well-preserved skeleton of Asteracanthus ornatissimus, preserved with teeth that markedly differ from other teeth referred to Asteracanthus, is described from the Tithonian Altmühltal Formation (Germany) by Stumpf et al. (2021), who interpret this specimen as indicating that Asteracanthus and Strophodus represent two valid genera distinct from all other hybodontiforms.[50]
- A study on the morphological diversity of teeth of lamniform sharks from mid-Cretaceous assemblages in Australia, and on its implications for the knowledge of the composition of mid-Cretaceous shark communities and their recovery in the aftermath of the Cenomanian-Turonian boundary event, is published by Bazzi, Kear & Siversson (2021).[51]
- A study on the biomechanics of teeth of five species of Otodus, aiming to assess the functional significance of morphological trends in otodontid teeth and to test whether the morphology of otodontid teeth enabled the transition from piscivory to predation on marine mammals and the evolution of titanic body sizes, is published by Ballell & Ferrón (2021)[52]
- A study on a bonebed in the Oligocene Chandler Bridge Formation (South Carolina, United States) with a large sample of Carcharocles angustidens dominated by small teeth is published by Miller, Gibson & Boessenecker (2021), who interpret this bonebed as a nursery area for C. angustidens.[53]
- A study on growth patterns, reproductive biology and likely lifespan of Otodus megalodon is published by Shimada et al. (2021).[54]
- Perez, Leder & Badaut (2021) present a novel method for estimating body size in fossil lamniform sharks, and attempt to determine the body size of Otodus megalodon.[55]
- Revision of the fossil record of the extant tiger shark and the extinct members of the tiger shark lineage is published by Türtscher et al. (2021).[56]
- Redescription of Striatolamia tchelkarnurensis is published by Malyshkina (2021).[57]
- Shark teeth which might represent the first occurrence of the blacknose shark in the Pacific Ocean are described from the Pliocene Upper Onzole Formation (Ecuador) by Collareta et al. (2021), who evaluate the implications of this finding for the knowledge of the evolutionary history of the blacknose shark and the whitenose shark.[58]
- Two fossil teeth of the blacktip shark are reported from lower Pliocene marine deposits of Tuscany (Italy) by Collareta et al. (2021), representing the first known occurrence of this species in the fossil record from both Europe and the Mediterranean Basin.[59]
- A study on the morphological diversity of extant and fossil shark teeth, and on their implications for the knowledge of the evolution of lamniform and carcharhiniform sharks throughout the last 83 million years, is published by Bazzi et al. (2021).[60]
- A study on the evolutionary history of sharks across the Cretaceous–Paleogene extinction event, as indicated by morphological diversity of shark teeth across the Cretaceous–Paleogene interval, is published by Bazzi et al. (2021).[61]
- Evidence of a previously unknown major extinction of sharks in the early Miocene, ~19 million years ago, is presented by Sibert & Rubin (2021);[62] the study is subsequently criticized by Naylor et al. (2021)[63][64] and Feichtinger et al. (2021).[65][66]
- A study on shark scales from mid-Holocene (~7,000-y-old) and modern reef sediments in Bocas del Toro (Panama), aiming to determine changes of shark abundance in this area since the mid-Holocene and their possible causes, is published by Dillon et al. (2021).[67]
Ray-finned fishes
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et comb. nov |
Valid |
Taverne & Capasso |
Akrabou Formation |
A member of the family Pycnodontidae. The type species is A. aldrovandii. |
||||
|
Sp. nov |
Valid |
Bakaev in Ivanov et al. |
Permian (Roadian) |
Cutoff Formation |
A member of the family Elonichthyidae. |
|||
|
Sp. nov |
Valid |
Capasso et al. |
Late Cretaceous (Maastrichtian) |
A member of Pycnodontiformes. |
||||
|
Anomoeodus caddoi[70] |
Sp. nov |
Valid |
Suarez et al. |
A member of Pycnodontiformes. |
||||
|
Sp. nov |
In press |
Díaz-Cruz, Alvarado-Ortega & Cantalice |
Late Cretaceous (Campanian) |
Angostura Formation |
A member of Aulopiformes belonging to the family Dercetidae. |
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|
Gen. et sp. nov |
Figueroa, Weinschütz & Friedman |
Middle Devonian or older |
Paraná Basin |
An early ray-finned fish. Genus includes new species A. ferox. |
||||
|
Sp. nov |
Valid |
Nam, Nazarkin & Bannikov |
Middle Miocene |
A species of Auxis. |
||||
|
Gen. et sp. nov |
In press |
Hacker & Shimada |
A member of Ichthyodectiformes. Genus includes new species B. carteri. |
|||||
|
Gen. et comb. nov |
Valid |
Schwarzhans, Milàn & Carnevale |
Kerteminde Marl |
A member of the family Macrouridae. The type species is "Hymenocephalus" rosenkrantzi Schwarzhans (2003). |
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|
Gen. et comb. nov |
Valid |
Taverne & Capasso |
A member of the family Pycnodontidae. The type species is "Proscinetes" pillae Capasso (2007). |
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|
Gen. et sp. nov |
Valid |
Štamberg & Steyer |
Permian |
Brive Basin |
A pygopterid. Genus includes new species B. chantepieorum. |
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|
Sp. nov |
Schwarzhans & Nielsen |
Early Miocene |
Navidad Formation |
A species of Capromimus. |
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|
Sp. nov |
In press |
Newman et al. |
Devonian (Givetian) |
|||||
|
Gen. et sp. nov |
Valid |
Cantalice, Than-Marchese & Villalobos-Segura |
A member of Acanthopterygii of uncertain phylogenetic placement. Genus includes new species C. alvaradoi. |
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|
Sp. nov |
Valid |
Murray & Holmes |
Late Eocene |
A species of Clarotes. |
||||
|
Sp. nov |
Valid |
López-Arbarello & Ebert |
A member of Chondrostei belonging to the family Coccolepididae. |
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|
Sp. nov |
Valid |
Su, Chang & Chen |
Miocene |
Xiacaowan Formation |
A species of Ctenopharyngodon. |
|||
|
Sp. nov |
Valid |
Su, Chang & Chen |
Miocene |
Xiejia Formation |
A species of Ctenopharyngodon. |
|||
|
Gen. et sp. nov |
Valid |
Su, Chang & Chen |
Ulanbulage Formation |
A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is D. saintjaquensis. |
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|
Gen. et sp. nov |
In press |
Abu El-Kheir et al. |
Late Cretaceous (Maastrichtian) |
A member of Pycnodontiformes. Genus includes new species D. tavernensis. |
||||
|
Sp. nov |
Valid |
Ebert |
Late Jurassic (Kimmeridgian) |
A member of Ophiopsiformes. |
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|
Gen. et sp. nov |
Valid |
Su, Chang & Chen |
Oligocene |
Dongying Formation |
A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is E. liui. |
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|
Gen. et sp. nov |
In press |
Wick |
Late Cretaceous (Campanian) |
A member of Characiformes. Genus includes new species E. malateres. |
||||
|
Sp. nov |
Valid |
Ma, Xu & Geng |
A member of the family Colobodontidae. |
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|
Gen. et sp. nov |
Valid |
Taverne |
Late Cretaceous (Santonian) |
A lanternfish. The type species is G. sorbinii. |
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|
Gen. et sp. nov |
Valid |
Chen et al. |
A member of Clupeomorpha belonging to the group Ellimmichthyiformes. The type species is G. superstes. |
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|
Gen. et sp. nov |
Valid |
Reichenbacher & Bannikov |
Miocene |
A member of the family Gobiidae. The type species is K. prikryli. |
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|
Nom. nov |
Valid |
Schultze in Schultze et al. |
A replacement name for Kellia Kazantseva-Selezneva (1980). |
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|
Gen. et sp. nov |
Valid |
Přikryl |
Oligocene–early Miocene |
A deep-sea smelt. The type species is K. brzobohatyi. |
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|
Nom. nov |
Valid |
Schultze in Schultze et al. |
A replacement name for Neuburgia Kazantseva-Selezneva (1980). |
|||||
|
Nom. nov |
Valid |
Schultze in Schultze et al. |
Triassic |
A replacement name for Oshia Sytchevskaya (1999). |
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|
Gen. et sp. nov |
Valid |
Taverne |
A member of the family Pantodontidae. The type species is P. vandersypeni. |
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|
Gen. et sp. nov |
Valid |
Taverne |
Late Cretaceous (Cenomanian) |
A member of the family Pantodontidae. The type species is P. haqelensis. |
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|
Sp. nov |
Valid |
Xu |
Middle Triassic (Anisian) |
Guanling Formation |
A member of Neopterygii belonging to the group Louwoichthyiformes. |
|||
|
Gen. et sp. nov |
Valid |
Taverne |
Late Cretaceous (Cenomanian) |
A member of the family Pantodontidae. The type species is P. libanicus. |
||||
|
Sp. nov |
Valid |
Bannikov |
Miocene |
A member of Perciformes belonging to the group Stromateoidei. |
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|
Gen. et comb. nov |
Valid |
Nazarkin & Bannikov |
Miocene |
A member of Percoidei of uncertain phylogenetic placement; a new genus for "Pentaceros" sakhalinicus Gretchina (1975). |
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|
Gen. et sp. nov |
In press |
Wick |
Late Cretaceous (Campanian) |
Aguja Formation |
A member of Characiformes. Genus includes new species P. laramidensis. |
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|
Sp. nov |
Valid |
Kevrekidis, Arratia & Reichenbacher in Kevrekidis et al. |
Late Miocene |
A member of the family Clupeidae. |
||||
|
Gen. et sp. nov |
Valid |
Reichenbacher & Bannikov |
Miocene |
A member of the family Gobiidae. The type species is P. manfredi. |
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|
Sp. nov |
Valid |
Ren & Xu |
Middle Triassic (Anisian) |
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|
Gen. et sp. nov |
Valid |
Yabumoto & Nazarkin |
Late Miocene |
Koshikawa Formation |
A member of the family Scorpaenidae. Genus includes new species R. sakurai. |
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|
Gen. et sp. et comb. nov |
Valid |
Reichenbacher & Bannikov |
Miocene |
A member of the family Gobiidae. The type species is S. compactus; genus also includes "Hesperichthys" iugosus Schwarzhans, Brzobohatý & Radwańska (2020). |
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|
Sp. nov |
Valid |
Renesto, Magnani & Stockar |
||||||
|
Gen. et comb. nov |
Valid |
Liu |
Early Eocene |
A Catostomidae sucker. |
||||
|
Gen. et 2 sp. nov |
Valid |
Reichenbacher & Bannikov |
Miocene |
A member of the family Gobiidae. The type species is Y. decoratus; genus also includes Y. naslavcensis. |
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Otolith taxa
| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov |
Schwarzhans & Nielsen |
Early Miocene |
A species of Achirus. |
|||||
|
Sp. nov |
Schwarzhans & Nielsen |
Early Miocene |
Lacui Formation |
A species of Achirus. |
||||
|
Sp. nov |
Schwarzhans & Nielsen |
Early Miocene |
Lacui Formation |
A species of Agonopsis. |
||||
| Aphanolebias bettinae[104] | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene |  Serbia |
A member of the family Valenciidae. | ||
| Archaemacruroides vanknippenbergi[105] | Sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation |  Belgium |
A member of Gadiformes of uncertain phylogenetic placement. | |
| Ariosoma mesohellenica[106] | Sp. nov | Valid | Agiadi, Koskeridou & Thivaiou | Miocene (Aquitanian) |  Greece |
A species of Ariosoma. | ||
| Chiloconger chilensis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation |  Chile |
A species of Chiloconger. | ||
| Citharichthys parvisulcus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds Lacui Formation |
Chile |
A species of Citharichthys. | ||
| Citharichthys vergens[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation |
Chile |
A species of Citharichthys. | ||
| Coelorinchus fidelis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún Beds Lacui Formation |
Chile |
A species of Coelorinchus. | ||
| Coelorinchus rapelanus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Coelorinchus. | ||
| Cottunculus primaevus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation |
Chile |
A species of Cottunculus. | ||
| Cretaserranus[105] | Gen. et sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation | Belgium |
A member of Perciformes, possibly belonging to the family Serranidae. Genus includes new species C. maastrichtensis. | |
| Gnathophis elongatus[106] | Sp. nov | Valid | Agiadi, Koskeridou & Thivaiou | Miocene (Aquitanian) | Greece |
A species of Gnathophis. | ||
| Gnathophis quinzioi[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Gnathophis. | ||
| Gobiosoma? axsmithi[107] | Sp. nov | Valid | Ebersole, Cicimurri & Stringer | Oligocene (Rupelian) | Byram Formation |  United States(  Alabama) |
A member of the family Gobiidae. | |
| Hellenigobius[108] | Gen. et sp. et comb. nov | Valid | Schwarzhans, Agiadi & Thivaiou | Miocene |  Czech Republic |
A member of the subfamily Gobionellinae. The type species is H. praeschismatus; genus also includes "Pomatoschistus" bunyatovi Bratishko et al. (2015). | ||
| Klincigobus haraldahnelti[104] | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene | Serbia |
A member of the family Gobiidae. | ||
| Kuhlia orientalis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation |
Chile |
A flagtail. | ||
| Lampanyctus ipunensis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds | Chile |
A species of Lampanyctus. | ||
| Larimichthys koae[109] | Sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation |  Taiwan |
A species of Larimichthys. | |
| Lepophidium chonorum[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation |
Chile |
A species of Lepophidium. | ||
| Lepophidium mapucheorum[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Lepophidium. | ||
| Maurolicus brevirostris[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Maurolicus. | ||
| Navidadichthys[78] | Gen. et sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
Possibly a member of the family Prototroctidae. The type species is N. mirus. | ||
| Nezumia epuge[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation |
Chile |
A species of Nezumia. | ||
| Nibea chaoi[109] | Sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation | Taiwan |
A species of Nibea. | |
| Paracarapus[78] | Gen. et sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A pearlfish. The type species is P. chilensis. | ||
| Physiculus pichi[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Physiculus. | ||
| Plesiogobius[108] | Gen. et sp. et comb. nov | Valid | Schwarzhans, Agiadi & Thivaiou | Miocene (Aquitanian) | Greece |
A member of the family Gobiidae belonging to the subfamily Gobiinae. The type species is P. felliensis. | ||
| Polyipnus bandeli[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Polyipnus. | ||
| Pseudonus humilis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds Navidad Formation |
Chile |
A species of Pseudonus. | ||
| Pterothrissus transpacificus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Pterothrissus. | ||
| Pythonichthys panulus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation | Chile |
A species of Pythonichthys. | ||
| Rhinocephalus cretaceus[105] | Sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation | Belgium |
A member of the family Merlucciidae. | |
| Rhynchoconger chiloensis[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Rhynchoconger. | ||
| Severnichthus[110][111] | Gen. et sp. nov | In press | Stringer & Schwarzhans | Late Cretaceous (Maastrichtian) | Severn Formation | United States (  Maryland) |
Possibly a member of Polymixiiformes. Genus includes new species S. bourdoni. | |
| Sirembola supersa[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation | Chile |
A cusk-eel. | ||
| Spectrunculus sparsus[78] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Chile |
A species of Spectrunculus. | ||
| Taosciaena[109] | Gen. et 2 sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation | Taiwan |
A member of the family Sciaenidae. The type species is T. jiangi; genus also includes T. hui. | |
| Toxopyge vracevicensis[104] | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene | Serbia |
A member of the family Gobiidae. |
Ray-finned fish research
- Putative paraxial ossicle of a member of the family Molidae, possibly representing the first fossil find of the genus Mola from the Mediterranean Basin reported to date, is described from the Miocene Pietra Leccese Formation (Apulia, Italy) by Collareta et al. (2021).[112]
- A platysomid specimen, representing the earliest deep-bodied actinopterygian reported to date, is described from the Carboniferous (Tournaisian) Horton Bluff Formation (Canada) by Wilson, Mansky & Anderson (2021), who evaluate the implications of this findings for the knowledge of the evolution of early ray-finned fishes.[113]
- A review of the fossil record of Early–Middle Triassic marine bony fishes, aiming to determine the implications of poor fossil record from the late Olenekian-early middle Anisian interval on the knowledge of the Triassic radiation of bony fishes, is published by Romano (2021).[114]
- A diverse assemblage of late Maastrichtian and Paleocene ray-finned fishes is described from Evrytania (Greece) by Argyriou & Davesne (2021).[115]
- A study on the diversity of fishes from upper Paleocene microfossil localities in the Ravenscrag Formation (Saskatchewan, Canada) is published by Sinha et al. (2021).[116]
- New fish fauna dating to the Paleocene–Eocene Thermal Maximum, indicating that diverse fish communities thrived in the paleotropics during this time period, is reported from Egypt by El-Sayed et al. (2021).[117]
- Heingård et al. (2021) report preservation of residues of both internal and integumentary tissues in the form of dark organic stains in fossil fish larvae from the Eocene (Ypresian) Stolleklint Clay (Ølst Formation, Denmark).[118]
- Revision of the fossil material of sturgeons from the Upper Miocene deposits of southern Ukraine is published by Hilton, Kovalchuk & Podoplelova (2021).[119]
- A study on the morphological diversity and evolution of pycnodontiforms is published by Cawley et al. (2021).[120]
- A study on fossil crushing dentitions of Pycnodus zeaformis and P. maliensis, providing evidence of a distinct pattern of gap-filling tooth addition in pycnodonts, with individual large teeth replaced by multiple small teeth, is published by Collins & Underwood (2021).[121]
- A study on the histology of teeth and bones of Neoproscinetes penalvai and bones of Tepexichthys aranguthyorum is published by Meunier et al. (2021).[122]
- A redescription of Atacamichthys greeni is published by Arratia et al. (2021), who interpret it as a stem-group teleost, and name the new family Atacamichthyidae.[123]
- Revision of members of the clade Archaeomaenidae is published by Bean (2021), who considers Madariscus robustus to be a junior synonym of Archaeomaene tenuis.[124]
- A study on genome size evolution in fossil stem-group teleosts (based on data from bone cell volumes in fossil specimens), aiming to determine the timing of whole-genome duplication in the evolutionary history of teleosts, is published by Davesne et al. (2021).[125]
- New fossil material of elopomorphs, including the earliest records of members of the genera Albula and Paralbula from Gondwana reported to date and one of the earliest records of the genus Egertonia, is described from the Upper Cretaceous Mahajanga Basin (Madagascar) by Ostrowski (2021).[126]
- A study on the evolutionary history of lanternfishes, primarily based on the fossil record of otoliths, is published by Schwarzhans & Carnevale (2021).[127]
- Armbruster & Lujan (2021) identify Taubateia paraiba as a member of Rhinelepinae.[128]
