2019 in paleomalacology

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This list 2019 in paleo malacology is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that were described during the year 2019, as well as other significant discoveries and events related to molluscan paleontology that would occur in 2019.

Description of an assemblage of conchiferan mollusks from the Cambrian Shackleton Limestone (Antarctica) is published by Claybourn et al. (2019).^[1]
A study on Early Triassic recovery of ammonites and gastropods after the Permian–Triassic extinction event is published by Pietsch et al. (2019).^[2]
A study on the impact of the Cenomanian-Turonian boundary event and other Cretaceous oceanic anoxic events on generic- and species-level molluscan diversity, extinction rates and ecological turnover is published by Freymueller, Moore & Myers (2019).^[3]
A juvenile ammonite specimen assigned to the genus Puzosia and four marine gastropods are reported from the Cretaceous Burmese amber of Myanmar by Yu et al. (2019).^[4]
A study on the timing and nature of recovery of benthic marine ecosystems of Antarctica after the Cretaceous–Paleogene extinction event, as indicated by data from fossils of benthic molluscs, is published by Whittle et al. (2019).^[5]

Ammonites

Research

Dimorphic Indonesian ammonite Macrocephalites keeuwensis is reported from the core of Jara Dome (Kutch district, India) by Jain (2019).^[6]
Description of new fossils of members of the genus Lytohoplites from the Neuquén Basin (Argentina), taxonomic revision of the Lytohoplites species occurring in this basin, and a study on the ontogeny, probable sexual dimorphism and spectrum of intraspecific variability of members of this genus, is published by Vennari & Aguirre-Urreta (2019).^[7]
A study on the intraspecific variation through the ontogeny of the Late Cretaceous ammonite Scaphites whitfieldi from the Western Interior of the United States is published by Klein & Landman (2019).^[8]
A lower jaw of Spathites puercoensis is described from the Turonian Carlile Member of the Mancos Shale (New Mexico, United States) by Landman et al. (2019).^[9]
A study on periodic variations in carbon and oxygen stable isotope profiles from Campanian and Maastrichtian ammonites belonging to the genus Baculites, and on their implications for inferring the life histories of these ammonites, is published by Ellis & Tobin (2019).^[10]
A study on the hydrostatic properties of the orthoconic morphotype of Baculites compressus and on their implications for the knowledge of the mode of life of this animal is published by Peterman et al. (2019).^[11]
A study on the ecology and growth of baculitid and scaphitid ammonites from the Maastrichtian Owl Creek Formation (Mississippi, United States), evaluating whether these taxa may have migrated, is published by Ferguson et al. (2019).^[12]
A study on a sutural anomaly of ammonite specimens known as sutural pseudoinversion is published by Rogov (2019).^[13]
A study on the systematics of Late Cretaceous ammonites from the southeastern San Juan Basin (Sandoval County, New Mexico) and on the stratigraphic framework of their fossils is published by Sealey & Lucas (2019).^[14]
A study on the ontogenetic trajectories of septal spacing between succeeding chambers in Hypophylloceras subramosum and Phyllopachyceras ezoense is published online by Iwasaki, Iwasaki & Wani (2019).^[15]
A study on the intra- and interspecific variation and ontogenetic trajectories of the kossmaticeratid species Maorites densicostatus and Maorites seymourianus is published by Morón-Alfonso (2019).^[16]
A pathological ammonite specimen characterized by a pronounced left-right asymmetry in both ornamentation and suture lines is described from the Toarcian of southern France by Jattiot et al. (2019).^[17]
A study on the morphology and function of hook-like structures found in the body chambers of six specimens of Rhaeboceras halli is published online by Kruta et al. (2019).^[18]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Albanesiceras^[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Perisphinctidae. The type species is A. damianoi.
Albanites americanus^[20]	Sp. nov	Valid	Brayard et al.	Early Triassic		United States ( Idaho)
Allocrioceras irregulare^[21]	Sp. nov	Valid	Kennedy & Kaplan	Late Cretaceous (Turonian)		Germany
Anagaudryceras calabozoi^[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Cerro Toro Formation Rabot Formation Santa Marta Formation	Antarctica (James Ross Island) Chile
Anagaudryceras sanctuarium^[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian/Maastrichtian boundary)	Snow Hill Island Formation	Antarctica (James Ross Island)
Anagaudryceras subcompressum^[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Maastrichtian)	Snow Hill Island Formation	Antarctica (James Ross Island)
Araxoceltites^[23]	Gen. et 3 sp. nov	Valid	Korn, Ghaderi & Tabrizi	Permian (Changhsingian)		Iran	Genus includes new species A. sanestapanus, A. laterocostatus and A. cristatus.
Araxoceras insolens^[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran
Araxoceras truncatum^[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran
Balearites berryae^[25]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous		Spain
Beckeroceras^[26]	Gen. et comb. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)	Dzhaus Beds	Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Erbenoceras" khanakasuensis Yatskov (1990).
Beulia^[27]	Gen. et sp. nov	Valid	Korn & Price	Devonian (Famennian)		Germany	Genus includes new species B. wunderlichi.
Borealiceras^[28]	Gen. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous		Germany	Genus includes "Crioceras" stadtlaenderi Müller (1892).
Caribouceras^[20]	Gen. et sp. nov	Valid	Brayard et al.	Early Triassic		United States ( Idaho)	Genus includes new species C. slugense.
Carpathiceras^[29]	Gen. et comb. et sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia	A member of Asteroceratinae. The type species is "Coroniceras (Paracoroniceras)" bordoti Guérin-Franiatte (1966); genus also includes new species C. misiki.
Carthaginites multituberculatus^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the family Turrilitidae.
Chieseiceras dolomiticum^[31]	Sp. nov	Valid	Brack & Rieber	Middle Triassic (Late Anisian)		Italy
Chtelnicaceras^[29]	Gen. et sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia	An ammonite with an intermediate morphology between Arietitidae and Oxynoticeratidae. Genus includes new species C. janus.
Damesites africanus^[32]	Sp. nov	Valid	Kennedy & Lobitzer	Late Cretaceous (Cenomanian)	Odukpani Formation	Nigeria	A member of the family Desmoceratidae.
Docidoceras higginsi^[33]	Sp. nov	Valid	Chandler	Middle Jurassic (Bajocian)		United Kingdom
Dzhaprakoceras brodheadense^[34]	Sp. nov	Valid	Work & Mason	Carboniferous (Tournaisian)	Borden Formation	United States ( Kentucky)
Eogunnarites? elegans^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the family Kossmaticeratidae.
Epophioceras nodosus^[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Euomphaloceras varicostatum^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Falciclymenia cabrierensis^[27]	Sp. nov	Valid	Korn & Price	Late Devonian		France
Finiclymenia czarnockii^[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Finiclymenia langei^[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Flickia bullata^[36]	Sp. nov	Valid	Kennedy & Lobitzer	Late Cretaceous (Cenomanian)	Odukpani Formation	Nigeria
Follotites bighillensis^[34]	Sp. nov	Valid	Work & Mason	Carboniferous (Tournaisian)	Borden Formation	United States ( Kentucky)
Gaudryceras submurdochi^[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Rabot Formation Santa Marta Formation	Antarctica (James Ross Island)
Glaphyrites taimyricus^[37]	Sp. nov	Valid	Konovalova & Borissenkov in Borissenkov, Konovalova & Kutygin	Carboniferous	Makarovskaya Formation	Russia ( Krasnoyarsk Krai)
Glaucoceras^[38]	Gen. et comb. nov	Valid	Matamales-Andreu et al.	Early Cretaceous (Hauterivian)		Spain Switzerland	A new genus for "Baculites" renevieri Ooster (1860).
Glyptoxoceras (?) tenuicostatum^[21]	Sp. nov	Valid	Kennedy & Kaplan	Late Cretaceous (Turonian)		Germany	A member of the family Diplomoceratidae.
Hoevelia^[27]	Gen. et sp. nov	Valid	Korn & Price	Devonian (Famennian)		Germany	Genus includes new species H. schindewolfi.
Homoeoplanulites todaginensis^[39]	Sp. nov	Valid	Poulton & Hall	Middle Jurassic (Callovian)		Canada ( British Columbia)
Hoploscaphites macer^[40]	Sp. nov	Valid	Landman et al.	Late Cretaceous (Maastrichtian)	Bearpaw Shale Pierre Shale	United States ( Colorado Montana Wyoming)
Ivoites meshchankinae^[26]	Sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Czech Republic Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae.
Kamerunoceras multinodosum^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Turonian)	Karai Formation	India
Kayutoceras pogrebitskyi^[37]	Sp. nov	Valid	Borissenkov & Kutygin in Borissenkov, Konovalova & Kutygin	Carboniferous	Olen'ynskaya Formation	Russia ( Krasnoyarsk Krai)
Kirsoceras arteseptatum^[41]	Sp. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Germany
Kirsoceras gelamenum^[41]	Sp. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Germany
Kitabobactrites^[26]	Gen. et sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Czech Republic Uzbekistan	A member of Bactritida belonging to the family Bactritidae. The type species is K. salimovae.
Kumetaceras^[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A possible member of the family Spiroceratidae. The type species is K. enigmaticum.
Kunnamiceras^[30]	Gen. et comb. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India Madagascar	A member of the subfamily Acanthoceratinae. The type species is "Ammonites" tropicus Stoliczka (1865); genus also includes "Protacanthoceras" imperatoris Wright & Kennedy (1980).
Latiwitchellia atacamensis^[42]	Sp. nov	Valid	Dietze & von Hillebrandt	Middle Triassic (Bajocian)		Chile
Lissoceras maerteni^[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras maizetense^[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras ovale^[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)	El Pedregal Formation	France Germany Hungary Spain	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras sturanii^[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France Italy Russia ( Dagestan) Turkmenistan	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras submediterraneum^[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)	El Pedregal Formation	France Spain	A member of Haploceratoidea belonging to the family Lissoceratidae.
Loyezia roberti^[25]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous (Hauterivian)		Spain	A member of the family Crioceratitidae.
Maeneceras tabasense^[44]	Sp. nov	Valid	Korn et al.	Probably late early Famennian		Iran	A member of the family Sporadoceratidae.
Medioclymenia media^[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Megatirolites^[45]	Gen. et comb. nov	Valid	Balini et al.	Early Triassic (Olenekian)	Sefid-Kuh Limestone Tartaly Formation	Iran Kazakhstan	A member of Ceratitida belonging to the superfamily Dinaritoidea and to the family Tirolitidae. The type species is "Tirolites" rossicus Kiparisova (1947); genus also includes "Tirolites" armatus Shevyrev (1968).
Menuites armatus^[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Pachydiscidae.
Mesodistichites^[47]	Gen. et comb. et sp. nov	Valid	Krystyn et al.	Late Triassic (Norian)	Nayband Formation	Iran	A member of Ceratitida belonging to the family Distichitidae. The type species is "Distichites" tozeri Seyed-Emami (1975); genus also includes new species M. evolutus.
Metabactrites rakhmonovi^[26]	Sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae.
Microderoceras crassiornatum^[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Microderoceras serratum^[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Nelomites^[41]	Nom. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Kazakhstan	A replacement name for Melonites Bogoslovskii (1971).
Neophylloceras nodosum^[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Phylloceratidae.
Noridiscites nodosus^[47]	Sp. nov	Valid	Krystyn et al.	Late Triassic (Norian)	Nayband Formation	Iran	A member of Ceratitida belonging to the family Noridiscitidae.
Oraniceras calvum^[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Oraniceras issae^[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Pachydiscus wrighti^[49]	Sp. nov	Valid	Kennedy	Late Cretaceous		United Kingdom
Paracrioceras damaisi^[28]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous
Parasolenoceras ribiraense^[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Diplomoceratidae.
Paroecotraustes minor^[19]	Sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Oppeliidae.
Pernaceras^[28]	Gen. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous		France	Genus includes "Ammonites" assimilis Matheron (1880).
Phlycticeras tenue^[19]	Sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Strigoceratidae.
Praechebbites^[50]	Gen. et sp. et comb. nov	Valid	Becker et al.	Devonian (Emsian)		Morocco China? France?	A member of the family Mimosphinctidae belonging to the new subfamily Chebbitinae. The type species is P. debaetsi; genus might also include "Anetoceras (Erbenoceras)" mattei Feist (1970) and "Teicherticeras" ilanense Shen (1975).
Prolecanites akiyoshiensis^[51]	Sp. nov	Valid	Fujikawa	Carboniferous	Akiyoshi Limestone	Japan
Protacanthoceras parva^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the subfamily Acanthoceratinae.
Pseudocrassiceras^[52]	Gen. et comb et sp. nov	Valid	Rulleau & Jattiot	Early Jurassic (Toarcian)		France Spain	A member of the family Hildoceratidae. The type species is "Ammonites" bayani Dumortier (1874); genus also includes "Ammonites" frantzi Reynès (1868), "Hildoceras" comense var. praegruneri Monestier (1931) (raised to the rank of a separate species Pseudocrassiceras praegruneri) and a new species P. arbaulti.
Pseudosonninia^[53]	Gen. et sp. nov	Valid	Parent et al.	Middle Jurassic (Callovian)	Los Molles Formation	Argentina	A member of the family Oppeliidae. The type species is P. chacaymelehuensis.
Puzosia (Bhimaites) falx^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Schlueterella stinnesbecki^[54]	Sp. nov	Valid	Ifrim	Late Cretaceous (Turonian-Coniacian)		Mexico	A member of the family Diplomoceratidae.
Semilissoceras^[43]	Gen. et 3 sp. et comb. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		Algeria France Portugal Spain United Kingdom	A member of Haploceratoidea belonging to the family Lissoceratidae. The type species is S. ellipticum; genus also includes new species S. turgidulum and S. costellatum, as well as "Oppelia" depereti Flamand (1911) and "Lissoceras" semicostulatum Buckman (1923).
Sicilisphinctes^[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Perisphinctidae. The type species is S. incertus.
Siemiradzkia mangoldi^[55]	Sp. nov	Valid	Metodiev	Middle Jurassic (Bathonian)	Bov Formation	Bulgaria France	A member of the family Perisphinctidae.
Sikeliceras^[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A possible member of the family Spiroceratidae. The type species is S. costatum.
Silesites yajimai^[56]	Sp. nov	Valid	Matsukawa	Early Cretaceous (Aptian)	Sanyama Formation	Japan
Sokurella elshankae^[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Sokurella pletniovkae^[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Stephanoceras (Skirroceras) englandi^[57]	Sp. nov	Valid	Chandler	Middle Jurassic (Lower Bajocian)	Inferior Oolite Formation	United Kingdom
Stephanoceras (Riccardiceras) eoeteosum^[57]	Sp. nov	Valid	Chandler	Middle Jurassic (Middle or Upper Aalenian)	Inferior Oolite Formation	United Kingdom
Uptonia schlatteri^[58]	Sp. nov	Valid	Fauré & Téodori	Early Jurassic (Pliensbachian)		France
Uzbekisphinctes^[26]	Gen. et comb. nov	Valid	Naglik, De Baets & Klug	Early Devonian	Daleje Shale Dzhaus Beds Middle Kaub Formation	China Czech Republic Germany Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Teicherticeras" rudicostatum Bogoslovsky (1980); genus also includes "Convoluticeras" discordans Erben (1965) and "Teicherticeras" primigenitum Erben (1965).
Vedioceras fusiforme^[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran	A member of the family Araxoceratidae.
Watinoceras elegans^[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Zelandites pujatoi^[22]	Sp. nov	Valid	Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Rabot Formation Santa Marta Formation	Antarctica (James Ross Island)

Other cephalopods

Research

A study on hydrostatic properties of early Paleozoic nautiloid and endoceratoid cephalopods, and on its implications for the knowledge of ecology of these cephalopods, is published by Peterman, Barton & Yacobucci (2019).^[59]
A study on the distribution and diversity of cephalopods during the Cambrian–Ordovician transition is published by Fang et al. (2019).^[60]
A study on cephalopod occurrences in South China and adjacent areas during the Middle to Late Ordovician, aiming to define biogeographic provinces, is published by Fang et al. (2019).^[61]
A study on changes of geographic range size and rates of speciation and extinction in nautiloid and ammonoid cephalopods living in the North American Midcontinent Sea from the Carboniferous (Pennsylvanian)-early Permian is published by Kolis & Lieberman (2019).^[62]
Ascocerid fossils are described from the Hirnantian?–Llandovery strata of the Eusebio Ayala and Vargas Peña formations (southeastern Paraguay) by Cichowolski et al. (2019), representing the first ascocerid record from high paleolatitudes of Gondwana.^[63]
A juvenile specimen of Dolorthoceras, representing the first Devonian cephalopod specimen preserving nacreous structures and the first ectocochleate cephalopod preserving fibrous structures within its conch, is described from the Frasnian Domanic Formation (Russia) by Doguzhaeva (2019).^[64]
A study on the response of cephalopod assemblages from the area of the present-day European Russia to environmental changes during the early Aptian Oceanic Anoxic Event is published by Rogov et al. (2019).^[65]
A study on the paleobiology of cephalopods from the Albian Mahajanga Basin of Madagascar will be published by Hoffmann et al. (2019).^[66]
A proposal for revision of the classification of nautiloid cephalopods presented in Treatise Part K is presented by King & Evans (2019), who name a new subclass Tarphyceratia, new order Rioceratida and new family Bactroceratidae.^[67]
A study on embryonic and early juvenile development of shell in Peismoceras is published by Manda & Turek (2019).^[68]
A study on changes of the morphological diversity of Late Permian coiled nautiloids is published online by Korn et al. (2019).^[69]
A study on the paleobiology of Angulithes mermeti, based on new material from the upper Cenomanian of Egypt, and on its implications for the knowledge of the evolution of Cenomanian lineage of Angulithes, is published by Wilmsen & Nagm (2019).^[70]
A study on the anatomy of Gordoniconus beargulchensis and on its implications for the knowledge of evolution of coleoids is published by Klug et al. (2019).^[71]
A study on a putative Sinemurian octocoral Mesosceptron from Montagna del Casale (Sicily, Italy) is published by Pignatti et al. (2019), who reinterpret the fossils of this taxon as incompletely preserved coleoid rostra, and consider Mesosceptron to be a subjective junior synonym of the genus Atractites.^[72]
A study on four specimens of Clarkeiteuthis conocauda from the Toarcian Posidonia Shale (Germany) preserved with small bony fish in their arm crowns, on the feeding behaviour of members of this species, and on the differences in the mode of life of C. conocauda, Passaloteuthis bisulcata and Hibolithes semisulcatus, is published by Jenny et al. (2019).^[73]
A study on belemnite body-size dynamics across the Pliensbachian–Toarcian boundary in the Peniche section (Lusitanian Basin, Portugal), evaluating its implications for the knowledge of the impact of the Pliensbachian–Toarcian boundary warming event on belemnite body size, is published by Rita et al. (2019).^[74]
New belemnite fossils are described from the Berriasian–Hauterivian of Arctic Canada and North-East Greenland by Mutterlose et al. (2019), who also evaluate the implications of these fossils for the knowledge of spatial distribution patterns of belemnites at northern high latitudes.^[75]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Andamanoceras^[76]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species A. densiseptum.
Basiloceras^[77]	Gen. et sp. nov	Valid	Pohle, Klug & Haas	Devonian (Eifelian and Givetian)		Morocco	A member of Oncocerida belonging to the family Acleistoceratidae. Genus includes new species B. goliath and B. david.
Beloitoceras thorslundi^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Oncoceratidae.
Cabaneroceras^[79]	Gen. et sp. nov	Valid	Kröger & Gutiérrez-Marco	Ordovician (Darriwilian)		Spain	A member of Intejocerida. Genus includes new species C. aznari.
Cameroceras motsognir^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Cimomia kurkurensis^[80]	Sp. nov	Valid	Hewaidy, Qot & Moneer	Early Paleocene	Kurkur Formation	Egypt	A member of the family Hercoglossidae.
Clothoceras^[78]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae. Genus includes new species C. thornquisti.
Danoceras skalbergensis^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Deltoidonautilus hassani^[80]	Sp. nov	Valid	Hewaidy, Qot & Moneer	Late Paleocene	Tarawan Formation	Egypt	A member of the family Hercoglossidae.
Dicoelites aprilis^[81]	Sp. nov	Valid	Dzyuba in Dzyuba, Mitta & Sherstyukov	Middle Jurassic (Bajocian)	Djangura Formation	Russia ( Karachay-Cherkessia)	A belemnite.
Dicoelites octobris^[81]	Sp. nov	Valid	Dzyuba in Dzyuba, Mitta & Sherstyukov	Middle Jurassic (Bajocian)	Djangura Formation	Russia ( Karachay-Cherkessia)	A belemnite.
Discoceras amtjaernense^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Discoceras nilssoni^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Endoceras naekki^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Engeseriteuthis^[82]	Gen. et sp. nov	Valid	Fuchs et al.	Late Jurassic (late Kimmeridgian to ?early Tithonian)	Geisental Formation	Germany	A member of Muensterelloidea. Genus includes new species E. arcuatus.
Eromangateuthis^[83]	Gen. et comb. nov	Valid	Fuchs	Early Cretaceous (Albian)	Allaru Formation	Australia	A member of Octobrachia belonging to the family Plesioteuthidae; a new genus for "Boreopeltis" soniae Wade (1993).
Etchesia^[84]	Gen. et sp. nov	Valid	Fuchs	Late Jurassic (Tithonian)	Kimmeridge Clay Formation	United Kingdom	A member of Coleoidea belonging to the family Muensterellidae. Genus includes new species E. martilli.
Furudaloceras^[78]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Cyrtogomphoceratidae. Genus includes new species F. tomtei.
Grafordites^[85]	Gen. et sp. nov	Valid	Niko & Mapes	Carboniferous (Kasimovian)	Graford Formation	United States ( Texas)	A member of the family Dentoceratidae. The type species is G. mcleodi.
Gujiaonautilus^[86]	Gen. et sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid. Genus includes new species G. longliensis.
Isbergoceras^[78]	Gen. et 2 sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Graciloceratidae. The type species is I. niger; genus also includes I. consobrinum.
Isorthoceras nikwis^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Isorthoceras sylphide^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Isorthoceras urdr^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Kullsbergoceras^[78]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Cyrtogomphoceratidae. Genus includes new species K. nissei.
Lambeoceras (Lambeoceras) balashovi^[87]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Lambeoceras (Lambeoceras) hanae^[87]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Lambeoceras (Lambeoceras) sibiricum^[87]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Langgunites^[76]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species L. mucronulatus.
Malayorthoceras^[76]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida. Genus includes new species M. gracilentum.
Muensterella jillae^[82]	Sp. nov	Valid	Fuchs et al.	Late Cretaceous (Turonian)	South Bosque Formation	United States ( Texas)
Muensterella spinosa^[82]	Sp. nov	Valid	Fuchs et al.	Late Jurassic (Tithonian)	Mörnsheim Formation	Germany
Muensterellina^[82]	Gen. et sp. nov	Valid	Fuchs et al.	Middle Jurassic (Callovian)	Oxford Clay Formation	United Kingdom	A member of Muensterelloidea. Genus includes new species M. johnjagti.
Neotainoceras zhaoi^[86]	Sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid.
Nodopleuroceras gujiaoense^[86]	Sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid.
Ordogeisonoceras uppsalaensis^[78]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Geisonoceratidae.
Sepia vandervoorti^[88]	Sp. nov	Valid	Košťák, Jagt & Schlögl	Miocene (late Langhian–earliest Serravallian)	Köselerli Formation	Turkey	A species of Sepia.
Shanthaiceras^[76]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species S. amplum.
Sibumasuoceras kilimense^[76]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida.
Sibumasuoceras scrivenori^[76]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida.
Tafadnatoceras^[89]	Gen. et sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Tiouririne Formation	Morocco	A member of Orthocerida belonging to the family Stereoplasmoceratidae. The type species is T. tiouririnense. Announced in 2019; the final version of the article naming it is not published yet.
Tofangoceras kedahense^[76]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tofangoceras rayense^[76]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tofangoceras staiti^[76]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tyrionella^[82]	Gen. et sp. nov	Valid	Fuchs et al.	Late Jurassic (Tithonian)	Altmühltal Formation	Germany	A member of Muensterelloidea. Genus includes new species T. fauseri.
Valkyrioceras^[78]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Valcouroceratidae. Genus includes new species V. dalecarlia.
Wadema tattai^[89]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of Actinocerida belonging to the family Wademidae. Announced in 2019; the final version of the article naming it is not published yet.

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acharax svalbardensis^[168]	Sp. nov	Valid	Hansen et al.	Late Pleistocene		Fram Strait	A bivalve belonging to the family Solemyidae.
Atomodesma? hautmanni^[126]	Sp. nov	Valid	Foster et al.	Early Triassic		China	A bivalve.
Atrina multicostata^[169]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Atrina.
Baxoitrigonia^[170]	Gen. et sp. nov		Stiller & Chen	Late Jurassic (Tithonian)	Lagongtang Formation	China	A bivalve. Genus includes new species B. baxoiensis.
Bengtsonella^[171]	Gen. et comb. nov	Junior homonym	Cooper & Leanza	Early Cretaceous (Albian)	Riachuelo Formation	Brazil	A bivalve belonging to the family Megatrigoniidae and the subfamily Megatrigoniinae. The type species is "Anditrigonia" britoi Hessel (2005). The generic name is preoccupied by Bengtsonella Muller & Hinz (1991) and Bengtsonella Mostler (1996); Ceccolini & Cianferoni (2021) coined a replacement name Bengtsonigonia.^[172]
Californigonia^[171]	Gen. et comb. nov	Valid	Cooper & Leanza	Jurassic (Callovian–early Oxfordian)	Bicknell Sandstone	Canada ( British Columbia) United States ( California)	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" plumasensis Hyatt (1892).
Callocardia (Nitidavenus) youssoufia^[98]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Callucina itaporangensis^[173]	Sp. nov	Valid	Ayoub-Hannaa et al.	Cretaceous	Cotinguiba Formation	Brazil United Kingdom	A bivalve belonging to the family Lucinidae.
Cardiocardita peregrina^[98]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Carditella pitufina^[174]	Sp. nov	Valid	Pérez	Early Pliocene		Argentina	A bivalve belonging to the family Condylocardiidae.
Chlamys (Praechlamys) prima^[169]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Chlamys.
Ckaraosippur^[175]	Gen. et comb. et sp. nov	Valid	Santelli & del Río	Miocene–Pliocene	Bahía Inglesa Formation Camarones Formation Navidad Formation	Argentina Chile	A scallop. The type specie is "Pecten" calderensis Möricke (1896); genus also includes new species C. camachoi.
Conchocele yatsuoensis^[176]	Sp. nov	Valid	Amano et al.	Early Miocene	Kurosedani Formation	Japan	A bivalve.
Costacallista discrepans^[98]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Craginella^[171]	Gen. et comb. nov	Valid	Cooper & Leanza	Late Jurassic (Kimmeridgian)	Malone Formation	Mexico United States ( Louisiana Texas)	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" goodellii Cragin (1897).
Damborenella^[171]	Gen. et comb. nov	Valid	Cooper & Leanza	Late Jurassic (Tithonian)		Argentina Chile Colombia Peru	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" eximia Philippi (1899); genus also includes "Trigonia" discors Philippi (1899), "Anditrigonia" lamberti Levy (1967) and "Anditrigonia eximia" tesselicaudata Leanza (1993) (raised to the rank of a separate species Damborenella tessellicaudata).
Dentalium mcganna^[177]	Sp. nov	Valid	Powell, Clites & Poust	Miocene	Sobrante Formation	United States ( California)	A species of Dentalium.
Dietotenhosen^[175]	Gen. et comb. nov	Valid	Santelli & del Río	Miocene–Pliocene	Bahía Inglesa Formation Coquimbo Formation Horcón Formation Hornillos Formation Huaricangana Formation La Portada Formation Pisco Formation Taime Formation	Chile Peru	A scallop. Genus includes D. hupeanus (Philippi, 1887) and D. remondi (Philippi, 1887).
Dosinia (Dosinisca) vredenburgi^[178]	Sp. nov	Valid	Borkar & Kulkarni	Miocene	Chhasra Formation Khari Nadi Formation Lower Mekran Series	India Pakistan	A bivalve belonging to the family Veneridae; a species of Dosinia.
Dosinia (Dosinisca) wynnei^[178]	Sp. nov	Valid	Borkar & Kulkarni	Miocene	Chhasra Formation Khari Nadi Formation Lower Mekran Series	India Pakistan	A bivalve belonging to the family Veneridae; a species of Dosinia.
Entolium reticulatum^[169]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve.
Eosphaera^[179]	Gen. et comb. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Jurassic (Callovian to Kimmeridgian)		Egypt India Madagascar Portugal	A bivalve belonging to the family Lucinidae. The type species is "Sphaera" madagascariensis Newton (1889).
Chaetopleura abbessi^[180]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Chattian) and Miocene (Burdigalian)^[181]		France	A chiton. Originally described as a species of Chaetopleura; subsequently transferred to the genus Ischnochiton by Dell'Angelo et al. (2018).^[181]
Chaetopleura gaasi^[180]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Rupelian)		France	A chiton, Originally described as a species of Chaetopleura; subsequently transferred to the new genus Spinochiton by Dell'Angelo et al. (2018).^[181]
Helcionella lemdadensis^[182]	Sp. nov	Valid	Geyer, Valent & Meier	Cambrian	Tannenknock Formation	Germany
Hoernesia? danisa^[126]	Sp. nov	Valid	Foster et al.	Early Triassic		China	A bivalve.
Homopleura^[183]	Gen. et comb. et 2 sp. nov	Valid	Masse & Fenerci-Masse	Cretaceous		Bulgaria France	A rudist bivalve. The type species is "Monopleura" affinis Matheron (1878); genus also includes "Monopleura" imbricata Matheron (1842) and "Monopleura" coquandi Matheron (1878), as well as new species H. rustrella and H. balkanica.
Inoceramus chiplonkari^[30]	Sp. nov	Valid	Walaszczyk in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian and Turonian)	Karai Formation	India Japan
Ischnochiton crovatoi^[184]	Sp. nov	Valid	Dell'Angelo, Sosso & Bonfitto	Pleistocene		Italy	A chiton, a species of Ischnochiton.
Ischnochiton fehsei^[180]	Sp. nov	Valid	Cherns & Schwabe	Eocene		France	A chiton, a species of Ischnochiton.
Lasaea firma^[185]	Sp. nov	Valid	Lesport & Lozouet in Lesport et al.	Miocene (Aquitanian)		France	A bivalve belonging to the family Lasaeidae.
Leptostega frankenwaldensis^[182]	Sp. nov	Valid	Geyer, Valent & Meier	Cambrian	Tannenknock Formation	Germany	A helcionelloid.
Neilo altamirano^[108]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A bivalve belonging to the family Malletiidae.
Neocrassina (Pruvostiella) madagascariensis^[179]	Sp. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Late Jurassic (Oxfordian)	Morondava Basin	Madagascar	A bivalve belonging to the family Astartidae.
Palaeonucula feruglioi^[186]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Parailsanella luonanensis^[187]	Sp. nov	Valid	Li et al.	Cambrian Series 2	Xinji Formation	China	A helcionellid.
Peronaea basteroti^[185]	Sp. nov	Valid	Lesport, Lozouet & Pacaud in Lesport et al.	Miocene (Aquitanian)		France	A bivalve, a species of Peronaea.
Phaenodesmia? piatnitzkyi^[186]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Pinna simionescui^[169]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Pinna.
Platymyoidea morrisi^[188]	Sp. nov	Valid	Hodges	Early Jurassic		United Kingdom	A bivalve.
Pleurophopsis matsumotoi^[189]	Sp. nov	Valid	Amano et al.	Late Oligocene to early Miocene	Nabae Group	Japan	A bivalve belonging to the family Vesicomyidae.
Pleurophopsis talarensis^[108]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A bivalve belonging to the family Vesicomyidae.
Praedicerocardium^[169]	Gen. et sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve. Genus includes new species P. vetulus.
Praenucula pojetai^[89]	Sp. nov	In press	Ebbestad et al.	Late Ordovician		Morocco	A bivalve belonging to the family Praenuculidae. Announced in 2019; the final version of the article naming it is not published yet.
Procardia inouei^[190]	Sp. nov	Valid	Amano	Paleocene	Katsuhira Formation	Japan	A bivalve belonging to the group Anomalodesmata and to the family Parilimyidae.
Pseudophopsis^[108]	Gen. et comb. nov	Valid	Kiel et al.	Eocene and Early Oligocene	Talara Basin	Cuba Peru	A bivalve belonging to the group Heterodonta and to the family Kalenteridae. The type species is "Pleurophopsis" peruviana Olsson (1931); genus also includes "Unio" bitumen Cooke (1919).
Quenstedtia madagascariensis^[179]	Sp. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Late Jurassic (Oxfordian)	Morondava Basin	Madagascar	A bivalve belonging to the family Quenstedtiidae.
Ramentoides xixiangensis^[191]	Sp. nov	Valid	Qin et al.	Cambrian		China	A member of Maikhanellidae.
Romaniamya^[169]	Gen. et sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve. Genus includes new species R. mahmudiaensis.
Rutitrigonia cintarojensis^[150]	Sp. nov	Valid	Cataldo et al.	Early Cretaceous (Barremian)	Huitrín Formation	Argentina	A bivalve.
Ryderia tehuelchana^[186]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Scythentolium anisicum^[169]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve.
Sogdianagonia^[192]	Nom. nov	Valid	Schneider & Kelly	Late Cretaceous (Cenomanian to Turonian)		Kyrgyzstan Uzbekistan	A bivalve belonging to the family Megatrigoniidae; a replacement name for Turkestanella Tashiro (1979). The type species is "Trigonia" turkestanensis Arkhangelsky (1916).
Solemya lucifuga^[193]	Sp. nov	Valid	Pérez-Barría & Nielsen	Early Miocene	Ranquil Formation	Chile	A bivalve, a species of Solemya.
Stenoplax monila^[180]	Sp. nov	Disputed	Cherns & Schwabe	Oligocene (Rupelian)		France	A chiton. Originally described as a species of Stenoplax; Dell'Angelo et al. (2020) considered this species to be a junior synonym of Lepidochitona oligocaena.^[194]
Tindaria kretensis^[195]	Sp. nov	Valid	Koskeridou, La Perna & Giamali	Early Pliocene		Greece	A bivalve.
Tonicella lira^[180]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Chattian)		France	A chiton, a species of Tonicella.
Vaccinites alceotarlaoi^[196]	Sp. nov	Valid	Pons et al.	Late Cretaceous (early Campanian)		Serbia Spain	A rudist bivalve.
Vaccinites bilottei^[196]	Sp. nov	Valid	Pons et al.	Late Cretaceous (late Campanian)		Bulgaria	A rudist bivalve.
Vaccinites pirenaicus^[196]	Sp. nov	Valid	Pons et al.	Late Cretaceous (late Campanian)		Spain	A rudist bivalve.
Wareniconcha mercenarioides^[197]	Sp. nov	Valid	Kase et al.	Pliocene		Philippines	A bivalve belonging to the family Vesicomyidae.
Xylophagella littlei^[198]	Sp. nov	Valid	Hryniewicz in Hryniewicz et al.	Late Paleocene	Basilika Formation	Norway	A bivalve belonging to the family Pholadidae.
Yoldiella spitsbergensis^[198]	Sp. nov	Valid	Amano in Hryniewicz et al.	Late Paleocene	Basilika Formation	Norway	A bivalve belonging to the family Yoldiidae.

2019 in paleomalacology

Ammonites

Research

New taxa

Other cephalopods

Research

New taxa

Gastropods

Research

New taxa

Other molluscs

Research

New taxa

References

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